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Eumetazoa

Taxonomy
Eumetazoa was named by Butchili (1910). It is extant.

It was synonymized subjectively with Animalia by Tsai and Fordyce (2015).

It was assigned to Metazoa by Peterson et al. (2005); and to Proepitheliozoa by Nielsen (2012).

Subtaxa
Ausiidae, Bilateralomorpha, Eoandromeda, Funisia, Neuralia, Placozoa (syn. Phagocytellozoa), Plexus, Rangeomorpha, Shufangtubulus, Tribrachiomorpha, Triploblastica, Wutubus
Sister taxa
Bilateralomorpha, Neuralia, Tribrachiomorpha, Triploblastica

Synonymy list
YearName and author
1910Eumetazoa Butchili p. 32
2005Eumetazoa Peterson et al. p. 38
2012Eumetazoa Nielsen p. 33 figs. Fig. 4.2

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RankNameAuthor
kingdomAnimalia()
—Eumetazoa
RankNameAuthor
—Eumetazoa

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Eumetazoa Butchili 1910
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Fm. †Ausiidae Hahn and Pflug 1985
+
G. †Ausia Hahn and Pflug 1985
+
†Ausia fenestrata Hahn and Pflug 1985
G. †Burykhia Fedonkin et al. 2012
+
†Burykhia hunti Fedonkin et al. 2012
G. †Finkoella Martyshyn and Uchman 2021
+
†Finkoella oblonga Martyshyn and Uchman 2021
†Finkoella ukrainica Martyshyn and Uchman 2021
G. †Pharyngomorpha Martyshyn and Uchman 2021
+
†Pharyngomorpha reticulata Martyshyn and Uchman 2021
Unr. †Bilateralomorpha Erwin et al. 2011
+
G. †Brachina Wade 1972
+
†Brachina delicata Wade 1972
G. †Eoandromeda Tang et al. 2008
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†Eoandromeda octobrachiata Tang et al. 2008
G. †Funisia Droser and Gehling 2008
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†Funisia dorothea Droser and Gehling 2008
+
Invalid names: Phagocytellozoa [synonym]
G. †Plexus Joel et al. 2014
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†Plexus ricei Joel et al. 2014
+
G. †Avalofractus Narbonne et al. 2009
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†Avalofractus abaculus Narbonne et al. 2009
G. †Beothukis Brasier and Antcliffe 2009
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†Beothukis mistakensis Brasier and Antcliffe 2009
G. †Bradgatia Boynton and Ford 1995
+
†Bradgatia linfordensis Boynton and Ford 1995
G. †Charnia Ford 1958
+
†Charnia gracilis Wu et al. 2024
†Charnia grandis Glaessner and Wade 1966
†Charnia masoni Ford 1958
†Charnia siberica Sokolov 1972
Invalid names: Blackbrookia Boynton and Ford 1995 [synonym], Pseudovendia Boynton and Ford 1979 [synonym], Shepshedia Boynton and Ford 1995 [synonym]
Unr. †Charnida Narbonne et al. 2009
+
G. †Culmofrons Laflamme et al. 2012
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†Culmofrons plumosa Laflamme et al. 2012
Fm. †Charniidae Glaessner 1979
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G. †Fractofusus Gehling and Narbonne 2007 [Spindle]
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†Fractofusus andersoni Gehling and Narbonne 2007 [Spindle]
†Fractofusus misrai Gehling and Narbonne 2007 [Spindle]
G. †Frondophyllas Bamforth and Narbonne 2009
+
†Frondophyllas grandis Bamforth and Narbonne 2009
G. †Hapsidophyllas Bamforth and Narbonne 2009
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†Hapsidophyllas flexibilis Bamforth and Narbonne 2009
G. †Parviscopa Hofmann et al. 2008
+
†Parviscopa bonavistensis Hofmann et al. 2008
G. †Pectinifrons Bamforth et al. 2008 [Pectinate]
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†Pectinifrons abyssalis Bamforth et al. 2008 [Pectinate]
