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Marchantiophytina (liverwort)

Taxonomy
Marchantiophyta was named as a phylum. It is extant.

It was reranked as the subphylum Marchantiophytina by Taylor et al. (2008).

It was assigned to Marchantiomorpha by Kenrick and Crane (1997); and to Bryophyta by Taylor et al. (2008).

Synonymy list
YearName and author
1997Marchantiophyta Kenrick and Crane
2008Marchantiophytina Taylor et al. p. 1028

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RankNameAuthor
kingdomPlantae
phylumBryophyta
RankNameAuthor
subphylumMarchantiophytina
subphylumMarchantiophytina

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subph. Marchantiophytina [liverwort]
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G. †Discites Harris 1931
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†Discites minutus Harris 1931
G. †Eohepatica Heard and Jones 1931
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†Eohepatica dyfriensis Heard and Jones 1931
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G. Fuscocephaloziopsis Fulford 1968
G. †Cheirorhiza Krasilov 1970
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†Cheirorhiza brittae Krasilov 1970
G. †Chrysotheca Miner 1935
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†Chrysotheca diskoensis Miner 1935
G. †Diettertia Brown and Robison 1974
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†Diettertia montanensis Brown and Robison 1974
G. †Laticaulina Krasilov 1970
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†Laticaulina papillosa Krasilov 1970
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G. Plagiochila Dumortier 1831
Or. Porellales Schljakov 1972
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Fm. Frullaniaceae Ordway 1881
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G. Frullania Raddi 1818
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†Frullania palaeoafricana Bouju et al. 2022
†Frullania shewanensis Bouju et al. 2022
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G. Cololejeunea Spruce 1884
G. Lejeunea Libert 1820
G. Leptolejeunea Spruce 1884
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G. Spruceanthus Verdoorn 1934
G. Thysananthus Lindenberg 1844
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G. Radula Dumortier 1822
G. †Jungermanniopsis Howe and Hollick 1922
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†Jungermanniopsis cockerellii Howe and Hollick 1922
G. †Jungermannites Göppert 1845
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†Jungermannites neesianus Goeppert 1845
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Fm. Marchantiidae Engler 1893
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G. †Costatheca Hall 1967
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†Costatheca lata Vangerow 1954
†Costatheca striata Dijkstra 1949
G. †Khasurtya Mamontov 2020
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†Khasurtya ginkgoides Mamontov and Ignatov 2020
G. †Khasurtythallus Mamontov 2019
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G. †Stoverisporites Norvick and Burger 1975
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†Stoverisporites lunaris Norvick and Burger 1975
G. †Triporoletes Mtchedlishvili 1960
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†Triporoletes asper Srivastava 1972
†Triporoletes blanensis Srivastava 1975
†Triporoletes cenomanianus Srivastava 1977
†Triporoletes laevigatus Playford 1971
†Triporoletes radiatus Playford 1971
†Triporoletes reticulatus Playford 1971
†Triporoletes simplex Playford 1971
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G. †Metzgeriites Steere 1946
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†Metzgeriites glebosus Harris 1931
†Metzgeriites montanensis Lapasha and Miller 1985
Or. Sphaerocarpales Cavers 1910
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G. †Aequitriradites Delcourt and Sprumont 1955
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†Aequitriradites acusus Dettmann 1963
†Aequitriradites baculatus Döring 1964
†Aequitriradites dubius Delcourt and Sprumont 1955
†Aequitriradites fusus Singh et al. 1964
†Aequitriradites hispidus Dettmann and Playford 1968
†Aequitriradites indicus Singh et al. 1964
†Aequitriradites longispinus Archangelsky and Archangelsky 2005
†Aequitriradites plicatus Archangelsky and Archangelsky 2005
†Aequitriradites spinulosus Cookson and Dettmann 1958
†Aequitriradites superspinulosus Archangelsky et al. 2008
†Aequitriradites triangulatus Singh et al. 1964
†Aequitriradites verrucosus Cookson and Dettmann 1958
Invalid names: Calobryales [empty], Marchantiophyceae [replaced], Monocleales [empty]
G. †Schizolepidella Halle 1913
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G. †Thallomia Heard and Jones 1931
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†Thallomia llandyfriensis Heard and Jones 1931
Invalid names: Treubiopsida [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
No ecological data are available

