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Ptychagnostidae

Artiopoda - Agnostida - Ptychagnostidae

Synonymy list
YearName and author
1939Ptychagnostidae Kobayashi
1996Ptychagnostidae Westrop et al. pp. 812, 816
1997Ptychagnostidae Whittington et al. p. 350
2003Ptychagnostidae Hong et al.
2017Ptychagnostidae Naimark and Pegel p. 1177

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
RankNameAuthor
classArtiopodaHou and Bergstrom
orderAgnostidaSalter 1864
suborderAgnostinaSalter 1864
superfamilyCondylopygoideaRaymond 1913
familyPtychagnostidaeKobayashi 1939
familyPtychagnostidaeKobayashi 1939

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Ptychagnostidae Kobayashi 1939
show all | hide all
G. †Acidusus Opik 1979
+
†Acidusus acidusus Öpik 1979
†Acidusus aculeatus Angelin 1851
†Acidusus cassis Öpik 1961
†Acidusus leptus Qian 1983
+
Invalid names: Ptychagnostus michaeli Robison 1984 [synonym]
†Acidusus occultatus Öpik 1979
G. †Allobodochus Opik 1979
G. †Aotagnostus Opik 1979
+
†Aotagnostus aotus Öpik 1979
†Aotagnostus culminosus Öpik 1979
+
Invalid names: Aotagnostus protentus Öpik 1979 [synonym]
†Aotagnostus lundgreni Tullberg 1880
+
Invalid names: Triplagnostus longitriangularis Lin 1991 [synonym]
†Aotagnostus magniceps Öpik 1979
†Aotagnostus minutus Lu and Lin 1989
G. †Goniagnostus Howell 1935
+
Subg. †Goniagnostus (Criotypus) Opik 1979
+
†Criotypus oxytorus Öpik 1979
†Goniagnostus fumicola Opik 1961
+
Invalid names: Goniagnostus nodibundus Opik 1967 [synonym]
+
Invalid names: Goniagnostus paenerugatus Öpik 1979 [synonym]
†Goniagnostus nathorsti Brogger 1878
†Goniagnostus scarabaeus Whitehouse 1939
†Goniagnostus spiniger Westergard 1931
†Goniagnostus venustus Zhou and Yang 1977
G. †Lejopyge Hawle and Corda 1847
+
†Lejopyge acantha Robison 1984
†Lejopyge armata Linnarsson 1869
†Lejopyge barrandei Hicks 1872
†Lejopyge cos Opik 1967
†Lejopyge elegans Tullberg 1880
†Lejopyge laevigatus Dalman 1828
+
Invalid names: Lejopyge calva Robison 1964 [synonym]
†Lejopyge multifora Opik 1979
†Lejopyge rigbyi Robison 1984
†Lejopyge sinensis Lu and Lin 1987
Invalid names: Miagnostus Jaekel 1909 [objective synonym]
G. †Myrmecomimus Öpik 1979
+
†Myrmecomimus saltus Öpik 1979
†Myrmecomimus solitarius Ergaliev and Ergaliev 2008
†Myrmecomimus tribulis Öpik 1979
G. †Onymagnostus Opik 1979
+
†Onymagnostus convexus Ergaliev and Ergaliev 2008
†Onymagnostus cuneiformis Ergaliev and Ergaliev 2008
†Onymagnostus hybridus Broegger 1878
†Onymagnostus mundus Öpik 1979
†Onymagnostus ovaliformis Ergaliev and Ergaliev 2008
†Onymagnostus rotundatus Ergaliev 1980
†Onymagnostus semiermis Öpik 1979
†Onymagnostus stenorrhachis Grönwall 1902
Invalid names: Agnostonymus Öpik 1979 [synonym]
G. †Pseudophalacroma Pokrovskaya 1958
+
†Pseudophalacroma dubium Whitehouse 1936
+
Invalid names: Pseudophalacroma crebra Pokrovskaya 1958 [synonym]
+
Invalid names: Pseudophalacroma triangularis Ju 1983 [synonym]
†Pseudophalacroma scanense Westergard 1946
G. †Ptychagnostus Jaekel 1909
+
Subg. †Ptychagnostus (Acidusus) Opik 1979
+
Subg. †Ptychagnostus (Ptychagnostus) Jaekel 1909
†Ptychagnostus affinis Brøgger 1878
†Ptychagnostus akanthodes Robison 1964
