PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Bembidion

Insecta - Coleoptera - Carabidae

Taxonomy

Bembidion was named by Latreille (1802). It is extant.

It was assigned to Elaphrii by Latreille (1802); to Carabiques by Dejean (1821); to Bembidiinae by Casey (1918); to Carabidae by Gemminger and Harold (1868), Scudder (1877), Scudder (1900), Piton (1940), Zhang (1989); to Bembidiina by Bouchard et al. (2011), Maddison (2012), Ortuño and Arillo (2015); and to Bembidiini by LeConte (1861), Liebherr and Porch (2015), Maddison et al. (2019).

Synonymy list

Year	Name and author
1802	Bembidion Latreille p. 82
1821	Bembidium Dejean p. 16
1861	Bembidium LeConte p. 36
1868	Bembicidium Gemminger and Harold
1877	Bembidium Scudder p. 759
1900	Bembidium Scudder p. 18
1918	Bembidion Casey p. 4
1940	Bembidium Piton p. 172
1989	Bembidion Zhang p. 101
2011	Bembidion Bouchard et al.
2012	Bembidion Maddison
2015	Bembidion Liebherr and Porch p. 208
2015	Bembidion Ortuño and Arillo p. 578
2019	Bembidion Maddison et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia

Rank	Name	Author
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Adephaga
family	Carabidae	()
subfamily	Trechinae	()
—	Trechitae
tribe	Bembidiini	Stephens 1827
genus	Bembidion

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Bembidion Latreille 1802

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Subg. Bembidion (Asioperyphus) Vysoky 1986

Bembidion (Asioperyphus) bimaculatum Kirby 1837

Bembidion (Asioperyphus) lenae Csiki 1928

Bembidion (Asioperyphus) sordidum Kirby 1837

Subg. Bembidion (Bembidion) Latreille 1802

Bembidion (Bembidion) mutatum Gemminger and Harold 1868

Bembidion (Bembidion) quadrimaculatum Linnaeus 1761

Subg. Bembidion (Bracteon) Bedel 1879

Bembidion (Bracteon) alaskense Lindroth 1962

Bembidion (Bracteon) balli Lindroth 1962

Bembidion (Bracteon) inaequale Say 1823

Bembidion (Bracteon) lapponicum Zetterstedt 1828

Bembidion (Bracteon) levettei Casey 1918

Bembidion (Bracteon) velox Linnaeus 1761

Invalid names: Bembidion (Chrysobracteon) Netolitzky 1914 [synonym]

Subg. Bembidion (Diplocampa) Bedel 1896

Bembidion (Diplocampa) assimile Gyllenhal 1810

Subg. †Bembidion (Eodontium) Schmidt and Michalik 2017

†Bembidion (Eodontium) bukejsi Schmidt and Michalik 2017

Subg. Bembidion (Eupetedromus) Netolitzky 1911

†Bembidion (Eupetedromus) alekseevi Schmidt and Michalik 2017

Subg. Bembidion (Furcacampa) Netolitzky 1931

Bembidion (Furcacampa) versicolor LeConte 1847

Subg. Bembidion (Hirmoplataphus) Lindroth 1963

Bembidion (Hirmoplataphus) salebratum LeConte 1848

Subg. Bembidion (Hydrium) LeConte 1847

Invalid names: Bembidion (Eurytrachelus) Motschulsky 1850 [synonym]