G. †Plumeropriscum Mason and Narbonne 2016
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†Plumeropriscum hofmanni Mason and Narbonne 2016
G. †Primocandelabrum Hofmann et al. 2008
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†Primocandelabrum hiemaloranum Hofmann et al. 2008
Invalid names: Hiemalora Fedonkin 1982 [synonym]
G. †Rangea Gurich 1930
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†Rangea schneiderhoehni Gurich 1929
†Rangea siberica Sokolov 1972
G. †Trepassia Narbonne et al. 2009
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†Trepassia wardae Narbonne and Gehling 2003
G. †Vinlandia Brasier et al. 2012
+
†Vinlandia antecedens Laflamme et al. 2007
G. †Shufangtubulus Dai and Hua 2024
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†Shufangtubulus inornatus Dai and Hua 2024
Unr. Triploblastica Lankester 1877
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Ph. Ctenophora Eschscholtz 1829 [comb jelly]
+
G. †Ctenorhabdotus Conway Morris and Collins 1996
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†Ctenorhabdotus campanelliformis Parry et al. 2021 [comb jelly]
Fm. †Dinomischidae Conway Morris 1977
+
G. †Calathites Luo et al. 1999
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†Calathites spinalis Luo et al. 1999
G. †Daihua Zhao et al. 2019
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†Daihua sanqiong Zhao et al. 2019
G. †Dinomischus Conway Morris 1977
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†Dinomischus isolatus Chen et al. 1989
†Dinomischus venustus Chen 1989
G. †Xianguangia Chen and Erdtmann 1991
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†Xianguangia sinica Chen and Erdtmann 1991
G. †Paleoctenophora Stanley and Sturmer 1983
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†Paleoctenophora brasseli Stanley and Sturmer 1983
Cl. †Scleroctenophora Ou et al. 2015
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G. †Batofasciculus Hou et al. 1999
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†Batofasciculus ramificans Hou et al. 1999
G. †Galeactena Ou et al. 2015
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†Galeactena hemispherica Ou et al. 2015
G. †Gemmactena Ou et al. 2015
+
†Gemmactena actinala Ou et al. 2015
G. †Maotianoascus Chen and Zhou 1997
+
†Maotianoascus octonarius Chen and Zhou 1997
G. †Thaumactena Ou et al. 2015
+
†Thaumactena ensis Ou et al. 2015
G. †Trigoides Luo et al. 1999
+
†Trigoides aclis Luo et al. 1999
G. †Sinoascus Chen and Zhou 1997
G. †Thalassostaphylos Parry et al. 2021 [comb jelly]
+
†Thalassostaphylos elegans Parry et al. 2021 [comb jelly]
G. †Yunnanoascus Hu et al. 2007
Invalid names: Tentaculata [empty]
Unr. †Eutriploblastica Zrzavy et al. 1998
+
+
Invalid names: Notandropora [synonym]
Unr. †Neotriploblastica Zrzavy et al. 1998
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Unr. †Eucoelomata
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Invalid names: Phoronozoa Zrzavy et al. 1998 [empty]
Unr. †Paracoelomata Zrzavy et al. 1998
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Invalid names: Neotrichozoa Zrzavy et al. 1998 [empty]
Invalid names: Rhabditophora [empty]
Invalid names: Nephrozoa Jondelius et al. 2002 [empty]
G. †Wutubus Chen et al. 2014
+
†Wutubus annularis Chen et al. 2014
Invalid names: Neuralia [empty], Tribrachiomorpha Grazhdankin 2014 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
No ecological data are available