†Ptychagnostus aksayensis Ergaliev and Ergaliev 2008
†Ptychagnostus atavus Tullberg 1880
†Ptychagnostus convexus Westergard 1946
†Ptychagnostus cuyanus Rusconi 1948
†Ptychagnostus intermedius Tullberg 1880
†Ptychagnostus karatauensis Ergaliev 1980
†Ptychagnostus punctuosus Angelin 1851
†Ptychagnostus seminula Whitehouse 1939
†Ptychagnostus spinosus Ergaliev 1980
Invalid names: Aristarius Öpik 1979 [synonym], Canotagnostus Rusconi 1951 [synonym], Zeteagnostus Opik 1979 [synonym]
G. †Tomagnostus Howell 1935
+
†Tomagnostus bothrus Robison 1994
†Tomagnostus fissus Lundgren 1879
†Tomagnostus perrugatus Grönwall 1902
†Tomagnostus sibiricus Pokrovskaya and Jegorova 1972
G. †Triplagnostus Howell 1935
+
†Triplagnostus gibbus Linnarsson 1869
†Triplagnostus scopus Öpik 1979
†Triplagnostus ultimus Öpik 1979
G. †Yakutiana Özdikmen 2009
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Entire body: yesf
Adult length: 1.0 to < 10f
Adult width: 1.0 to < 10f
Locomotion: slow-movingo
Life habit: epifaunalo
Diet: detritivoreo
Vision: blindf
Comments: BODY SIZE: Based on component genera; 3 of 3 genera share 10 to < 100 length, 3 of 3 share 10 to < 100 width.f
Created: 2009-06-11 18:42:48
Modified: 2009-12-13 04:44:17
Source: f = family, o = order
References: Kiessling 2004, Hendy 2009

Age range: base of the Solvan to the top of the Arcadian or 508.20000 to 495.90000 Ma

Collections (364 total)


Time interval Ma Country or state Original ID and collection number
Cambrian538.8 - 486.85Argentina (Mendoza) Ptychagnostus sp., Ptychagnostus intermedius (200107)
Cambrian - Tremadocian538.8 - 477.1Norway (Haukelifjell) Lejopyge armata (192870)
Branchian516.2 - 506.5United Kingdom (England) Ptychagnostus triangulatus, Tomagnostus corrugatus, Tomagnostus gracilis, Tomagnostus fissus, Ptychagnostus barrandei (225240)
Delamaran509.7 - 505.3USA (Utah) Ptychagnostus intermedius, Ptychagnostus gibbus (10566)
Amgan508.2 - 504.5Russian Federation Triplagnostus anabarensis (235430)
Amgan508.2 - 504.5Kazakhstan Onymagnostus cuneiformis (9970 9973 9976 9977) Tomagnostus fissus (9971)
Amgan508.2 - 504.5Russian Federation (Siberia) Ptychagnostus gibbus (10002)
Amgan508.2 - 504.5USA (Utah) Ptychagnostus seminula, Ptychagnostus intermedius, Ptychagnostus gibbus (225065 225070)
Solvan508.2 - 504.5Canada (Newfoundland and Labrador) Onymagnostus seminula (10819 10827 10832) Onymagnostus seminula, Ptychagnostus gibbus (10836) Tomagnostus sp. (10816 10822 10831 10850) Tomagnostus sp., Onymagnostus seminula, Ptychagnostus intermedius (10826)
Solvan508.2 - 504.5Norway Ptychagnostus punctuosus, Ptychagnostus gibbus (155475)
Solvan - Menevian508.2 - 499.5Czechia Ptychagnostus vinicensis (73529)
Taijangian506.5 - 504.5China (Sichuan) Ptychagnostus atavus (127059 127060) Ptychagnostus atavus, Ptychagnostus leptus (127058)
Taijangian506.5 - 504.5China (Zhejiang) Ptychagnostus sp., Ptychagnostus atavus, Ptychagnostus sinicus (125682) Triplagnostus gibbus (125536)
Taijangian506.5 - 504.5China (Hunan) Ptychagnostus affinis (49154) Ptychagnostus atavus (49150 49151 49153 49269 49270 49271 49272) Ptychagnostus atavus, Ptychagnostus affinis (49155 49268) Ptychagnostus atavus, Ptychagnostus cuyanus (49156 49159) Ptychagnostus atavus, Ptychagnostus cuyanus, Ptychagnostus gibbus (49158 49273) Ptychagnostus atavus, Ptychagnostus gibbus (49149)