Subg. Bembidion (Melomalus) Casey 1918

Bembidion (Melomalus) planatum LeConte 1848

Subg. Bembidion (Metallina) Motschulsky 1850

Bembidion (Metallina) dyschirinum LeConte 1861

Bembidion (Metallina) lampros Herbst 1784

Subg. Bembidion (Neobembidion) Bousquet 2006

Bembidion (Neobembidion) constricticolle Hayward 1897

Subg. Bembidion (Nesocidium) Sharp 1903

Bembidion (Nesocidium) ignicola Blackburn 1879

Bembidion (Nesocidium) pacificum Blackburn 1878

Subg. Bembidion (Notaphus) Dejean 1821

Bembidion (Notaphus) coloradense Hayward 1897

Bembidion (Notaphus) dorsale Say 1823

Bembidion (Notaphus) nigripes Kirby 1837

Bembidion (Notaphus) varium Olivier 1795

Subg. Bembidion (Ochthedromus) LeConte 1847

†Bembidion (Ochthedromus) davidae Pierce 1944

Subg. Bembidion (Peryphanes) Jeannel 1941

Bembidion (Peryphanes) grapii Gyllenhal 1827

Bembidion (Peryphanes) yukonum Fall 1926

Subg. Bembidion (Peryphus) Dejean 1821

Bembidion (Peryphus) tetracolum Say 1823

Bembidium (Peryphus) tibiale Duftschmid 1812

Subg. Bembidion (Philochthemphanes) Netolitzky 1943

†Bembidion (Philochthemphanes) christelae Ortuño and Arillo 2010

Invalid names: Bembidion (Archaeophilochthus) Ortuño and Arillo 2010 [synonym]

Subg. Bembidion (Philochthus) Stephens 1828

Bembidion (Philochthus) biguttatum Fabricius 1779

Bembidion (Philochthus) guttula Fabricius 1792

Subg. Bembidion (Phyla) Motschulsky 1844

Bembidion (Phyla) obtusum Audinet-Serville 1821

Subg. Bembidion (Plataphus) Motschulsky 1864

Bembidion (Plataphus) arcticum Lindroth 1963

Bembidion (Plataphus) breve Motschulsky 1845

Bembidion (Plataphus) fellmanni Mannerheim 1823

Bembidion (Plataphus) gebleri Gebler 1833

Bembidion (Plataphus) haruspex Casey 1918

Bembidion (Plataphus) planiusculum Mannerheim 1843

Bembidion (Plataphus) rusticum Casey 1918

Invalid names: Bembidion (Plataphodes) Ganglbauer 1891 [synonym]

Subg. Bembidion (Princidium) Motschulsky 1864

Bembidion (Princidium) punctulatum Drapiez 1821

Subg. Bembidion (Semicampa) Netolitzky 1910

Bembidion (Semicampa) morulum LeConte 1863

Subg. Bembidion (Terminophanes) Müller-Motzfeld 1998

Bembidion (Terminophanes) mckinleyi Fall 1926

Subg. Bembidion (Testedium) Motschulsky 1864

Bembidion (Testedium) bipunctatum Linnaeus 1761

Subg. Bembidion (Trepanedoris) Netolitzky 1918

Bembidion (Trepanedoris) doris Panzer 1797

Bembidion (Trepanedoris) fortestriatum Motschulsky 1845

Bembidion (Trepanedoris) frontale LeConte 1847

Bembidion (Trepanedoris) pseudocautum Lindroth 1963

Subg. Bembidion (Trepanes) Motschulsky 1864

Bembidion (Trepanes) articulatum Panzer 1796

Bembidion (Trepanes) octomaculatum Goeze 1777

Bembidion aeneum Germar 1824

†Bembidion damnosum Scudder 1900

†Bembidion everestae Pierce 1944

Bembidion femoratum Sturm 1825

†Bembidion festivum Zhang et al. 1994

Bembidion gilvipes Sturm 1825

Bembidion lunatum Duftschmid 1812

Bembidion minimum Fabricius 1792

Bembidion obliquum Sturm 1825

Bembidion schueppeli Dejean 1831

Bembidion semipunctatum Donovan 1806

†Bembidion strenum Zhang 1989

Bembidion virens Gyllenhal 1827

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Priabonian to the top of the Holocene or 37.71000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 33.9 Ma

Collections (91 total)

