PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Cladosictis

Mammalia - Sparassodonta

Taxonomy

Cladosictis was named by Ameghino (1887). It is not extant.

It was assigned to Hathlyacynidae by Ameghino (1894); to Hathliacyninae by Marshall (1978); to Hathlyacyninae by Marshall (1981); to Borhyaenidae by Stirton (1953) and McKenna and Bell (1997); and to Hathliacynidae by de Muizon (1994), de Muizon (1999) and Goin et al. (2007).

Species

Acyon bardus, Anatherium defossus, C. centralis, C. defossa (syn. Agustylus bardus), C. patagonica (syn. Protoproviverra ensidens, Hathlyacynus linchyi, Agustylus cynoides, Hathliacynus lustratus, Amphithereutes obscurus, Ictioborus fenestratus, Hathliacynus fischeri, Agustylus primaevus, C. lateralis, Hathliacynus cultridens, Agustylus minus, C. dissimilis, C. petersoni, Hathliacynus rollieri, C. trouessarti)

Synonyms

Anatherium defossus was named by Ameghino (1887) [Not the original recombination]. Its type specimen is MACN 669, a mandible (fragment of right mandibular ramus with roots of i1-3, base of c, base of p1, p2 complete), and it is a 3D body fossil. Its type locality is Cerro Observatorio, which is in a Santacrucian terrestrial horizon in the Santa Cruz Formation of Argentina.

It was recombined as Cladosictis defossa by Marshall (1978); it was considered a nomen dubium by Forasiepi et al. (2006).
Acyon bardus was named by Ameghino (1889) [title: Enumeracion synoptique des especes de mammiferes fossiles des formaciones eocenes de Patagonie Ameghino first named Acyon bardus in 1889, and then recombined this species as Agustylus bardus in this reference. ]. Its type specimen is MACN 9, a mandible (fragment of left mandibular ramus with roots of m1-2, m3 complete, m4 missing tip of protoconid—crowns of m3-4 are heavily worn), and it is a 3D body fossil. Its type locality is Santa Cruz, which is in a Santacrucian terrestrial horizon in the Santa Cruz Formation of Argentina.

It was recombined as Agustylus bardus by Ameghino (1894); it was synonymized subjectively with Cladosictis defossa by Marshall (1978); it was synonymized subjectively with Anatherium defossus by Marshall (1981); it was considered a nomen dubium by Forasiepi et al. (2006).
Anatherium was named by Ameghino (1887) [Enumeracion sistematica de las especies de mamiferos fosiles coleccionados por Carlos Ameghino en los terrenos eocenos de la Patagonia austral y depositados en el Museo La Plata].

It was synonymized subjectively with Cladosictis by Marshall (1978) and Forasiepi et al. (2006).

It was assigned to Creodonta by Ameghino (1891); to Hathlyacynidae by Ameghino (1894); to Hathlyacyninae by Marshall (1981); and to Borhyaenidae by McKenna and Bell (1997).
Hathliacynus was named by Ameghino (1887) [title: Enumeracion sistematica de las especies de mamiferos fosiles coleccionados por Carlos Ameghino en los terrenos eocenos de la Patagonia austral y depositados en el Museo La Plata]. It is the type genus of Hathliacynidae.

It was synonymized subjectively with Cladosictis by Marshall (1978) and Marshall (1981).

It was assigned to Hathlyacynidae by Ameghino (1894).
Agustylus was named by Ameghino (1887) [title: Enumeracion sistematica de las especies de mamiferos fosiles coleccionados por Carlos Ameghino en los terrenos eocenos de la Patagonia austral y depositados en el Museo La Plata].

It was synonymized subjectively with Cladosictis by Marshall (1978) and Marshall (1981).

It was assigned to Creodonta by Ameghino (1891); and to Amphiproviverridae by Ameghino (1894).
Ictioborus was named by Ameghino (1891) [title: Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el eoceno inferior de la Patagonia austral].

It was synonymized subjectively with Cladosictis by Marshall (1978) and Marshall (1981).

It was assigned to Creodonta by Ameghino (1891); and to Acyonidae by Ameghino (1894).

