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Hathlyacininae
Taxonomy
Hathliacynidae was named by Ameghino (1894). It is not extant. Its type is Hathliacynus. It was considered monophyletic by de Muizon (1999).
It was reranked as the subfamily Hathliacyninae by Marshall (1977) and Marshall (1978); it was reranked as the subfamily Hathlyacyninae by Patterson and Marshall (1978), Marshall (1981) and Villarroel and Marshall (1982); it was reranked as the subfamily Hathlyacininae by Hoffstetter and Petter (1983); it was corrected as Hathlyacynidae by Goin et al. (2013).
It was assigned to Borhyaenidae by Marshall (1977), Marshall (1978), Patterson and Marshall (1978), Marshall (1981), Villarroel and Marshall (1982) and Hoffstetter and Petter (1983); to Borhyaenoidea by de Muizon (1994) and de Muizon (1999); to Sparassodonta by Ameghino (1894) and Oliveira and Goin (2011); and to Sparassodonta by Martín and Tejedor (2007), Goin et al. (2007), Goin et al. (2010) and Goin et al. (2013).
It was reranked as the subfamily Hathliacyninae by Marshall (1977) and Marshall (1978); it was reranked as the subfamily Hathlyacyninae by Patterson and Marshall (1978), Marshall (1981) and Villarroel and Marshall (1982); it was reranked as the subfamily Hathlyacininae by Hoffstetter and Petter (1983); it was corrected as Hathlyacynidae by Goin et al. (2013).
It was assigned to Borhyaenidae by Marshall (1977), Marshall (1978), Patterson and Marshall (1978), Marshall (1981), Villarroel and Marshall (1982) and Hoffstetter and Petter (1983); to Borhyaenoidea by de Muizon (1994) and de Muizon (1999); to Sparassodonta by Ameghino (1894) and Oliveira and Goin (2011); and to Sparassodonta by Martín and Tejedor (2007), Goin et al. (2007), Goin et al. (2010) and Goin et al. (2013).
Subtaxa
Acyon, Acyonidae, Amphoproviverridae, Andinogale, Borhyaenidium, Chasicostylus, Cladictidae, Cladosictidae, Cladosictinae, Cladosictis (syn. Agustylus, Hathliacynus, Anatherium, Ictioborus), Contrerascynus (syn. Simpsonia), Notictis, Notocynus, Notogale, Patene (syn. Ischyrodidelphis), Perathereutes, Procladosictis, Pseudocladosictis, Pseudocladosictis determinabile, Pseudonotictis, Sallacyon, Sipalocyon (syn. Amphithereutes, Amphiproviverra, Protoproviverra, Thylacodictis)
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1889 | Acyonidae Ameghino p. 894 |
| 1894 | Hathlyacynidae Ameghino p. 126 |
| 1894 | Amphiproviverridae Ameghino p. 131 |
| 1894 | Acyonidae Ameghino p. 139 |
| 1894 | Hathliacynidae Ameghino p. 382 |
| 1894 | Amphoproviverridae Ameghino p. 389 |
| 1902 | Pseudocladosictis Ameghino p. 47 |
| 1902 | Pseudocladosictis determinabile Ameghino p. 47 |
| 1923 | Cladictidae Winge p. 77 |
| 1927 | Cladosictinae Cabrera p. 273 |
| 1935 | Cladosictidae Ameghino p. 131 |
| 1977 | Hathliacyninae Marshall p. 419 |
| 1978 | Hathliacyninae Marshall |
| 1978 | Hathlyacyninae Patterson and Marshall |
| 1981 | Hathlyacyninae Marshall |
| 1982 | Hathlyacyninae Villarroel and Marshall |
| 1983 | Hathlyacininae Hoffstetter and Petter |
| 1994 | Hathliacynidae de Muizon |
| 1999 | Hathliacynidae de Muizon p. 507 |
| 2007 | Hathliacynidae Goin et al. |
| 2007 | Hathliacynidae Martín and Tejedor p. 747 |
| 2010 | Hathliacynidae Goin et al. |