Time interval	Ma	Country or state	Original ID and collection number
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	B. alekseevi (139928) B. bukejsi (152022) B. christelae (123462)
Burdigalian	20.45 - 15.98	USA (California)	B. davidae (161515) B. everestae (161516)
Burdigalian	20.45 - 15.98	China (Shandong)	B. festivum (138946) B. strenum (128552)
Late/Upper Miocene	11.63 - 5.333	Russian Federation	B. sp. 1 (153434) B. sp. 4, B. sp. 5, B. sp. 1, B. sp. 2, B. sp. 3, B. sp. (113713)
Messinian	7.246 - 5.333	USA (Alaska)	B. sp., B. sp., B. spp. (123352)
Gelasian	2.58 - 1.8	Greenland	B. mckinleyi (160159) B. spp., B. dyschirinum, B. sordidum (160108) B. spp., B. fellmanni (160152) B. spp., B. fellmanni, B. mckinleyi, B. sordidum (160151) B. spp., B. levettei, B. velox, B. lampros, B. yukonum (160153) B. spp., B. sordidum (160147) B. spp., B. sp., B. spp., B. levettei, B. lapponicum, B. balli, B. vitiosum, B. dyschirinum, B. fellmanni, B. difficile, B. arcticum, B. planatum, B. planiusculum, B. gebleri, B. salebratum, B. mckinleyi, B. grapii, B. yukonum, B. bimaculatum, B. sordidum (160109) B. spp., B. sp., B. spp., B. velox, B. dyschirinum, B. fellmanni, B. arcticum, B. planiusculum, B. gebleri, B. lenae, B. yukonum, B. sordidum (160146) B. spp., B. spp., B. levettei, B. alaskense, B. dyschirinum, B. fellmanni, B. planatum, B. planiusculum, B. gebleri, B. mckinleyi, B. yukonum, B. bimaculatum, B. sordidum (160144)
Early/Lower Pleistocene	2.58 - 0.774	Russian Federation	B. spp. (163069 163089)
Chibanian	0.774 - 0.129	Japan	B. sp. (165704 165705 165707 165709)
Chibanian	0.774 - 0.129	Russian Federation	B. sp. (163091 163095)
Chibanian	0.774 - 0.129	Japan (Kumamoto)	B. sp. (164901)
Middle Pleistocene	0.774 - 0.129	United Kingdom (Warwickshire)	B. assimile (169391 169392 169393) B. sp., B. assimile, B. schueppeli (169388)
Middle Pleistocene	0.774 - 0.129	United Kingdom	B. doris, B. articulatum, B. octomaculatum (168266) B. doris, B. punctulatum, B. tetracolum, B. assimile (168264) B. lampros, B. doris, B. assimile, B. minimum, B. articulatum, B. obtusum (168265) B. lampros, B. doris, B. punctulatum, B. tetracolum, B. quadrimaculatum (168267) B. lampros, B. doris, B. varium, B. semipunctatum, B. tetracolum, B. gilvipes, B. biguttatum (168270) B. lampros, B. gilvipes, B. assimile, B. minimum, B. articulatum (168268) B. lampros, B. semipunctatum, B. assimile, B. articulatum (168269) B. lampros, B. varium, B. assimile (168272) B. lunatum, B. gilvipes, B. assimile (168271)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Wisconsin)	B. sp., B. mutatum, B. grapei (169406)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation	B. sp. (153443 153444 153446 163101 163102 163104 163106 163110 173201 173202 173205 173206 173208 173216)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Washington)	B. fortestriatum (169379) B. haruspex, B. planiusculum (169374) B. planiusculum (169365) B. rusticum (169359) B. sp. (169360 169370 169376 169378) B. spp., B. bimaculatum, B. inaequale, B. nigripes (157092 167662)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (British Columbia)	B. spp., B. dorsale (167673)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Pennsylvania)	B. fortestriatum (137209 137212) B. mutatum (137210) B. versicolor (137211)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (Worcestershire)	B. bipunctatum (163075) B. spp., B. lampros, B. bipunctatum, B. obliquum, B. virens, B. femoratum, B. lunatum, B. aeneum (163070)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (Ontario)	B. balli (174230) B. sp. 1, B. sp. 2, B. balli (174229) Bembidium damnosum (152577)
Late/Upper Pleistocene	0.129 - 0.0117	Ukraine	Bembidium tibiale, Bembidium bipunctatum (153476)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (England)	B. doris (164906 164909) B. doris, B. guttula (164905) B. guttula (164904)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Colorado)	B. bimaculatum, B. coloradense, B. constricticolle (169336) B. sp. (169333) B. sp., B. breve (169347 169350)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation (Astrakhan)	B. sp., B. minimum (125794)
Tarantian	0.129 - 0.0117	USA (Pennsylvania)	B. anguliferum group (137201 137204) B. anguliferum group, B. versicolor, B. pseudocautum, B. frontale (137206) B. incertum group, B. mutatum (137205)
Holocene	0.0117 - 0.0	USA (Hawaii)	B. ignicola, B. pacificum (224098)