Synonymy list

Year	Name and author
1887	Anatherium defossus Ameghino
1887	Cladosictis Ameghino
1887	Agustylus Ameghino p. 7
1887	Hathliacynus Ameghino p. 7
1887	Anatherium Ameghino p. 8
1889	Acyon bardus Ameghino p. 292
1891	Agustylus Ameghino p. 29
1891	Anatherium Ameghino p. 29
1891	Anatherium defossus Ameghino p. 29
1891	Ictioborus Ameghino p. 29
1894	Anatherium Ameghino p. 126
1894	Cladosictis Ameghino p. 126
1894	Hathlyacynus Ameghino p. 126
1894	Agustylus Ameghino p. 135
1894	Agustylus bardus Ameghino p. 136
1894	Ictioborus Ameghino p. 139
1953	Cladosictis Stirton p. 614
1978	Cladosictis Marshall
1978	Cladosictis defossa Marshall
1981	Anatherium Marshall
1981	Anatherium defossus Marshall
1981	Cladosictis Marshall
1994	Cladosictis de Muizon
1997	Anatherium McKenna and Bell
1997	Cladosictis McKenna and Bell
1999	Cladosictis de Muizon p. 507
2007	Cladosictis Goin et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()

Rank	Name	Author
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
subclass	Metatheria	()
—	Marsupialiformes
order	Sparassodonta	()
subfamily	Hathlyacininae	()
genus	Cladosictis	Ameghino 1887

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Cladosictis Ameghino 1887

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†Cladosictis centralis Ameghino 1902

†Cladosictis defossa Ameghino 1887

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Invalid names: Agustylus bardus Ameghino 1889 [synonym]

†Cladosictis patagonica Ameghino 1887

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Invalid names: Agustylus cynoides Ameghino 1887 [synonym], Agustylus minus Ameghino 1935 [synonym], Agustylus primaevus Mercerat 1891 [synonym], Amphithereutes obscurus Ameghino 1935 [synonym], Cladosictis dissimilis Mercerat 1891 [synonym], Cladosictis lateralis Ameghino 1894 [synonym], Cladosictis petersoni Sinclair 1906 [synonym], Cladosictis trouessarti Ameghino 1891 [synonym], Hathliacynus cultridens Mercerat 1891 [synonym], Hathliacynus fischeri Mercerat 1891 [synonym], Hathliacynus lustratus Ameghino 1887 [synonym], Hathliacynus rollieri Mercerat 1891 [synonym], Hathlyacynus linchyi Mercerat 1891 [synonym], Ictioborus fenestratus Ameghino 1891 [synonym], Protoproviverra ensidens Ameghino 1891 [synonym]

Invalid names: Acyon bardus Ameghino 1889 [nomen dubium], Agustylus Ameghino 1887 [synonym], Anatherium Ameghino 1887 [synonym], Anatherium defossus Ameghino 1887 [nomen dubium], Hathliacynus Ameghino 1887 [synonym], Ictioborus Ameghino 1891 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Maximum body mass:

3.7 kgs

Minimum body mass:

3.7 kgs

Body mass estimate:

3.7 kg

Environment:

terrestrialsubo

Locomotion:

actively mobilec

Life habit:

scansorialsubc

Diet:

carnivoresubo

Reproduction:

viviparoussubc

Created:

2013-05-24 23:51:13

Modified:

2013-05-24 08:51:13

Source: s = species, subo = suborder, subc = subclass, c = class, subp = subphylum

References: Lillegraven 1979, Hendy et al. 2009, Carroll 1988, Goin 1997, Vizcaíno et al. 2010, Clemens 1979

Age range: base of the Burdigalian to the top of the Friasian or 20.44000 to 15.50000 Ma

Collections (33 total)

Time interval	Ma	Country or state	Original ID and collection number
Middle Eocene - Early/Lower Miocene	47.8 - 15.97	Argentina (Chubut)	C. sp. (176461)
Colhuehuapian	21.0 - 17.5	Argentina (Neuquén)	C. sp. (89029)
Colhuehuapian	21.0 - 17.5	Argentina (Chubut)	C. sp., C. centralis (141368)
Colhuehuapian	21.0 - 17.5	Argentina (Río Negro)	C. patagonica (89093)
Burdigalian	20.44 - 15.97	Chile (Aisén)	C. sp. (71843)
Santacrucian	17.5 - 16.3	Argentina (Santa Cruz)	Agustylus cynoides, C. patagonica (176593) Agustylus primaevus, Hathliacynus fischeri, C. dissimilis (176563) C. patagonica (27313 142529 176565 176567 176569 176572 176576 176581 176584 176590 176594 176598 176599 176602 176635 176636 210153 210638) C. patagonica, Agustylus minus, Ictioborus fenestratus, Protoproviverra ensidens, Anatherium defossus (176570) C. patagonica, C. lateralis (142533) C. patagonica, C. petersoni (176585) C. patagonica, Hathliacynus lustratus, Hathliacynus cultridens, Hathliacynus rollieri, Amphithereutes obscurus, Acyon bardus (176575) C. patagonicus, C. patagonica (176578) Proviverra trouessarti (176596)
Friasian	16.3 - 15.5	Chile (Aisén)	C. patagonica (139845)
Langhian	15.97 - 13.82	Argentina (Rio Negro)	C. patagonica (198569)