| 2011 | Hathliacynidae Oliveira and Goin |
| 2013 | Hathlyacynidae Goin et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. †Hathlyacininae Ameghino 1894
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G. †Acyon Ameghino 1887
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†Acyon myctoderos Forasiepi et al. 2006
†Acyon tricuspidatus Ameghino 1887
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Invalid names: Anatherium herrerae Marshall 1981 [synonym], Ictioborus destructor Ameghino 1894 [synonym]
G. †Borhyaenidium Pascual and Bocchino 1963
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†Borhyaenidium altiplanicus Villarroel and Marshall 1983
†Borhyaenidium musteloides Pascual and Bocchino 1963
†Borhyaenidium riggsi Marshall 1981
G. †Cladosictis Ameghino 1887
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†Cladosictis centralis Ameghino 1902
†Cladosictis patagonica Ameghino 1887
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Invalid names: Agustylus cynoides Ameghino 1887 [synonym], Agustylus minus Ameghino 1935 [synonym], Agustylus primaevus Mercerat 1891 [synonym], Amphithereutes obscurus Ameghino 1935 [synonym], Cladosictis dissimilis Mercerat 1891 [synonym], Cladosictis lateralis Ameghino 1894 [synonym], Cladosictis petersoni Sinclair 1906 [synonym], Cladosictis trouessarti Ameghino 1891 [synonym], Hathliacynus cultridens Mercerat 1891 [synonym], Hathliacynus fischeri Mercerat 1891 [synonym], Hathliacynus lustratus Ameghino 1887 [synonym], Hathliacynus rollieri Mercerat 1891 [synonym], Hathlyacynus linchyi Mercerat 1891 [synonym], Ictioborus fenestratus Ameghino 1891 [synonym], Protoproviverra ensidens Ameghino 1891 [synonym]
Invalid names: Acyon bardus Ameghino 1889 [nomen dubium], Agustylus Ameghino 1887 [synonym], Anatherium Ameghino 1887 [synonym], Anatherium defossus Ameghino 1887 [nomen dubium], Hathliacynus Ameghino 1887 [synonym], Ictioborus Ameghino 1891 [synonym]
G. †Contrerascynus Mones 2014
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†Contrerascynus borhyaenoides Contreras 1990
Invalid names: Simpsonia Contreras 1990 [replaced]
G. †Notictis Ameghino 1889
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†Notictis ortizii Ameghino 1889
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Invalid names: Didelphys curvidens Burmeister 1891 [objective synonym]
G. †Patene Simpson 1935
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†Patene campbelli Goin and Candela 2004
†Patene coluapiensis Simpson 1935
†Patene simpsoni Paula Couto 1952
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Invalid names: Ischyrodidelphis castellanosi Paula Couto 1952 [synonym]
Invalid names: Ischyrodidelphis Paula Couto 1952 [synonym]
G. †Perathereutes Ameghino 1891
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†Perathereuthes amputans Ameghino 1891
†Perathereuthes obtusus Ameghino 1891
G. †Procladosictis Ameghino 1902
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†Procladosictis anomala Ameghino 1902
Invalid names: Procladosictis erecta Ameghino 1902 [nomen vanum]
G. †Sipalocyon Ameghino 1887
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†Sipalocyon externus Ameghino 1902
†Sipalocyon gracilis Ameghino 1887
