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Euoplocephalus tutus

Reptilia - Ankylosauridae

Taxonomy
Stereocephalus tutus was named by Lambe (1902). Its type specimen is NMC 210, a skull (Skull roof of the antorbital region, and a partial first cervical half ring), and it is a 3D body fossil. Its type locality is near mouth of Berry Creek (NMC), which is in a Campanian terrestrial horizon in the Dinosaur Park Formation of Canada. It was considered monophyletic by Vickaryous et al. (2004).

It was recombined as Ankylosaurus tutus by Brown (1914), Kuhn (1964); it was recombined as Palaeoscincus tutus by Hennig (1915); it was recombined as Euoplocephalus tutus by Lambe (1910), Lambe (1914), Lambe (1915), Lambe (1919), Gilmore (1923), Parks (1924), Huene (1927), Sternberg (1928), Hay (1930), Russell (1930), Russell (1964), Russell (1966), Coombs (1978), Sereno (1986), Coombs (1986), Tumanova (1987), Vickaryous and Ryan (1997), Eberth (1997), Currie (2000), Ford (2000), Eberth et al. (2001), Carpenter (2001), Barrett (2001), Ford and Kirkland (2001), Garcia and Pereda-Suberbiola (2003), Vickaryous et al. (2004), Ryan and Evans (2005), Currie (2005), Sullivan and Fowler (2006), Vickaryous (2006), Arbour (2007), Longrich (2008), Burns (2008), Osi and Makádi (2009), Carpenter et al. (2011), Burns and Sullivan (2011), Burns and Sullivan (2011), Arbour et al. (2012), Penkalski and Blows (2013), Arbour and Currie (2013), Arbour and Currie (2013), Kirkland et al. (2013), Penkalski (2014), Arbour et al. (2014), Blows (2014), Arbour et al. (2014), Han et al. (2014), Hill et al. (2015), Arbour and Currie (2016), Borinder et al. (2016), Kineer et al. (2016), Herne et al. (2018), Penkalski (2018), Sakagami and Kawabe (2020).

Synonymy list
YearName and author
1902Stereocephalus tutus Lambe p. 55 figs. Pl. 10, 11, 21
1905Stereocephalus tutus Lambe p. 23
1908Stereocephalus tutus Brown p. 197
1910Euoplocephalus tutus Lambe p. 151
1914Ankylosaurus tutus Brown p. 378
1914Euoplocephalus tutus Lambe p. 132
1915Palaeoscincus tutus Hennig p. 14
1915Euoplocephalus tutus Lambe p. 117
1917Stereocephalus tutus Sternberg p. 210
1919Euoplocephalus tutus Lambe p. 39
1921Stereocephalus tutus Gilmore p. 589
1923Euoplocephalus tutus Gilmore pp. 47-48
1924Euoplocephalus tutus Parks p. 7
1927Euoplocephalus tutus Huene p. 261
1928Euoplocephalus tutus Sternberg p. 100
1930Euoplocephalus tutus Hay p. 238
1930Euoplocephalus tutus Russell p. 149
1964Ankylosaurus tutus Kuhn p. 45
1964Euoplocephalus tutus Russell p. 13
1965Europlocephalus tutus Simmons p. 70
1966Euoplocephalus tutus Russell p. 19
1978Euoplocephalus tutus Coombs p. 148 fig. 3
1986Euoplocephalus tutus Coombs, Jr. p. 170
1986Euoplocephalus tutus Sereno
1987Euoplocephalus tutus Tumanova p. 15
1997Euoplocephalus tutus Eberth p. 201
1997Euoplocephalus tutus Vickaryous and Ryan p. 490
2000Euoplocephalus tutus Currie p. 277
2000Euoplocephalus tutus Ford p. 175
2001Euoplocephalus tutus Barrett p. 40 fig. 2.8
2001Euoplocephalus tutus Carpenter p. 457
2001Euoplocephalus tutus Eberth et al. p. 58
2001Euoplocephalus tutus Ford and Kirkland p. 254
2003Euoplocephalus tutus Garcia and Pereda-Suberbiola p. 161
2004Euoplocephalus tutus Vickaryous et al. p. 365
2005Euoplocephalus tutus Currie p. 5
2005Euoplocephalus tutus Ryan and Evans p. 315
2006Euoplocephalus tutus Sullivan and Fowler p. 259
2006Euoplocephalus tutus Vickaryous p. 1011
2007Euoplocephalus tutus Arbour p. 41A
2008Euoplocephalus tutus Burns p. 1107
2008Euoplocephalus tutus Longrich p. 994
2009Euoplocephalus tutus Osi and Makádi p. 235 fig. 6
2011Euoplocephalus tutus Burns and Sullivan p. 177
2011Euoplocephalus tutus Carpenter et al. p. 5
2012Euoplocephalus tutus Arbour et al. p. 55
2013Euoplocephalus tutus Arbour and Currie p. 27
2013Euoplocephalus tutus Kirkland et al. p. 7
2013Euoplocephalus tutus Penkalski and Blows p. 178
2014Euoplocephalus tutus Arbour et al. p. 2
2014Euoplocephalus tutus Arbour et al. p. 634
2014Euoplocephalus tutus Blows p. 61 fig. 5
2014Euoplocephalus tutus Han et al. p. 12 fig. 7
2014Euoplocephalus tutus Penkalski p. 617
2015Euoplocephalus tutus Hill et al. p. 900
2016Euoplocephalus tutus Arbour and Currie p. 418–419
2016Euoplocephalus tutus Borinder et al. p. 66
2016Euoplocephalus tutus Kineer et al. p. 42
2018Euoplocephalus tutus Herne et al. p. 54
2018Euoplocephalus tutus Penkalski p. 264
2020Euoplocephalus tutus Sakagami and Kawabe p. 12