Taijangian506.5 - 504.5China (Liaoning) Ptychagnostus sinicus (126817)
Wuliuan506.5 - 504.5Sweden (Jämtland) Triplagnostus gibbus (240064)
Middle Cambrian506.5 - 497.0USA (Alaska) Lejopyge calva (64770) Lejopyge laevigata (64768) Lejopyge sp. (64769) Ptychagnostus aculeatus, Lejopyge laevigata (64772) Ptychagnostus sp. (64766)
Middle Cambrian506.5 - 497.0Czechia (Jince) Onymagnostus hybridus (76482) Onymagnostus hybridus, Tomagnostus perrugatus (76481)
Middle Cambrian506.5 - 497.0USA (Utah) Ptychagnostus atavus (86743 156506) Ptychagnostus atavus, Ptychagnostus gibbus (86739 86741) Ptychagnostus atavus, Ptychagnostus seminula (156504) Ptychagnostus atavus, Ptychagnostus seminula, Ptychagnostus intermedius (156505) Ptychagnostus gibbus (86729 86731 86732 86733 86737 86738 86742) Ptychagnostus intermedius (156507) Ptychagnostus intermedius, Ptychagnostus gibbus (86728 86730 86735 86736) Ptychagnostus richmondensis (77269 77270 77277)
Middle Cambrian506.5 - 497.0USA (Nevada) Ptychagnostus atavus (156503) Ptychagnostus atavus, Ptychagnostus intermedius (156502) Ptychagnostus atavus, Ptychagnostus seminula, Ptychagnostus intermedius (156500 156501)
Middle Cambrian506.5 - 497.0Canada (Newfoundland and Labrador) Ptychagnostus atavus (9682)
Middle Cambrian506.5 - 497.0Greenland Lejopyge barrandei (10475 10477 10485) Ptychagnostus affinis, Lejopyge barrandei, Tomagnostus perrugatus (10484)
Middle Cambrian506.5 - 497.0Argentina (Mendoza) Ptychagnostus aculeatus, Lejopyge laevigata (9811) Ptychagnostus cuyanus (9815 9816)
Middle Cambrian506.5 - 497.0India Goniagnostus sp., Lejopyge sp., Lejopyge armata, Ptychagnostus aculeatus, Lejopyge acantha (114635) Lejopyge armata, Ptychagnostus aculeatus (114634) Ptychagnostus aculeatus, Lejopyge acantha, Pseudophalacroma ovale (114665)
Middle Cambrian506.5 - 497.0Italy (Iglesiente) Ptychagnostus sp. (139880)
St Davids506.5 - 497.0United Kingdom (England) Agnostus lundgreni (55509)
St Davids506.5 - 497.0North Korea Lejopyge armata (49082)
St Davids506.5 - 497.0USA (Utah) Ptychagnostus sp. (216)
St Davids506.5 - 497.0USA (Montana) Tomagnostus sp., Ptychagnostus atavus (212)
St Davids506.5 - 497.0Canada (British Columbia) Ptychagnostus burgessensis (215) Ptychagnostus sp., Lejopyge calva (122017)
St Davids506.5 - 497.0Greenland Lejopyge seminula, Tomagnostus gibbus, Tomagnostus fissus, Ptychagnostus intermedius, Phalagnostus ovalis, Lejopyge barrandei, Tomagnostus bothrus (10478)
Taijangian - Guzhangian506.5 - 497.0China (Anhui) Goniagnostus sp., Triplagnostus sp. (127474)
Taijangian - Wangcunian506.5 - 497.0China (Henan) Goniagnostus sp., Ptychagnostus sp. (127073)
Arcadian506.5 - 495.9Canada (Newfoundland and Labrador) Ptychagnostus fissus (9689) Ptychagnostus hybridus (9635) Ptychagnostus seminula, Ptychagnostus fissus (9690)
Arcadian506.5 - 495.9Canada (New Brunswick) Lejopyge hybridus, Ptychagnostus atavus, Tomagnostus perrugatus (64540)