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Invalid names: Amphiproviverra crassa Ameghino 1894 [synonym], Amphiproviverra minuta Ameghino 1894 [synonym], Hathliacynus lynchi Mercerat 1891 [synonym], Perathereutes amputans Ameghino 1891 [synonym], Perathereutes obtusus Ameghino 1891 [synonym], Protoproviverra manzaniana Ameghino 1891 [synonym], Protoviverra manzaniana Ameghino 1891 [synonym], Sipalocyon altiramis Ameghino 1894 [synonym], Sipalocyon curtus Ameghino 1894 [synonym], Sipalocyon longus Ameghino 1894 [synonym], Sipalocyon mixtus Ameghino 1894 [synonym], Thylacodictis exilis Mercerat 1891 [synonym]
†Sipalocyon obusta Ameghino 1891
Invalid names: Amphiproviverra Ameghino 1891 [synonym], Amphithereutes Ameghino 1935 [synonym], Protoproviverra Ameghino 1891 [synonym], Thylacodictis Mercerat 1891 [synonym]
Invalid names: Acyonidae Ameghino 1889 [invalid subgroup], Amphoproviverridae Ameghino 1894 [invalid subgroup], Cladictidae Winge 1923 [invalid subgroup], Cladosictidae Ameghino 1935 [invalid subgroup], Cladosictinae Cabrera 1927 [invalid subgroup], Pseudocladosictis Ameghino 1902 [nomen vanum], Pseudocladosictis determinabile Ameghino 1902 [nomen vanum]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: s = species, subo = suborder, subc = subclass, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Lillegraven 1979, Carroll 1988, Vizcaíno et al. 2010, Goin 1997, Clemens 1979 | |||||
Age range: base of the Early/Lower Eocene to the top of the Montehermosan or 55.80000 to 3.80000 Ma
Collections (77 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Ypresian | Argentina (Chubut) | Patene indet. (133040) | |
| Early/Lower Eocene | Brazil (Rio de Janeiro) | Hathlyacynidae indet., Ischyrodidelphis castelllanosi, Patene simpsoni (118314) | |
| Early/Lower Eocene | France (Marne) | Protoproviverra palaeonictides (181633) | |
| Mustersan | Argentina (Chubut) | Procladosictis anomala (176286) | |
| Lutetian | Argentina (Salta) | Patene simpsoni (142855 211419) | |
| Lutetian | Argentina (Chubut) | Hathliacynidae Gen. et sp. nov. 6 (176059) Hathliacynidae gen. et sp. nov. 6 (133039) Patene sp. (44956) | |
| Middle Eocene - Early/Lower Miocene | Argentina (Chubut) | Cladosictis sp. (176461) | |
| Bartonian | Argentina (Chubut) | Patene coluapiensis (176186) Patene coluapiensis, Pseudocladosictis determinabile (13779) Procladosictis erecta (176163) | |
| Tinguirirican | Argentina (Chubut) | Hathliacynidae indet. (176200) | |
| Rupelian | Peru (Departamento de Ucayali) | Patene campbelli (149523) | |
| Deseadan | Bolivia | Sallacyon hoffstetteri, Notogale mitis, Andinogale sallensis (95688) | |
| Deseadan | Argentina (Chubut) | Pharsophorus mitis, Notogale mitis (14211) | |
| Deseadan | Argentina (Santa Cruz) | Sallacyon hoffstetteri (50074) | |
| Deseadan | Bolivia (La Paz) | Notogale mitis (14208 14209 133615) Sallacyon hoffstetteri (133596) | |
| Early/Lower Miocene | Argentina (Chubut) | Sipalocyon gracilis, Sipalocyon externus (236575) | |
| Colhuehuapian | Argentina (Río Negro) | Cladosictis patagonica (89093) | |
| Colhuehuapian | Argentina (Chubut) | Cladosictis sp., Sipalocyon sp., Hathliacynidae indet., Cladosictis centralis, Sipalocyon externus, Anatherium herrerae (141368) | |