Age range

Maximum range based only on fossils: base of the Poundian to the top of the Early/Lower Maastrichtian or 570.00000 to 66.00000 Ma
Minimum age of oldest fossil (stem group age): 541.0 Ma

Collections (130 total)


Time interval Ma Country or state Original ID and collection number
Neoproterozoic1000.0 - 541.0China (Guizhou) Eumetazoa indet. (97560 97561)
Ediacaran635.0 - 541.0China (Shaanxi) Shufangtubulus inornatus (237603)
Ediacaran635.0 - 541.0Namibia Rangea schneiderhoehni (6712 10397 236924 237207) Rangea schneiderhoeni (207853 207854 207855) Rangea sp. (9881)
Ediacaran635.0 - 541.0Norway Hiemalora sp. (51870)
Ediacaran635.0 - 541.0Russian Federation (Sakha) Charnia masoni (237144) Hiemalora stellaris, Charnia masoni, Hiemalora pleiomorphus (99226)
Ediacaran635.0 - 541.0China (Hubei) Charnia sp., Charniidae indet., Hiemalora pleiomorpha, Paracharnia dengyingensis, Wutubus annularis, Rangea sp. (235125) Hiemalora sp., Wutubus annularis (6853) Rangea schneiderhoehni, Paracharnia dengyingensis, Wutubus annularis, Charnia gracilis, Charnia sp. (236902)
Ediacaran635.0 - 541.0Russian Federation (Arkhangelsk) Charnia masoni, Charnia sp. (91644) Rangea schneiderhoehni (207834 237633) Rangea sp. (91650) Rangea sp., Ausia sp. (91643)
Ediacaran635.0 - 541.0United Kingdom (England) Charnia grandis, Bradgatia linfordensis (236883) Primocandelabrum sp., Charnia masoni, Bradgatia linfordensis (228457) Pseudovendia charnwoodensis (10287 10473)
Ediacaran635.0 - 541.0Namibia (Southeastern Namibia) Bradgatia sp. (89797)
Ediacaran635.0 - 541.0United Kingdom Charnia masoni, Bradgatia linfordensis (153338)
Ediacaran635.0 - 541.0Canada (Newfoundland and Labrador) Bradgatia linfordensis (80327 89121) Bradgatia linfordensis, Fractofusus misrai, Pectinifrons abyssalis, Hapsidophyllas sp., Hapsidophyllas flexibilis (89116) Bradgatia linfordensis, Fractofusus misrai, Plumeropriscum hofmanni (89122) Bradgatia linfordensis, Pectinifrons abyssalis (89114 89115 191945) Bradgatia linfordensis, Pectinifrons abyssalis, Beothukis sp. (191946) Charnia antecedens (80324 80353) Charnia antecedens, Bradgatia linfordensis (80326) Charnia antecedens, Bradgatia linfordensis, Fractofusus andersoni, Blackbrookia sp. (80341) Charnia masoni (89209) Charnia masoni, Bradgatia linfordensis, Fractofusus andersoni, Beothukis mistakensis (89119) Charnia masoni, Charnia antecedens, Charnia wardi (51741) Charnia masoni, Charnia grandis, Charnia antecedens (80351) Charnia masoni, Charnia wardi (51740) Charnia masoni, Fractofusus andersoni, Beothukis mistakensis, Charnia wardi, Trepassia wardae (89096) Charnia masoni, Pectinifrons abyssalis (89118) Charnia masoni, Pectinifrons abyssalis, Hapsidophyllas flexibilis (89111) Fractofusus andersoni (80337) Fractofusus andersoni, Blackbrookia sp. (80340) Hiemalora sp., Primocandelabrum hiemaloranum, Charnia masoni, Bradgatia linfordensis, Fractofusus misrai, Fractofusus andersoni, Pectinifrons abyssalis, Avalofractus abaculus, Beothukis mistakensis, Hapsidophyllas flexibilis, Culmofrons plumosa, Plumeropriscum hofmanni (89095) Hiemalora sp., Primocandelabrum hiemaloranum, Charnia masoni, Fractofusus andersoni, Beothukis mistakensis, Trepassia wardae, Culmofrons plumosa (236918) Hiemalora sp., Primocandelabrum sp., Bradgatia sp., Vinlandia sp. (160728) Hiemalora stellaris (80344 80352) Hiemalora stellaris, Bradgatia linfordensis (80323) Hiemalora stellaris, Charnia antecedens (80322 80333 80347) Hiemalora stellaris, Charnia masoni, Charnia antecedens (80349) Hiemalora stellaris, Charnia masoni, Charnia antecedens, Bradgatia linfordensis (80325) Hiemalora stellaris, Hapsidophyllas flexibilis (207808) Hiemalora stellaris, Primocandelabrum sp., Charnia antecedens (80335) Hiemalora stellaris, Primocandelabrum sp., Primocandelabrum hiemaloranum, Charnia masoni, Blackbrookia sp. (80332) Hiemalora stellaris, Primocandelabrum sp., Primocandelabrum hiemaloranum, Charnia masoni, Charnia antecedens, Parviscopa bonavistensis, Bradgatia linfordensis, Fractofusus misrai (80350) Primocandelabrum sp., Charnia masoni, Bradgatia linfordensis (80342) Primocandelabrum sp., Charnia masoni, Charnia antecedens, Parviscopa bonavistensis, Bradgatia linfordensis, Fractofusus misrai (80348) Rangeomorpha indet., Charnia sp., Charnia masoni, Bradgatia linfordensis, Avalofractus abaculus, Beothukis mistakensis, Trepassia wardae (90710)