Tuesaian505.3 - 503.0Kazakhstan Ptychagnostus affinis (118091) Ptychagnostus atavus (118088 118090 118094) Ptychagnostus atavus, Ptychagnostus affinis (118092 118093) Ptychagnostus atavus, Ptychagnostus intermedius, Ptychagnostus karatauensis (118086) Ptychagnostus atavus, Ptychagnostus karatauensis (118087) Ptychagnostus intermedius (9966 9967) Ptychagnostus intermedius, Ptychagnostus affinis (9969) Ptychagnostus intermedius, Ptychagnostus convexus, Onymagnostus rotundatus, Onymagnostus convexus, Onymagnostus cuneiformis, Onymagnostus ovaliformis, Triplagnostus gibbus, Tomagnostus sibiricus (118085) Ptychagnostus intermedius, Ptychagnostus convexus, Onymagnostus rotundatus, Onymagnostus convexus, Onymagnostus ovaliformis, Triplagnostus gibbus, Tomagnostus sibiricus (118054) Ptychagnostus punctuosus, Ptychagnostus atavus, Ptychagnostus karatauensis (118089) Tomagnostus fissus (9968)
Floran504.5 - 503.0Australia (Queensland) Aotagnostus aotus (225225) Ptychagnostus atavus, Ptychagnostus intermedius (90972) Triplagnostus diremptus, Onymagnostus semiermis, Criotypus lemniscatus (225224) Triplagnostus fretus, Triplagnostus posterus, Triplagnostus stramineus, Triplagnostus quasigibbus, Ptychagnostus atavus, Zeteagnostus incautus, Triplagnostus scopus, Criotypus paenerugatus, Aristarius retrocornutus (225241) Triplagnostus posterus, Triplagnostus sp. indet., Ptychagnostus atavus, Zeteagnostus incautus, Criotypus oxytorus, Criotypus lemniscatus, Criotypus paenerugatus (225175)
Floran504.5 - 503.0Australia (Northern Territory) Criotypus sp., Onymagnostus sp., Zeteagnostus sinicus, Triplagnostus gibbus (225075)
Drumian504.5 - 500.5Poland Ptychagnostus gibbus (72272)
Drumian504.5 - 500.5Sweden (Jämtland) Goniagnostus nathorsti, Lejopyge calva (240067) Ptychagnostus affinis (240065 240066)
Drumian504.5 - 500.5Canada (New Brunswick) Onymagnostus hybridus (240657) Onymagnostus hybridus, Tomagnostus perrugatus (240658) Tomagnostus perrugatus (240659)
Drumian504.5 - 500.5Sweden Goniagnostus nathorsti, Lejopyge laevigata (240063) Goniagnostus nathorsti, Lejopyge laevigata, Acidusus aculeatus (240062)
Drumian504.5 - 500.5Norway Ptychagnostus atavus (155478) Ptychagnostus sp. (155476)
Floran - Undillan504.5 - 500.5Antarctica Ptychagnostus punctuosus (69520)
Floran - Undillan504.5 - 500.5Australia (Queensland) Onymagnostus angulatus, Aotagnostus aotus (225217)
Mayan504.5 - 499.5Kazakhstan (Balkhash) Lejopyge laevigata (118218)
Mayan504.5 - 499.5Russian Federation (Altai (Region)) Lejopyge avius, Lejopyge laevigata (10060) Lejopyge laevigata (10062)
Mayan504.5 - 499.5Russian Federation (Gorniy Altay) Goniagnostus sp. (10173)
Mayan504.5 - 499.5Russian Federation (Sakha) Pseudophalacroma crebra (225247)
Mayan504.5 - 499.5Sweden (Scania) Goniagnostus spiniger (225259)
Mayan504.5 - 499.5Russian Federation (Gorniy Altay Republic) Lejopyge sp. (10175)
Mayan504.5 - 499.5Russian Federation Lejopyge armata, Lejopyge laevigata (10118) Pseudophalacroma crebra (225248)
Menevian504.5 - 499.5United Kingdom (Wales) Ptychagnostus ciceroides, Ptychagnostus punctuosus (193783) Ptychagnostus davidis, Ptychagnostus ciceroides, Ptychagnostus punctuosus (193782) Ptychagnostus granulatum, Tomagnostus fissus (198942)
