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Helophoridae

Insecta - Coleoptera - Helophoridae

Taxonomy
Helophorida was named by Leach (1815). It is extant.

It was reranked as the tribe Helophorina by Sahlberg (1880); it was reranked as the subfamily Helophorinae by Lawrence and Newton (1995) and Bouchard et al. (2011); it was corrected as Helophoridae by Short and Fikáček (2013) and Cai et al. (2022).

It was assigned to Hydrophilidae by Sahlberg (1880), Lawrence and Newton (1995) and Bouchard et al. (2011); and to Hydrophiloidea by Short and Fikáček (2013) and Cai et al. (2022).

Subtaxa
Helophorus

Synonymy list
YearName and author
1880Helophorina Sahlberg p. 61
1995Helophorinae Lawrence and Newton, Jr.
2011Helophorinae Bouchard et al. p. 154
2013Helophoridae Short and Fikáček
2022Helophoridae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyHydrophiloidea
familyHelophoridae()
familyHelophoridae()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Helophoridae Leach 1815
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G. Helophorus Fabricius 1775
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Invalid names: Helophorus (Gephelophorus) lomnicki d'Orchymont 1927 [synonym], Helophorus (Trichelophorus) pleistocenicus Lomnicki 1894 [synonym]
Subg. Helophorus (Meghelophorus) Kuwert 1886
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Invalid names: Helophorus (Praehelephorus) dzieduszyckii Lomnicki 1894 [synonym]
Invalid names: Helophorus (Praehelephorus) Lomnicki 1894 [synonym]
Subg. †Helophorus (Mesohelophorus) Ponomarenko 1977
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Invalid names: Mesohelophorus sibiricus Ponomarenko 1977 [replaced]
Subg. †Helophorus (Mesosperchus) Ponomarenko 1977
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†Helophorus (Mesosperchus) gracilis Prokin et al. 2010
†Helophorus (Mesosperchus) inceptivus Fikáček et al. 2012
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Invalid names: Mesohelophorus elongatus Ponomarenko 1990 [synonym]
†Helophorus (Mesosperchus) yixianus Fikáček et al. 2012
Subg. Helophorus (Orphelophorus) d'Orchymont 1927
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Invalid names: Helophorus (Orphelophorus) wandereri d'Orchymont 1927 [synonym]
Subg. Helophorus (Trichelophorus) Kuwert 1886
Helophorus auricollis Eschscholtz 1822
Helophorus browni McCorkle 1973
Helophorus eclectus d'Orchymont 1945
Helophorus lacustris Le Conte 1850
Helophorus nubilus Fabricius 1777
Helophorus oblongus Le Conte 1850
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Invalid names: Helophorus (Rhopalhelophorus) kuwerti Lomnicki 1894 [synonym]
Helophorus orientalis Motschulsky 1860
Helophorus praenanus Lomnicki 1894
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Invalid names: Helophorus polonicus Lomnicki 1894 [synonym]
Helophorus splendidus Sahlberg 1880
Helophorus tuberculatus Gyllenhal 1808
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Tithonian to the top of the Late/Upper Pleistocene or 152.10000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma

Collections (74 total)


Time interval Ma Country or state Original ID and collection number
Tithonian152.1 - 145.0Russian Federation (Chita) Mesohelophorus elongatus (125407) Mesosperchus tarsalis (119424 123197 152174) Mesosperchus tarsalis, Mesohelophorus elongatus (114307)
Tithonian152.1 - 145.0Mongolia Helophorus inceptivus (131803) Helophorus spp., Helophorus costalis (104748) Helophorus spp., Helophorus inceptivus, Helophorus costalis (121394)
Tithonian152.1 - 145.0Russian Federation Mesosperchus tarsalis (125408 138150)
Hauterivian132.9 - 129.4Russian Federation (Chita) Mesosperchus tarsalis (151559)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Hydrophilopsia gracilis, Helophorus yixianus (111716)
Barremian129.4 - 125.0Russian Federation (Chita) Mesosperchus tarsalis (151642)
Aptian125.0 - 113.0Russian Federation (Buryatia) Mesohelophorus sibiricus (105536 105969)
Early/Lower Aptian121.4 - 122.46Russian Federation (Chita) Mesosperchus tarsalis (152173)
Aquitanian - Burdigalian23.03 - 15.97Russian Federation Helophorus sibiricus (117611)
Late/Upper Miocene11.608 - 5.333Russian Federation Helophorus tuberculatus (153434)
Messinian7.246 - 5.333USA (Alaska) Helophorus sp. (123352)
Gelasian2.588 - 1.806Greenland Helophorus khnzoriani (160143) Helophorus spp., Helophorus khnzoriani (160144) Helophorus spp., Helophorus tuberculatus, Helophorus khnzoriani (160109)
Early/Lower Pleistocene2.588 - 0.781Russian Federation Helophorus splendidus (163069)
Chibanian0.774 - 0.129Russian Federation Helophorus obscurellus (163093) Helophorus splendidus (163094 163097)
Middle Pleistocene0.774 - 0.129United Kingdom Helophorus sp. (168265 168266 168267 168270 168271) Helophorus spp. (168264 168269)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Helophorus sp. (169389)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Helophorus brevipalpis (164903 164920)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Helophorus sp. (167675)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Helophorus sempervarians (137207 137210 137211 137213) Helophorus sp., Helophorus sempervarians (137208) Helophorus tuberculatus (137209) Helophorus tuberculatus, Helophorus sempervarians (137212)
Late/Upper Pleistocene0.129 - 0.0117USA (Alaska) Helophorus splendidus (169385)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Helophorus aquaticus, Helophorus nubilus, Helophorus brevipalpis (163070)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Helophorus splendidus (153437 153439 153443 153446 173208)
Late/Upper Pleistocene0.129 - 0.0117Germany Helophorus lomnicki, Helophorus wandereri (156167)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Helophorus auricollis (169350) Helophorus eclectus (169347)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Helophorus lacustris (169359 169363) Helophorus oregonensis (169370 169372 169374 169376) Helophorus oregonensis, Helophorus lacustris (169366) Helophorus oregonensis, Helophorus sp., Helophorus lacustris (169361) Helophorus sp. (169362) Helophorus sp., Helophorus lacustris (169360) Helophorus spp. (157092 167662 169375 169379)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Helophorus pleistocenicus, Helophorus dzieduszyckii, Helophorus praenanus, Helophorus polonicus, Helophorus kuwerti, Helophorus aquaticus, Helophorus lomnicki (153476) Helophorus sp., Helophorus sibiricus, Helophorus aquaticus, Helophorus praenanus, Helophorus grandis, Helophorus glacialis, Helophorus aspericollis, Helophorus orientalis, Helophorus browni, Helophorus discrepans (113753)
Tarantian0.129 - 0.0117USA (Pennsylvania) Helophorus sp. (137201 137204) Helophorus sp., Helophorus sempervarians (137205 137206)