Age range

Maximum range based only on fossils: base of the Anisian to the top of the Langhian or 246.70000 to 13.82000 Ma
Minimum age of oldest fossil (stem group age): 241.464 Ma

Collections (247 total)


Time interval Ma Country or state Original ID and collection number
Anisian246.7 - 241.464Israel Aequitriradites minor (31494)
Early/Lower Jurassic201.4 - 174.7Antarctica Schizolepidella sp. 1 (29347)
Middle Jurassic - Late/Upper Cenomanian174.7 - 93.9Egypt (Western Desert) Aequitriradites verrucosus, Aequitriradites spinulosus (139303)
Bathonian - Oxfordian168.2 - 154.8Egypt (Western Desert) Aequitriradites spinulosus (139310 139311 139312 139313 139315 139316 139317)
Callovian - Oxfordian165.3 - 154.8Russian Federation Cheirorhiza sp. 1, Laticaulina sp. 1 (30671)
Late/Upper Jurassic - Early/Lower Cretaceous161.5 - 100.5India (Gujarat) Aequitriradites sp. (224625)
Early/Lower Berriasian143.1 - 141.0Denmark (Bornholm) Aequitriradites spinulosus (83134)
Neocomian - Barremian143.1 - 121.4Sudan (South Kurdufan) Aequitriradites acusus (172845 172847 172850 172851 172852 172853 172857 172860 172861 172864 172865)
Early/Lower Cretaceous143.1 - 100.5Argentina (San Luis) Triporoletes reticulatus (151090)
Early/Lower Cretaceous143.1 - 100.5India (Gujarat) Aequitriradites fusus, Aequitriradites triangulatus (224620) Aequitriradites verrucosus (32379)
Cretaceous143.1 - 66.0Netherlands (South Limburg) Chrysotheca striata (31831) Chrysotheca tenuis, Chrysotheca quadriplicata, Chrysotheca dentata, Chrysotheca striata (31829) Chrysotheca tenuis, Chrysotheca quadriplicata, Chrysotheca striata (31832)
Cretaceous143.1 - 66.0Greenland Chrysotheca diskoensis (32492)
Cretaceous143.1 - 66.0Netherlands Chrysotheca subreticulata (32718)
Early/Lower Cretaceous - Late/Upper Cretaceous143.1 - 66.0Antarctica (Antarctic peninsula) Triporoletes reticulatus (172746)
Late/Upper Berriasian - Early/Lower Valanginian139.1 - 132.6United Kingdom (England) Triporoletes sp. (32353)
Hauterivian132.6 - 125.77Portugal Aequitriradites spinulosus (73484) Aequitriradites spinulosus, Aequitriradites dubius (73496)
Hauterivian132.6 - 125.77Egypt (Western Desert) Aequitriradites spinulosus (139320 139322) Aequitriradites verrucosus (139323)
Late/Upper Hauterivian132.6 - 125.77Argentina (Neuquén) Aequitriradites sp. (150584)
Barremian125.77 - 121.4Egypt (El Wadi El Gadid) Aequitriradites spp. (139497 139498 139501) Aequitriradites spp., Aequitriradites spinulosus (139500) Triporoletes reticulatus, Aequitriradites spp. (139495 139496) Triporoletes reticulatus, Aequitriradites spp., Aequitriradites spinulosus, Aequitriradites acusus (139493 139499)
Barremian125.77 - 121.4Portugal Aequitriradites baculatus (73324) Aequitriradites spinulosus (73498)
Early/Lower Barremian125.77 - 121.4United Kingdom (England) Triporoletes reticulatus, Aequitriradites verrucosus (45193)
Barremian - Early/Lower Aptian125.77 - 119.57China (Nei Mongol) Stoverisporites sp. (46976)
Late/Upper Barremian - Early/Lower Aptian125.77 - 119.57Argentina (Neuquén) Triporoletes reticulatus (149818)
