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Macraulacinae

Insecta - Coleoptera - Eucnemidae

Synonymy list
YearName and author
1923Macraulacinae Fleutiaux
1993Macraulacinae Muona
1995Macraulacinae Lawrence and Newton, Jr.
2011Macraulacinae Bouchard et al.
2016Macraulacinae Webster p. 449
2019Macraulacinae Otto p. 2
2020Macraulacinae Muona p. 4
2021Macraulacinae Muona p. 9
2022Macraulacinae Cai et al.
2022Macraulacinae Muona p. 2
2023Macraulacinae Otto p. 13

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
RankNameAuthor
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderElateriformiaCrowson 1960
superfamilyElateroidea
familyEucnemidaeEschscholtz 1829
subfamilyMacraulacinaeFleutiaux 1923
subfamilyMacraulacinaeFleutiaux 1923

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Macraulacinae Fleutiaux 1923
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Tr. Echthrogasterini Cobos 1965
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G. Hemiopsida Macleay 1872
Tr. Euryptychini Mamaev 1976
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G. †Euryptychoides Muona 2021
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†Euryptychoides magnus Muona 2021
G. Euryptychus Leconte 1852
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†Euryptychus brunhildae Muona 1993
†Euryptychus burmensis Muona 2020
†Euryptychus decoratus Muona 2021
†Euryptychus excavatus Muona 2021
†Euryptychus gerhildae Muona 1993
†Euryptychus helmwigae Muona 2021
†Euryptychus mysticus Muona 2020
†Euryptychus siegfriedi Muona 1993
†Euryptychus sieglindae Muona 2021
†Euryptychus waltrautae Muona 2021
G. †Muonabuntor Li et al. 2020
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†Muonabuntor grandinotalis Li et al. 2020
Tr. Jenibuntorini Muona 1993
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G. Jenibuntor Muona 1993
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†Jenibuntor pusillus Muona 2020
Tr. Macraulacini Fleutiaux 1923
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G. Asiocnemis Mamaev 1976
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†Asiocnemis colombicus Muona 2022
G. Ceratus de Bonvouloir 1872
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†Ceratus wotani Muona 1993
G. Deltometopus de Bonvouloir 1871
G. Dromaeolus Kiesenwetter 1858
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†Dromaeolus argenteus Otto 2023
G. Fornax Laporte 1835
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†Fornax dominicensis Otto 2023
†Fornax ledensis Scudder 1895
†Fornax relictus Wickham 1916
†Fornax serropalpoides Otto 2023
G. Plesiofornax Coquerel 1866
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†Plesiofornax caribica Otto 2023
G. Spinifornax Fleutiaux 1926
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†Spinifornax donneri Muona 1993
G. Thambus Bonvouloir 1871
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†Thambus woodruffi Muona 2022
Tr. Nematodini Leiler 1976
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G. Coomanius Fleutiaux 1924
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†Coomanius balticus Muona 2021
G. Nematodes Berthold 1827
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†Nematodes miocenensis Otto 2023
†Nematodes thoracicus Otto 2023
Tr. Orodotini Muona 1993
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G. †Erdaia Muona 1993
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†Erdaia bispinulosa Muona 2021
†Erdaia guntheri Muona 1993
†Erdaia hageni Muona 1993
†Erdaia longicornis Muona 2021
†Erdaia longitarsis Muona 2021
G. †Neusiokia Muona 2022
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Tr. †Throscogeniini Yablokov-Khnzorian 1962
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G. †Throscogenius Yablokov-Khnzorian 1962
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†Throscogenius takhtajani Yablokov-Khnzorian 1962
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Early/Lower Cenomanian to the top of the Holocene or 99.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 93.5 Ma

Collections (14 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Cenomanian99.6 - 93.5Myanmar Euryptychus acutangulus, Euryptychus burmensis, Euryptychus elegantulus, Euryptychus mysticus, Jenibuntor pusillus (200411) Muonabuntor grandinotalis (161726)
Early/Lower Campanian83.5 - 70.6USA (North Carolina) Neusiokia appalachiensis (152037)
Priabonian38.0 - 33.9Denmark Euryptychus siegfriedi (123762)
Priabonian38.0 - 33.9Lithuania Euryptychus excavatus (123460) Throscogenius takhtajani (113470)
Priabonian38.0 - 33.9Germany (Halle) Fornax sp., Erdaia guntheri, Euryptychus siegfriedi, Euryptychus gerhildae (129227)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Erdaia bispinulosa, Erdaia longicornis, Erdaia longitarsis, Euryptychus decoratus, Euryptychus helmwigae, Euryptychus waltrautae, Euryptychus sieglindae, Euryptychoides magnus, Coomanius balticus, Nematodes schwertleitae (200410) Fornax sp., Macraulacini indet., Hemiopsida sp., Erdaia sp., Erdaia hageni, Euryptychus brunhildae, Spinifornax donneri, Ceratus wotani, Asiocnemis sp., Dromaeolus sp. (147612)
Chadronian37.2 - 33.9USA (Colorado) Fornax relictus (113749)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Euryptychus antilliensis, Euryptychus hispaniolus, Plesiofornax caribica, Fornax dominicensis, Fornax serropalpoides, Dromaeolus argenteus, Nematodes miocenensis, Nematodes thoracicus (229111) Thambus woodruffi (229103)
Late/Upper Pleistocene0.129 - 0.0117Canada (Ontario) Fornax ledensis (152573)
Holocene0.0117 - 0.0Colombia (Santander) Asiocnemis colombicus (229104)