PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Calosoma

Insecta - Coleoptera - Carabidae

Taxonomy

Calosoma was named by Weber (1801). It is extant.

It was assigned to Eleutherata by Weber (1801); to Metallici by Latreille (1802), Latreille (1804); to Carabiques by Dejean (1821); to Carabidae by Scudder (1900), Theobald (1937), Zhang (1989); and to Carabini by LeConte (1861), Tamutis et al. (2011), Bouchard et al. (2011).

Synonymy list

Year	Name and author
1801	Calosoma Weber p. 20
1802	Calosoma Latreille p. 91
1804	Calosoma Latreille p. 140
1821	Calosoma Dejean p. 7
1861	Calosoma LeConte p. 10
1900	Calosoma Scudder p. 16
1937	Calosoma Theobald p. 295
1989	Calosoma Zhang p. 95
2011	Calosoma Bouchard et al.
2011	Calosoma Tamutis et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta

Rank	Name	Author
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Adephaga
family	Carabidae	()
subfamily	Carabinae
tribe	Carabini
genus	Calosoma

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Calosoma Weber 1801

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Subg. Calosoma (Campalita) Motschulsky 1865

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†Campalita deplanatum Heer 1860

†Calosoma agassizi Barthelemy-Lapommeraye 1846

†Calosoma brunneum Hong 1985

†Calosoma calvini Wickham 1909

†Calosoma caraboides Heer 1860

†Calosoma catenulatum Heer 1860

†Calosoma ceresti Nel 1987

†Calosoma cockerelli Wickham 1910

†Calosoma emmonsi Scudder 1900

†Calosoma escheri Heer 1860

†Calosoma escrobiculatum Heer 1860

†Calosoma fernquisti Cockerell 1925

†Calosoma grasui Paucă and Ciobanu 1978

†Calosoma heeri Scudder 1895

Calosoma inquisitor Linnaeus 1758

†Calosoma jaccardi Heer 1860

†Calosoma kimi Lee and Nam 2021

Calosoma maderae Fabricius 1775

†Calosoma nauckianum Heer 1860

Calosoma semilaeve LeConte 1852

Calosoma sycophanta Linnaeus 1758 [forest caterpillar hunter]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Chadronian to the top of the Rancholabrean or 37.00000 to 0.01400 Ma
Minimum age of oldest fossil (stem group age): 33.9 Ma

Collections (32 total)

Time interval	Ma	Country or state	Original ID and collection number
Priabonian	37.71 - 33.9	United Kingdom (England)	C. sp. (123960)
Chadronian	37.0 - 33.9	USA (Colorado)	C. calvini (106379) C. calvini, C. cockerelli (124002) C. emmonsi (113749) C. emmonsii (106384)
MP 23	33.9 - 27.3	France	C. agassizi (122535) C. agassizi, Carabus ceresti (122984 155667)
Rupelian	33.9 - 27.3	France (Alpes-de-Haute)	C. agassizi (155208)
Rupelian	33.9 - 27.3	Romania	C. grasui (113962)
Chattian	27.3 - 23.04	France (Bouches-Du-Rhone)	C. agassizi (113712 122582 123943)
Chattian	27.3 - 23.04	France (Provence-Alpes-Côte d'Azur)	C. agassizi (122487)
Late/Upper Oligocene	27.3 - 23.04	France (Puy-de-Dome)	C. agassizi (155204)
Late/Upper Oligocene	27.3 - 23.04	France (Aix-en-Provence)	C. agassizi (114117)
Early/Lower Miocene	23.04 - 15.98	South Korea	C. kimi (223984)
Burdigalian	20.45 - 15.98	China (Shandong)	C. maderae (128549 128550 128551 128552) Shanwangicarabus brunneus (130412)
Burdigalian	20.45 - 15.98	Greece	C. exscrobiculatum (144526)
Langhian	15.98 - 13.82	USA (Washington)	C. fernquisti (122750)
Middle Miocene	15.98 - 11.63	Japan	C. sp. (153619)
Sarmatian	12.8 - 11.63	Switzerland	C. jaccardi, C. caraboides (153996)
Sarmatian	12.8 - 11.63	Germany (Baden-Württemberg)	C. escrobiculatum, C. nauckianum, C. catenulatum, C. deplanatum, C. escheri (117252) C. heeri (113830)
Vallesian - Turolian	11.63 - 5.09	Spain (Lleida)	Campalita deplanatum (140806)
Piacenzian	3.6 - 2.58	Germany (Niedersachsen)	C. maderae, C. inquisitor, C. sycophanta (109196)
Early/Lower Pleistocene	2.58 - 0.774	Japan	C. inquisitor (153617)
Rancholabrean	0.21 - 0.014	USA (California)	C. semilaeve (150336)