Late/Upper Barremian - Early/Lower Aptian125.77 - 119.57Germany (Nordrhein-Westphalen) Costatheca lata (22727)
Barremian - Aptian125.77 - 113.2Sudan (West Kurdufan) Aequitriradites acusus (145178 145179 145180)
Barremian - Aptian125.77 - 113.2Sudan (South Kurdufan) Aequitriradites acusus (172838 172839 172842 172844)
Barremian - Albian125.77 - 100.5China (Guizhou) Triporoletes sp. (182652)
Aptian121.4 - 113.2Portugal Aequitriradites spinulosus (73577) Aequitriradites spinulosus, Aequitriradites dubius (73582)
Aptian121.4 - 113.2Russian Federation (Buryatia) Khasurtythallus monosolenioides, Khasurtya ginkgoides (217431)
Aptian121.4 - 113.2Argentina (Santa Cruz) Aequitriradites verrucosus, Aequitriradites baculatus, Aequitriradites spinulosus (150335) Stoverisporites lunaris (149982)
Aptian121.4 - 113.2Canada (Alberta) Aequitriradites verrucosus (194481)
Aptian - Early/Lower Albian121.4 - 110.1Argentina (Santa Cruz) Aequitriradites superspinulosus (149849) Aequitriradites superspinulosus, Aequitriradites verrucosus (149850)
Aptian - Albian121.4 - 100.5Sudan (West Kurdufan) Aequitriradites acusus (145162 145163 145164 145165 145166 145170 145172)
Aptian - Albian121.4 - 100.5Argentina (Santa Cruz) Triporoletes reticulatus, Aequitriradites baculatus, Aequitriradites spinulosus (150615)
Aptian - Albian121.4 - 100.5Sudan (South Kurdufan) Aequitriradites acusus (145154 145155 145156 145161)
Aptian - Cenomanian121.4 - 93.9Libya (Cyrenaica) Aequitriradites sp. (142508)
Late/Upper Aptian119.57 - 113.2USA (Montana) Diettertia montanensis (21126 21130 21136 21139 21140 21153 21154 21155 21156) Metzgeriites montanensis, Diettertia montanensis (21151)
Late/Upper Aptian - Early/Lower Albian119.57 - 110.1India (Gujarat) Aequitriradites indicus (32378)
Albian - Aptian113.2 - 113.2Cote D'Ivoire (Abidjan) Aequitriradites sp. (137356)
Early/Lower Albian113.2 - 110.1Argentina (Santa Cruz) Triporoletes reticulatus (149846)
Early/Lower Albian - Middle Albian113.2 - 106.3Argentina (Santa Cruz) Triporoletes reticulatus, Aequitriradites superspinulosus (149746)
Albian113.2 - 100.5Canada (Nunavut) Costatheca diskoensis (32593)
Albian113.2 - 100.5Antarctica Triporoletes reticulatus (153431 153432)
Albian113.2 - 100.5Australia (South Australia) Aequitriradites spinolosus, Aequitriradites verrucosus (164488 164530) Aequitriradites spinulosus (164523) Aequitriradites verrucosus (164524 164529) Aequitriradites verrucosus, Aequitriradites spinulosus (164489 164490 164522 164525 164526 164528)
Albian - Cenomanian113.2 - 93.9Antarctica Triporoletes reticulatus (153422)
Albian - Cenomanian113.2 - 93.9Sudan (Sotuh Kurdufan) Aequitriradites acusus (145152)
Early/Lower Albian - Early/Lower Cenomanian113.2 - 93.9Argentina (Santa Cruz) Triporoletes reticulatus, Aequitriradites sp. 1, Aequitriradites sp. 2, Aequitriradites superspinulosus, Aequitriradites verrucosus, Aequitriradites baculatus, Aequitriradites spinulosus (149928)
Albian - Turonian113.2 - 89.8Australia (Victoria) Triporoletes reticulatus (163583)