Ediacaran635.0 - 541.0Canada (British Columbia) Bradgatia sp. (231439)
Ediacaran635.0 - 541.0Russian Federation Brachina sp. (51906) Burykhia hunti (237205) Charnia sp. (51905) Hiemalora sp., Charnia sp. (51904) Podolimirus sp. (10759)
Ediacaran635.0 - 541.0Australia (South Australia) Brachina delicata (10280 10286) Eoandromeda sp., Plexus sp. (237281) Funisia dorothea, Eoandromeda octobrachiata (89838 237220) Funisia sp. (191896) Plexus ricei (237282) Plexus sp. (237279 237280) Rangea grandis (89842) Rangeomorpha plumes, Charnia sp., Rangea sp. (191904) Rangeomorpha plumes, Charnia sp., Rangea sp., Funisia dorothea, Eoandromeda octobrachiata (191905) Rangeomorpha plumes, Rangea sp. (191906)
Ediacaran635.0 - 541.0Norway (Finnmark Fylke) Hiemalora sp. (10248)
Ediacaran635.0 - 541.0Canada (Newfoundland) Bradgatia sp., Frondophyllas grandis, Charnia sp. B (207806) Charnia sp. A, Frondophyllas grandis, Charnia sp. B (207807) Hiemalora sp., Primocandelabrum sp., Primocandelabrum hiemaloranum, Charnia masoni, Bradgatia sp., Pectinifrons abyssalis, Frondophyllas grandis, Hapsidophyllas flexibilis, Culmofrons plumosa (228550)
Ediacaran635.0 - 541.0Ukraine (Podolia) Finkoella ukrainica, Finkoella oblonga (236909) Finkoella ukrainica, Finkoella oblonga, Pharyngomorpha reticulata (236907)
Ediacaran635.0 - 541.0USA (Nevada) Wutubus annularis (223400) Wutubus sp. (231326)
Ediacaran635.0 - 541.0China (Guizhou) Eoandromeda octobrachiata (91632)
Ediacaran635.0 - 541.0Canada (Northwest Territories) Hiemalora pleiomorphus (10432) Hiemalora sp., Primocandelabrum hiemaloranum, Charnia masoni, Fractofusus andersoni, Beothukis mistakensis (231368)
Ediacaran635.0 - 541.0Brazil (Ceará) Pectinifrons abyssalis (236885)
Ediacaran635.0 - 541.0Canada (Nova Scotia) Charnia masoni, Trepassia wardae (169292) Trepassia wardae (236917) Vinlandia antecedens (236916)
Poundian570.0 - 538.8Canada (Newfoundland and Labrador) Charnia masoni (10285)
Early/Lower Cambrian538.8 - 509.0China (Yunnan) Batofasciculus ramificans (225161) Batofasciculus ramificans, Galeactena hemispherica (225156) Galeactena hemispherica (225157)
Meishucunian530.7 - 521.0China (Shaanxi) Shufangtubulus sp. (107774)
Atdabanian521.0 - 516.0China (Yunnan) Batofasciculus ramificans, Maotianoascus octonarius, Gemmactena actinala (225162) Dinomischus sp., Batofasciculus sp. (86929) Xianguangia sinica, Trigoides aclis, Calathites spinalis (68083) Xianguangia sp., Dinomischus sp. (86930) Yunnanoascus haikouensis (96869)
Nangaoian521.0 - 514.0China (Hubai) Dinomischus sp., Xianguangia sinica (219687)
Nangaoian521.0 - 514.0China (Yunnan) Dinomischus isolatus (132330 194163)
Stage 3521.0 - 514.0China (Yunnan) Batofasciculus ramificans, Maotianoascus octonarius, Trigoides aclis, Gemmactena actinala, Thaumactena ensis (225155) Calathites spinalis (235227 235228)
Delamaran511.2 - 506.5Canada (British Columbia) Ctenorhabdotus sp. (84418 84422 84427) Dinomischus isolatus (121959) Dinomischus sagittiformis (84426 84428)
Middle Cambrian509.0 - 497.0Canada (Brithish Columbia) Dinomischus sp. (188661)
Middle Cambrian509.0 - 497.0Canada (British Columbia) Ctenophora indet. (171308)
Middle Cambrian509.0 - 497.0USA (Utah) Thalassostaphylos elegans, Ctenorhabdotus campanelliformis (221991)
Menevian504.5 - 499.5USA (Utah) Ctenophora gen et sp. indet. A (230681)
Early/Lower Emsian409.1 - 402.5Germany Ctenophora indet. (26904)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0Brazil (São Paulo) Acanthocephala indet. (57000)