| Colhuehuapian | Argentina (Neuquén) | Cladosictis sp., Sipalocyon sp., Hathliacynidae indet. (89029) | |
| Burdigalian | Chile (Aisén) | Cladosictis sp. (71843) | |
| Burdigalian - Langhian | Peru (Cuzco) | Sipalocyon sp. (144513) | |
| Santacrucian | Argentina (Santa Cruz) | Cladosictis patagonica (142529 176567 176572 176576 176584 176598 176635 176636 210153) Cladosictis patagonica, Hathliacynus lustratus, Hathliacynus cultridens, Hathliacynus rollieri, Amphithereutes obscurus, Acyon bardus (176575) Cladosictis patagonicus, Cladosictis patagonica, Hathlyacynidae indet. (176578) Hathliacynus kobyi, Sipalocyon gracilis, Thylacodictis exilis, Agustylus primaevus, Hathliacynus fischeri, Cladosictis dissimilis, Acyon tricuspidatus, Pseudonotictis pusillus (176563) Hathlyacynidae indet. (176571) Protoproviverra manzaniana, Sipalocyon gracilis, Cladosictis patagonica, Ictioborus destructor (176594) Proviverra trouessarti (176596) Sipalocyon gracilis (87314 176582 210637 210786 210787) Sipalocyon gracilis, Amphiproviverra crassa, Cladosictis patagonica, Cladosictis lateralis, Perathereutes pungens (142533) Sipalocyon gracilis, Cladosictis patagonica (176569 176581 176590) Sipalocyon gracilis, Cladosictis patagonica, Acyon tricuspidatus (27313) Sipalocyon gracilis, Cladosictis patagonica, Cladosictis petersoni (176585) Sipalocyon gracilis, Cladosictis patagonica, Perathereutes pungens (210638) Sipalocyon gracilis, Hathliacynus lynchi, Agustylus cynoides, Cladosictis patagonica, Acyon tricuspidatus (176593) Sipalocyon gracilis, Sipalocyon curtus (176566) Sipalocyon gracilis, Sipalocyon mixtus, Sipalocyon altiramis, Sipalocyon longus, Cladosictis patagonica, Abderites altiramis (176565) Sipalocyon pusillus, Sipalocyon gracilis, Perathereutes amputans, Perathereutes obtusus, Amphiproviverra minuta, Cladosictis patagonica, Agustylus minus, Ictioborus fenestratus, Protoproviverra ensidens, Perathereutes pungens, Protoproviverra obusta, Anatherium defossus (176570) Sipalocyon sp., Sipalocyon gracilis, Cladosictis patagonica, Hathlyacyninae indet. (176599) Sipalocyon sp., Sipalocyon gracilis, Cladosictis patagonica, Perathereutes pungens (176602) | |
| Santacrucian - Friasian | Chile (Bío Bío) | Sipalocyon sp. (140107) | |
| Friasian | Chile (Aisén) | Sipalocyon gracilis, Cladosictis patagonica (139845) | |
| Langhian | Argentina (Rio Negro) | Cladosictis patagonica (198569) | |
| Middle Miocene | Argentina (Chubut) | Pseudonotictis chubutensis (186561) | |
| Laventan | Bolivia (Tarija) | Acyon myctoderos (38071) | |
| Tortonian | Argentina (Entre Rios) | Notictis ortizi (13626) | |
| Tortonian | Bolivia (La Paz) | Borhyaenidium altiplanicus (90023) | |
| Chasicoan | Argentina (Buenos Aires) | Chasicostylus castroi (79180) | |
| Chasicoan | Argentina (San Juan) | Simpsonia borhyaenoides (146533) | |
| Huayquerian | Argentina (Buenos Aires) | Borhyaenidium sp. (61298) | |
| Huayquerian | Argentina (La Pampa) | Borhyaenidium musteloides (87198) | |
| Huayquerian | Argentina (Entre Rios) | Notictis ortizi (55600) | |
| Montehermosan | Argentina (Catamarca) | Borhyaenidium riggsi (176757) | |
| Montehermosan | Argentina (Buenos Aires) | Notocynus hermosicus (13503 198863) Notocynus sp. (199153) |