Middle Albian - Late/Upper Albian110.1 - 100.5Antarctica Stoverisporites microverrucatus (153430)
Middle Albian - Late/Upper Albian110.1 - 100.5Canada (Nunavut) Costatheca diskoensis (32590)
Middle Albian - Late/Upper Albian110.1 - 100.5Australia (South Australia) Aequitriradites verrucosus, Aequitriradites spinulosus (163902)
Middle Albian - Late/Upper Albian110.1 - 100.5Morocco (Tarfaya) Triporoletes reticulatus (144143)
Middle Albian - Early/Lower Cenomanian110.1 - 93.9Egypt (Western Desert) Aequitriradites spinulosus (144955)
Late/Upper Albian - Early/Lower Cenomanian106.3 - 93.9Senegal (Dakar) Aequitriradites sp. (137360)
Late/Upper Albian - Early/Lower Cenomanian106.3 - 93.9Egypt (Western Desert) Aequitriradites spinulosus (139325) Aequitriradites verrucosus, Aequitriradites spinulosus (139332)
Cenomanian100.5 - 93.9USA (Minnesota) Costatheca sp. (31943 31945 31950 31951)
Cenomanian100.5 - 93.9USA (New Jersey) Aequitriradites insolitus (34114) Aequitriradites insolitus, Aequitriradites spinulosus (34141)
Cenomanian100.5 - 93.9USA (Kansas) Costatheca diskoensis (32607 32609 32638 32643 32644 32646 32647 32648 32652 32654 32677 32679 32681 32685 32687 32693) Costatheca lata (32680 32682 32692)
Cenomanian100.5 - 93.9Australia (Northern Territory) Stoverisporites micriverrucatus (163482) Stoverisporites microverrucatus (163508 163510 163513) Stoverisporites microverrucatus, Triporoletes laevigatus (163481 163506 163509 163511 163512) Triporoletes reticulatus, Stoverisporites microverrucatus (163507) Triporoletes reticulatus, Stoverisporites microverrucatus, Triporoletes laevigatus (163483 163484 163505 163514 163515)
Cenomanian100.5 - 93.9Cote D'Ivoire (Abidjan) Aequitriradites sp. (137355)
Cenomanian100.5 - 93.9USA (Iowa) Chrysotheca dakotaensis (32479) Chrysotheca levis, Chrysotheca dakotaensis, Chrysotheca diskoensis (32524)
Late/Upper Cenomanian100.5 - 93.9Argentina (Mendoza) Triporoletes reticulatus (149812)
Middle Cenomanian100.5 - 93.9South Sudan Aequitriradites acusus (200009)
Cenomanian - Cretaceous100.5 - 66.0USA (Kansas) Costatheca diskoensis (32655)
Late/Upper Cretaceous - Miocene100.5 - 5.333China (Xingjian) Triporoletes sp. (140039)
Turonian93.9 - 89.8Australia (Northern Territory) Triporoletes reticulatus, Stoverisporites microverrucatus, Triporoletes laevigatus (163819)
Turonian - Maastrichtian93.9 - 66.0Cote D'Ivoire (Abidjan) Aequitriradites sp. (137357)
Coniacian - Early/Lower Santonian89.8 - 83.6Antarctica Stoverisporites microverrucatus (169331 169332 169334 169335 169337 169338 169339 169340 169341 169342 169343 169344 169346 169348 169349 169351 169352)
Coniacian - Maastrichtian89.8 - 66.0Argentina (Chubut) Zlivisporis reticulatus (233992)
Santonian85.7 - 83.6Netherlands (Limburg) Costatheca dentata, Costatheca tenuis, Costatheca striata (31798) Costatheca quadriplicata (31780)
Santonian - Campanian85.7 - 72.2Antarctica Aequitriradites spinulosus, Triporoletes simplex (150509) Stoverisporites microverrucatus (169354)