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Eugnathostomata
—Teleostomi(Bonaparte 1836)
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
subclassEureptilia()
RankNameAuthor
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Ornithischia()
—Genasauria
—Thyreophora()
—Ankylosauria()
familyAnkylosauridae
subfamilyAnkylosaurinaeNopcsa 1918
tribeEuoplocephalini
genusEuoplocephalus
speciestutus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Euoplocephalus tutus Lambe 1902
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Diagnosis
ReferenceDiagnosis
T. A. Tumanova 1987Premaxillary beak very wide, anterior blunt, width equal to or exceeding space between rear [maxillary] teeth; maxillary shelves well developed; plane surface of palatal bone tilted laterally; anterior wall of medial section of pterygoids slanted forward; paroccipital processes laterally oriented; occipital condyle broad oval, oriented posteroventrally; quadrate and paroccipital processes not fused; occipital plane tilted slightly forward.
V. M. Arbour and P. J. Currie 2013Differs from Anodontosaurus lambei and Scolosaurus cutleri in lacking subcircular caputegulae at the bases of the quadratojugal and squamosal horns (postocular caputegulae); differs from Anodontosaurus lambei in lacking interstitial osteoderms at the bases of the primary osteoderms of the first cervical half ring, and in having semicircular major osteoderms in dorsal view on the tail club; differs from Dyoplosaurus acutosquameus in having laterally-directed sacral ribs; differs from Scolosaurus cutleri in having oval to subcircular-based keeled medial and lateral primary half ring osteoderms and in having a proportionately shorter postacetabular process of the ilium; differs from Ankylosaurus magniventris in having anteriorly-directed nares, and in lacking a continuous keel between the squamosal horn and supraorbitals.
V. M. Arbour and P. J. Currie 2016Dif- fers from Anodontosaurus lambei and Scolosaurus cutleri in lacking subcircular caputegulae at the bases of the quadratojugal and squamosal horns (postocular caputegulae); differs from Anodontosaurus lambei in lacking interstitial osteoderms at the bases of the primary osteoderms of the first cervical half ring, and in having semicircular major osteoderms in dorsal view on the tail club; differs from Dyoplosaurus acutosquameus in having laterally directed sacral ribs; differs from Scolosaurus cutleri in having oval to subcircular-based keeled medial and lateral primary half ring osteoderms and in having a proportionately shorter postacetabular process of the ilium; differs from Ankylosaurus magniventris in having anteriorly directed nares, and in lacking a continuous keel between the squamosal horn and supraorbitals.
P. Penkalski 2018A moderate-sized (5 m) ankylosaurine with well-defined scales in the frontal region; erect squamosal boss with central apex and kinked (sigmoidal) keel; skull trapezoidal in dorsal view; nasal plate large, centrally convex; some skull plates with incipient nodes or keels, mostly in nasofrontal region; supraorbital bosses pointed in dorsal view and distinct, especially in lateral view; prominent quadratojugal horn with central apex; premaxillary palate divided; humerus less robust than in other euoplocephalins; deltopectoral crest does not extend past humeral midpoint; six oval, sharply keeled, fused osteoderms with central apices on h.r.1; keels on medial and lateral osteoderms sigmoidal (kinked); free osteoderms weakly ornamented, with fine (1-2 mm) pitting; many osteoderms oval and sharply keeled; some osteoderms with offset, L-shaped or folded keel and/ or double-edged perimeter.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yesg
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: herbivoreo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2005-08-25 06:07:00
Modified: 2005-08-25 08:07:00
Source: g = genus, o = order
References: Marsh 1875, Peczkis 1995

Age range: Late/Upper Campanian or 83.60000 to 72.20000 Ma

Collections (6 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Campanian83.6 - 72.2Canada (Alberta) Anodontosaurus lambei (135807) Euoplocephalus sp. (48618) Euoplocephalus tutus (59004 223397) Scelidosaurus sp. (type locality: 155228)
Middle Campanian83.6 - 72.2Canada (Alberta) Euoplocephalus tutus (196420)