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Pseudhipparion retrusum
Taxonomy
Hippotherium retrusum was named by Cope (1889). Its type specimen is AMNH 8350, a set of teeth (two upper molars), and it is a 3D body fossil.
It was recombined as Hipparion retrusum by Trouessart (1898), Hay (1902); it was recombined as Neohipparion retrusum by Matthew (1909), Merriam (1917); it was recombined as Protohippus retrusus by Osborn (1918) and Hay (1930); it was recombined as Nannippus retrusus by McGrew and Meade (1938), Stirton (1940) and Lance (1950); it was recombined as Griphippus retrusus by Quinn (1955); it was considered a nomen dubium by Macdonald (1992); it was recombined as Pseudhipparion retrusum by Ameghino (1904), Webb (1969), Webb and Hulbert (1986), Kelly and Lander (1988), MacFadden (1998).
It was recombined as Hipparion retrusum by Trouessart (1898), Hay (1902); it was recombined as Neohipparion retrusum by Matthew (1909), Merriam (1917); it was recombined as Protohippus retrusus by Osborn (1918) and Hay (1930); it was recombined as Nannippus retrusus by McGrew and Meade (1938), Stirton (1940) and Lance (1950); it was recombined as Griphippus retrusus by Quinn (1955); it was considered a nomen dubium by Macdonald (1992); it was recombined as Pseudhipparion retrusum by Ameghino (1904), Webb (1969), Webb and Hulbert (1986), Kelly and Lander (1988), MacFadden (1998).
Synonymy list
| Year | Name and author |
|---|---|
| 1889 | Hippotherium retrusum Cope p. 446 |
| 1898 | Hipparion retrusum Trouessart |
| 1902 | Hipparion retrusum Hay p. 620 |
| 1904 | Pseudhipparion retrusum Ameghino |
| 1907 | Hippotherium retrusum Gidley p. 906 |
| 1909 | Neohipparion retrusum Matthew |
| 1917 | Neohipparion retrusum Merriam |
| 1918 | Protohippus retrusus Osborn p. 142 figs. Text Fig. 114 |
| 1930 | Protohippus retrusus Hay |
| 1938 | Nannippus retrusus McGrew and Meade p. 201 |
| 1940 | Nannippus retrusus Stirton p. 186 |
| 1950 | Nannippus retrusus Lance |
| 1955 | Griphippus retrusus Quinn |
| 1969 | Pseudhipparion retrusum Webb |
| 1986 | Pseudhipparion retrusum Webb and Hulbert, Jr. |
| 1988 | Pseudhipparion retrusum Kelly and Lander |
| 1998 | Pseudhipparion retrusum MacFadden p. 548 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Pseudhipparion retrusum Cope 1889
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| H. F. Osborn 1918 | (Matthew, 1913, Osborn, 1918) (1) Molar size intermediate, somewhat larger than in P. placidus but
apparently with shorter crowns; (2) crowns strongly curved; (3) protocone separate from protoloph in m2, united in m1; (4) protocone and hypocone united by longitudinal isthmus; (5) protocone flattened oval, posterior in position; (6) pli caballin fold present on little worn teeth, absent on half worn teeth; (7) two folds and minor accessory folds on each side of metaloph projecting into pre- and postfossettes; (8) a single inferior fold (crochet) in prefossette; (9) parastyle narrow, mesostyle broad at base and narrowing rather slightly toward crown. (Matthew, 1913) "These two type teeth cannot go together as placed; they may not belong to the same animal; the right hand tooth (A) might be Merychippus; the left hand tooth, if normal, does not agree well with any of the genera. If normal it represents an undescribed genus on the criteria used to separate the Miocene and Pliocene genera. If placed in Protohippus it should be in quotes." |
Measurements
No measurements are available
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| Source: g = genus, subf = subfamily, subc = subclass, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, MacFadden and Cerling 1996, Carroll 1988, MacFadden 1998, Ji et al. 2002, Lillegraven 1979 | |||||
Age range: base of the Barstovian to the top of the Clarendonian or 16.30000 to 9.40000 Ma
Collections (28 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Miocene | USA (Nebraska) | Pseudhipparion retrusum (18274) | |
| Miocene | USA (South Dakota) | Pseudhipparion retrusum (18183) | |
| Barstovian | USA (Nebraska) | Pseudhipparion retrusum (18124 18216 18442 18443) | |
| Barstovian - Clarendonian | USA (Nebraska) | Pseudhipparion retrusum (18077) | |
| Clarendonian | USA (Nebraska) | Pseudhipparion retrusum (18078 18082 18106 18148 18159 18176 18217 18242 18243 18260 18334 18372 18413 18414 18466) Pseudhipparion sp. (18314 18315 18316) | |
| Clarendonian | USA (South Dakota) | Pseudhipparion retrusum (18428) | |
| Clarendonian - Hemphillian | USA (Wyoming) | Merychippus sp. (18424) Pseudhipparion retrusum (18146) |