Campanian83.6 - 72.2Canada (Alberta) Aequitriradites spinulosus (207022) Costatheca dakotaensis (33036) Costatheca dentata (33006 33011) Costatheca diskoensis (33030) Costatheca diskoensis, Costatheca striata (33000) Costatheca elongata, Costatheca sp. (33023) Costatheca striata (33035)
Campanian83.6 - 72.2Antarctica Triporoletes reticulatus (153425)
Late/Upper Campanian83.6 - 72.2Canada (Alberta) Aequitriradites ornatus (22655) Aequitriradites ornatus, Aequitriradites spinulosus (36264)
Late/Upper Campanian83.6 - 72.2Antarctica Aequitriradites sp. (150182)
Campanian - Maastrichtian83.6 - 66.0Antarctica Stoverisporites microverrucatus (153427)
Campanian - Maastrichtian83.6 - 66.0Canada (Nunavut) Costatheca dakotaensis (32603)
Late/Upper Campanian - Early/Lower Maastrichtian83.6 - 66.0Argentina (Mendoza) Triporoletes asper (168549)
Late/Upper Campanian - Early/Lower Maastrichtian83.6 - 66.0Argentina (Neuquén) Triporoletes reticulatus (150621)
Late/Upper Campanian - Early/Lower Maastrichtian83.6 - 66.0Antarctica Aequitriradites spinulosus (150237)
Late/Upper Campanian - Maastrichtian83.6 - 66.0Antarctica (Seymour Island) Triporoletes reticulatus (167585 167586)
Late/Upper Campanian - Danian83.6 - 61.66Antarctica Triporoletes reticulatus (167588)
Early/Lower Maastrichtian72.2 - 66.0Argentina (Santa Cruz) Stoverisporites microverrucatus (231750) Triporoletes reticulatus (231736 231737 231742 231743 231744 231746 240581 240582 240583) Triporoletes reticulatus, Costatheca striata (231725) Triporoletes sp. (231735)
Late/Upper Maastrichtian72.2 - 66.0Argentina (Chubut) Zlivisporis reticulatus (234014)
Maastrichtian72.2 - 66.0Canada (Nunavut) Costatheca dakotaensis, Costatheca diskoensis, Costatheca elongata (32600) Costatheca dakotaensis, Costatheca tenuis, Costatheca diskoensis, Costatheca levis (32601) Costatheca dakotaensis, Costatheca tenuis, Costatheca levis (32602)
Maastrichtian72.2 - 66.0Antarctica Triporoletes reticulatus (153424)
Maastrichtian72.2 - 66.0Argentina (Santa Cruz) Triporoletes reticulatus (234760)
Maastrichtian72.2 - 66.0Canada (Alberta) Costatheca dakotaensis (33056) Costatheca elongata (33076) Costatheca tenuis (33058) Costatheca verrucata (33079 33080)
Late/Upper Maastrichtian - Early/Lower Paleocene72.2 - 61.66Canada (Alberta) Costatheca tenuis, Costatheca striata (33085)
Danian66.0 - 61.66Argentina (Chubut) Aequitriradites spinulosus (152264)
Early/Lower Eocene56.0 - 48.07Southern Ocean Triporoletes reticulatus (169272 169274)
MP 1348.07 - 41.03Germany (Eifel) Frullania sp. (24201)
Chadronian37.0 - 33.9USA (Colorado) Jungermanniopsis cockerellii (24209)
Early/Lower Oligocene - Early/Lower Miocene33.9 - 15.98Australia (Victoria) Jungermanniales indet. (169007)
Early/Lower Miocene23.04 - 15.98Ethiopia Thysananthus aethiopicus (227286) Thysananthus aethiopicus, Lejeunea abyssinicoides, Frullania shewanensis, Frullania palaeoafricana (224753)
Langhian15.98 - 13.82China (Fujian) Cololejeunea sp., Leptolejeunea sp., Spruceanthus sp., Radula sp., Plagiochila sp., Fuscocephaloziopsis sp. (220273)