Menevian504.5 - 499.5Denmark (Bornholm) Ptychagnostus punctuosus, Tomagnostus perrugatus (75946) Ptychagnostus subsp. indet., Ptychagnostus punctuosus (75947)
Menevian504.5 - 499.5USA (Utah) Ptychagnostus aculeatus, Lejopyge calva, Ptychagnostus cassis (10572) Ptychagnostus atavus (10567) Ptychagnostus gibbus (10507)
Menevian504.5 - 499.5Sweden (Scania) Acidusus atavus, Onymagnostus seminula, Tomagnostus fissus, Tomagnostus perrugatus (176679) Acidusus sp., Onymagnostus sp., Acidusus atavus, Tomagnostus fissus, Phalagnostus ovalis, Onymagnostus hybridus, Onymagnostus mundus, Ptychagnostus affinis, Tomagnostus perrugatus (179691) Ptychagnostus punctuosus, Ptychagnostus elegans, Ptychagnostus stenorrhachis, Ptychagnostus lundgreni (225250)
Menevian504.5 - 499.5Greenland (Nyeboe Land) Ptychagnostus ciceroides, Ptychagnostus punctuosus (9935)
Drumian - Marjumian504.5 - 497.0Mexico Ptychagnostus sp., Ptychagnostus atavus (236876)
Marjumian504.5 - 497.0Greenland (Peary Land) Lejopyge armata (49483) Lejopyge laevigata (49435 49449 49454)
Marjumian504.5 - 497.0Canada (Newfoundland and Labrador) Ptychagnostus atavus (9681) Ptychagnostus fissus (9632 9633 9680) Ptychagnostus hybridus (9673) Ptychagnostus seminula (9684 9687 9691)
Marjumian504.5 - 497.0USA (New York) Ptychagnostus punctuosus, Ptychagnostus elegans (221)
Marjumian504.5 - 497.0USA (Utah) Lejopyge hybridus, Ptychagnostus punctuosus, Ptychagnostus atavus, Pseudophalacroma rigbyi, Ptychagnostus cuyanus, Ptychagnostus leptus, Pseudophalacroma lundgreni, Ptychagnostus occultatus, Ptychagnostus affinis, Ptychagnostus akanthodes (230679) Lejopyge seminula, Ptychagnostus atavus, Ptychagnostus cuyanus, Ptychagnostus gibbus, Ptychagnostus sinicus, Ptychagnostus occultatus (230682) Ptychagnostus akanthodes (225239) Ptychagnostus germanus (77207) Ptychagnostus punctuosus, Lejopyge lundgreni, Lejopyge rigbyi (225249) Ptychagnostus punctuosus, Ptychagnostus occultatus (77206) Ptychagnostus sp. (56144)
Marjumian504.5 - 497.0USA (South Carolina) Ptychagnostus sp., Tomagnostus fissus (75062 75063)
Marjumian504.5 - 497.0Canada (British Columbia) Ptychagnostus sp., Lejopyge calva (122072)
Wangcunian504.5 - 497.0China (Henan) Goniagnostus sp., Phalagnostus ovalis (127074)
Wangcunian504.5 - 497.0China (Jiangsu) Triplagnostus sp. (127368)
Wangcunian504.5 - 497.0China (Zhejiang) Goniagnostus spiniger, Pseudophalacroma dubium, Pseudophalacroma triangularis (125539) Pseudophalacroma dubium, Pseudophalacroma triangularis (125538) Pseudophalacroma triangularis (125532) Ptychagnostus atavus (125537) Ptychagnostus sp. (125531)
Wangcunian504.5 - 497.0China (Hunan) Goniagnostus longispinus, Pseudophalacroma triangularis (128922) Goniagnostus nathorsti, Lejopyge calva (128336 128339) Goniagnostus sp., Goniagnostus fumicola (128923) Lejopyge armata (49291 128341) Lejopyge armata, Goniagnostus nathorsti (128340) Lejopyge armata, Lejopyge calva (128333) Lejopyge armata, Pseudophalacroma ovale (49165) Lejopyge calva (128334 128335 128338) Pseudophalacroma dubium (49281 128317) Pseudophalacroma ovale (49164 49285 49286 128318 128319 128331 128332) Pseudophalacroma ovale, Goniagnostus nathorsti (49163 49289 