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Scolosaurus cutleri

Reptilia - Ankylosauridae

Taxonomy
Scolosaurus cutleri was named by Nopcsa (1928). Its type specimen is BMNH R.5161, a partial skeleton (nearly complete skeleton with in situ osteoderms and skin impressions, lacking skull, distal half of tail, right forelimb and right hind limb), and it is a 3D body fossil. Its type locality is RTMP Quarry 80, Deadlodge Canyon, which is in a Campanian/Campanian channel sandstone/claystone in the Dinosaur Park Formation of Canada.

It was synonymized subjectively with Euoplocephalus tutus by Coombs (1986), Carpenter (2001), Ford and Kirkland (2001), Vickaryous et al. (2004), Ryan and Evans (2005), Currie (2005).

Synonymy list
YearName and author
1928Scolosaurus cutleri Nopcsa p. 54 figs. Pl. 6-7
1929Scolosaurus cutleri Swinton p. 22
1931Scolosaurus cutleri Huene p. 198
1932Scolosaurus cutleri Anonymous p. 574
1933Scolosaurus cutleri Sternberg p. 233
1934Scolosaurus cutleri Swinton p. 45
1954Scolosaurus cutleri Maleev
1956Scolosaurus cutleri Maleev
1964Scolosaurus cutleri Kuhn p. 51
1964Scolosaurus cutleri Tatarinov p. 577
1966Scolosaurus cutleri Russell p. 32
1970Scolosaurus cutleri Swinton p. 249
2000Scolosaurus cutleri Ford p. 175
2013Scolosaurus cutleri Arbour and Currie p. 31
2013Scolosaurus cutleri Kirkland et al. p. 7
2013Scolosaurus cutleri Penkalski and Blows p. 172
2014Scolosaurus cutleri Arbour et al. p. 2
2014Scolosaurus cutleri Arbour et al. p. 648
2014Scolosaurus cutleri Blows p. 58
2014Scolosaurus cutleri Han et al. p. 12 fig. 9
2014Scolosaurus cutleri Penkalski p. 617
2016Scolosaurus cutleri Arbour and Currie p. 420
2017Scolosaurus cutleri Arbour and Mallon p. 772
2018Scolosaurus cutleri Penkalski p. 265

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Eugnathostomata
—Teleostomi(Bonaparte 1836)
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
subclassEureptilia()
RankNameAuthor
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Ornithischia()
—Genasauria
—Thyreophora()
—Ankylosauria()
familyAnkylosauridae
subfamilyAnkylosaurinaeNopcsa 1918
tribeEuoplocephalini
genusScolosaurus
speciescutleri

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Scolosaurus cutleri Nopcsa 1928
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Diagnosis
ReferenceDiagnosis
P. Penkalski and W. T. Blows 2013Large (6 m) ankylosaurid with osteoderms that are predominantly conical, subconical, or mammillary in shape; presacral rod of four dorsosacral vertebrae with noticeably constricted intervertebral faces; ilia weakly divergent; ischium with unusually straight proximal border and slight posterior inflection of shaft distally; forelimb well armoured with conical spines. Differs from Dyoplosaurus and Euoplocephalus in the virtual absence of oval to subrectangular, low-keeled osteoderms; also differs from Euoplocephalus in having rugose, perforate osteoderms, in having an anteroposteriorly broad first cervical half-ring with osteoderms that have a posteriorly situated apex, in having a more robust humerus with a more distally expanded deltopectoral crest, and in having a more elongate, sigmoidal radius; differs from Dyoplosaurus in having oval sacral fenestrae, in the orientation of the third sacral ribs, in having subconical and mammillary spines in the pelvic and caudal regions instead of laterally compressed, high-pitched scutes, in the shape of the radius, and in having more common, hoof-shaped pedal unguals; differs from Ankylosaurus primarily in the complete lack of large, plate-like keeled osteoderms.
V. M. Arbour and P. J. Currie 2013Differs from Anodontosaurus lambei and Euoplocephalus tutus in the morphology of the squamosal horns, which are proportionately longer, backswept, and with distinct apices; differs from Euoplocephalus tutus in having small circular caputegulae at the bases of the squamosals and quadratojugals; differs from Anodontosaurus lambei, Euoplocephalus tutus, and Dyoplosaurus acutosquameus in having a proportionately longer postacetabular process of the ilium; differs from Anodontosaurus lambei and Euoplocephalus tutus in having proportionately large circular medial osteoderms with a low central prominences, and compressed, half-moon shaped lateral/distal osteoderms on the cervical half rings; differs from Dyoplosaurus acutosquameus in having laterally-directed sacral ribs; differs from Dyoplosaurus acutosquameus in having conical, osteoderms with centrally positioned apices on the lateral sides of the anterior portion of the tail; differs from Anodontosaurus and Dyoplosaurus in having a circular tail club knob in dorsal view, rather than a tail club knob that is wider than long (Anodontosaurus) or longer than wide (Dyoplosaurus); differs from Ankylosaurus magniventris in having anteriorly-directed nares, and in lacking a continuous keel between the squamosal horn and supraorbitals.
V. M. Arbour and P. J. Currie 2016Differs from Anodontosaurus lambei and Euoplocephalus tutus in the morphology of the squamosal horns, which are proportionately longer, backswept and with distinct apices; differs from Euoplocephalus tutus in having small circular caputegulae at the bases of the squamosals and quadratojugals; differs from Anodontosaurus lambei, Dyoplosaurus acutosquameus and Euoplocephalus tutus in having a proportionately longer postacetabular process of the ilium; differs from Anodontosaurus lambei and Euoplocephalus tutus in having proportionately large cir- cular medial osteoderms with low central prominences, and compressed, half-moon-shaped lateral/distal osteoderms on the cervical half rings; differs from Dyoplosaurus acutosquameus in having laterally directed sacral ribs; differs from Dyoplosaurus acutosquameus in having conical osteoderms with centrally positioned apices on the lateral sides of the anterior portion of the tail; differs from Anodontosaurus and Dyoplosaurus in having a circular tail club knob in dorsal view, rather than a tail club knob that is wider than long (Anodontosaurus) or longer than wide (Dyoplosaurus).
P. Penkalski 2018Weak intervertebral swellings on presacral rod; nine vertebrae in synsacrum (4-3-2, plus cd 1); otherwise as for genus.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: herbivoreo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2004-12-14 12:50:52
Modified: 2004-12-14 14:50:52
Source: o = order
Reference: Marsh 1875

Age range: Late/Upper Campanian or 83.60000 to 72.20000 Ma

Collections (2 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Campanian83.6 - 72.2Canada (Alberta) Ankylosauria indet. (59023)
Middle Campanian - Late/Upper Campanian83.6 - 72.2Canada (Alberta) Scolosaurus cutleri (type locality: 59015)