PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Leptoceratopsidae (leptoceratopsid)

Osteichthyes - Leptoceratopsidae

Taxonomy

Leptoceratopsinae was named by Nopcsa (1923). Its type is Leptoceratops. It was considered monophyletic by Ryan et al. (2023).

It was reranked as the family Leptoceratopsidae by Makovicky (2001), You and Dodson (2003), Chinnery (2004), Chinnery and Horner (2007), Butler et al. (2008), Han et al. (2012), Farke et al. (2014), Zheng et al. (2015), Morschhauser et al. (2019), Arbour and Evans (2019), Ryan et al. (2023).

It was assigned to Ceratopsidae by Nopcsa (1923), Nopcsa (1928), Nopcsa (1929), Huene (1934), Huene (1934); to Ceratopsoidea by You and Dodson (2003); to Coronosauria by Morschhauser et al. (2019), Arbour and Evans (2019); and to Neoceratopsia by Makovicky (2001), Chinnery (2004), Chinnery and Horner (2007), Butler et al. (2008), Han et al. (2012), Farke et al. (2014), Zheng et al. (2015), Ryan et al. (2023).

Subtaxa

Cerasinops, Ferrisaurus, Gremlin, Gryphoceratops, Helioceratops, Ischioceratops, Koreaceratops, Leptoceratops (type genus), Montanoceratops, Prenoceratops, Udanoceratops, Unescoceratops, Zhuchengceratops

Synonymy list

Year	Name and author
1923	Leptoceratopsinae Nopcsa p. 126
1928	Leptoceratopsinae Nopcsa p. 185
1929	Leptoceratopsinae Nopcsa p. 71
1934	Leptoceratopsinae Huene p. 15
2001	Leptoceratopsidae Makovicky p. 254
2003	Leptoceratopsidae You and Dodson
2004	Leptoceratopsidae Chinnery pp. 584-585 fig. 9
2007	Leptoceratopsidae Chinnery and Horner p. 638 fig. 6
2008	Leptoceratopsidae Butler et al. p. 22 figs. 2-4
2012	Leptoceratopsidae Han et al. p. 1391 fig. 14
2014	Leptoceratopsidae Farke et al.
2015	Leptoceratopsidae Zheng et al. p. 6 fig. 3
2019	Leptoceratopsidae Arbour and Evans
2019	Leptoceratopsidae Morschhauser et al. p. 129 fig. 10
2023	Leptoceratopsidae Ryan et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
class	Osteichthyes
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()

Rank	Name	Author
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Cerapoda
—	Marginocephalia
—	Ceratopsia	()
infraorder	Neoceratopsia
family	Leptoceratopsidae	()
family	Leptoceratopsidae	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Leptoceratopsidae Nopcsa 1923 [leptoceratopsid]

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G. †Cerasinops Chinnery and Horner 2007

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†Cerasinops hodgskissi Chinnery and Horner 2007

G. †Ferrisaurus Arbour and Evans 2019

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†Ferrisaurus sustutensis Arbour and Evans 2019

G. †Gremlin Ryan et al. 2023

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†Gremlin slobodorum Ryan et al. 2023

G. †Gryphoceratops Ryan et al. 2012

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†Gryphoceratops morrisoni Ryan et al. 2012

G. †Helioceratops Jin et al. 2009

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†Helioceratops brachygnathus Jin et al. 2009

G. †Ischioceratops He et al. 2015

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†Ischioceratops zhuchengensis He et al. 2015

G. †Koreaceratops Lee et al. 2010

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†Koreaceratops hwaseongensis Lee et al. 2010

G. †Leptoceratops Brown 1914

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†Leptoceratops gracilis Brown 1914

G. †Montanoceratops Sternberg 1951

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†Montanoceratops cerorhynchus Brown and Schlaikjer 1942

G. †Prenoceratops Chinnery 2004

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†Prenoceratops pieganensis Chinnery 2004

G. †Udanoceratops Kurzanov 1992

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†Udanoceratops tschizhovi Kurzanov 1992

G. †Unescoceratops Ryan et al. 2012

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†Unescoceratops koppelhusae Ryan et al. 2012

G. †Zhuchengceratops Xu et al. 2010

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†Zhuchengceratops inexpectus Xu et al. 2010

Diagnosis

Reference	Diagnosis
B. F. Nopcsa 1928	no horns; fringe well developed, mandible robust

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Maximum body mass:

1000 kgg

Minimum body mass:

100 kgg

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

gregariouso

Environment:

terrestrialsubo

Locomotion:

actively mobileo

Life habit:

ground dwellingsubo

Diet:

herbivoresubo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2005-09-15 11:08:09

Modified:

2005-09-15 13:08:09

Source: g = genus, subo = suborder, o = order

References: Marsh 1875, Peczkis 1995, Kiessling 2004, Norman and Weishampel 1991

Age range: base of the Albian to the top of the Early/Lower Maastrichtian or 113.20000 to 66.00000 Ma

Collections (29 total)

Time interval	Ma	Country or state	Original ID and collection number
Aptian - Albian	121.4 - 100.5	China (Jilin)	Neoceratopsia indet. (72321)
Albian	113.2 - 100.5	South Korea (Gyeonggi-do)	Koreaceratops hwaseongensis (103988)
Late/Upper Santonian	85.7 - 83.6	Canada (Alberta)	Gryphoceratops morrisoni (125521)
Campanian	83.6 - 72.2	China (Shandong)	Zhuchengceratops inexpectus, Ischioceratops zhuchengensis (91473)
Early/Lower Campanian	83.6 - 72.2	Sweden (Skåne)	Leptoceratopsidae indet. (74334 74335)
Early/Lower Campanian - Middle Campanian	83.6 - 72.2	USA (North Carolina)	Leptoceratopsidae indet. (179206)
Late/Upper Campanian	83.6 - 72.2	Canada (Alberta)	Prenoceratops sp. (52991)
Late/Upper Campanian	83.6 - 72.2	USA (Montana)	Neoceratopsia indet. (69931)
Late/Upper Campanian	83.6 - 72.2	Mongolia (Omnogov)	Udanoceratops tschizhovi (65344)
Middle Campanian	83.6 - 72.2	Canada (Alberta)	Gremlin slobodorum (45670) Protoceratopsidae indet. (60173)
Middle Campanian	83.6 - 72.2	USA (Montana)	Cerasinops hodgskissi (100851)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	Mongolia (Dornogov)	Udanoceratops tschizhovi (65338)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	USA (Montana)	Ceratopsia indet. (53999) Leptoceratops sp. (54000)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Canada (Alberta)	Ceratopsia indet. (51965)
Early/Lower Maastrichtian	72.2 - 66.0	USA (Montana)	Ceratopsia indet. (55266) Montanoceratops cerorhynchus (55267)
Late/Upper Maastrichtian	72.2 - 66.0	Canada (Alberta)	Ceratopsia indet. (64628) Leptoceratops gracilis (64353) Leptoceratops sp. (14569) Montanoceratops sp. (64791)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Montana)	Leptoceratops gracilis (73947 73948)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Wyoming)	Leptoceratops gracilis (34747) Leptoceratops sp. (53824)
Late/Upper Maastrichtian	72.2 - 66.0	USA (South Dakota)	Leptoceratops sp. (224484)
Maastrichtian	72.2 - 66.0	Canada (British Columbia)	Cerapoda indet. (84008)