49290) Pseudophalacroma ovale, Goniagnostus nathorsti, Pseudophalacroma dubium, Pseudophalacroma lundgreni, Pseudophalacroma scanense (49283) Pseudophalacroma ovale, Lejopyge calva (49288) Pseudophalacroma ovale, Pseudophalacroma dubium (49162) Pseudophalacroma ovale, Pseudophalacroma lundgreni (49284) Ptychagnostus aculeatus (128337) Ptychagnostus atavus (128330) Ptychagnostus atavus, Pseudophalacroma triangularis, Triplagnostus longitriangularis (128921) Ptychagnostus leptus (49277 49278 49280) Ptychagnostus punctuosus, Pseudophalacroma dubium, Goniagnostus scarabaeus (49282) Ptychagnostus punctuosus, Ptychagnostus atavus (49275) Ptychagnostus punctuosus, Ptychagnostus atavus, Pseudophalacroma ovale, Ptychagnostus cuyanus, Pseudophalacroma dubium, Ptychagnostus leptus (49161) Ptychagnostus punctuosus, Ptychagnostus cuyanus (49274)
Wangcunian - Guzhangian504.5 - 497.0China (Henan) Ptychagnostus aculeatus (127076)
Wangcunian - Guzhangian504.5 - 497.0China (Zhejiang) Lejopyge armata (125685) Lejopyge armata, Goniagnostus spiniger (125684) Lejopyge armata, Goniagnostus spiniger, Aotagnostus minutus (125535 125541) Lejopyge armata, Pseudophalacroma sp., Goniagnostus spiniger, Pseudophalacroma dubium (125554) Lejopyge armata, Ptychagnostus aculeatus, Goniagnostus spiniger (125540 125683)
Wangcunian - Guzhangian504.5 - 497.0China (Hubei) Goniagnostus nathorsti (130258)
Wangcunian - Guzhangian504.5 - 497.0China (Hunan) Goniagnostus fumicola (128926 128927) Goniagnostus venustus (128928) Lejopyge armata (128925)
Wangcunian - Guzhangian504.5 - 497.0China (Sichuan) Lejopyge laevigata (127067)
Undillan503.0 - 500.5Australia (Northern Territory) Ptychagnostus punctuosus, Aotagnostus culminosus (225215)
Undillan503.0 - 500.5Australia (Queeland) Ptychagnostus punctuosus, Acidusus occultatus (225234)
Undillan503.0 - 500.5Australia (Queensland) Acidusus navus, Ptychagnostus punctuosus, Goniagnostus nathorsti, Acidusus germanus (225238) Acidusus occultatus, Myrmecomimus tribulis (225233) Aotagnostus aotus, Aristarius ultimus, Myrmecomimus saltus (225246) Aotagnostus culminosus (225219) Aotagnostus ponebrevis, Triplagnostus diremptus, Onymagnostus semiermis, Criotypus lemniscatus (225229) Aotagnostus protentus, Myrmecomimus tribulis (225242) Goniagnostus nathorsti, Acidusus occultatus, Aristarius ultimus (225232) Myrmecomimus mixtus, Aotagnostus culminosus, Ptychagnostus affinis, Acidusus acidusus (225237) Myrmecomimus sp. indet., Ptychagnostus punctuosus, Aotagnostus culminosus, Aotagnostus protentus (225222) Myrmecomimus tribulis (225244) Onymagnostus angulatus, Onymagnostus semiermis, Criotypus lemniscatus (225228) Onymagnostus angulatus, Ptychagnostus mesostatus, Ptychagnostus punctuosus, Goniagnostus nathorsti, Aotagnostus aotus (225216) Onymagnostus semiermis, Ptychagnostus scarifatus (225227) Onymagnostus seminula, Aotagnostus magniceps, Myrmecomimus tribulis (225223) Ptychagnostus coartatus, Ptychagnostus atavus, Triplagnostus gibbus, Zeteagnostus incautus (225230) Ptychagnostus idmon, Triplagnostus diremptus, Onymagnostus semiermis, Ptychagnostus scarifatus (225226) Ptychagnostus indet., Aotagnostus protentus, Myrmecomimus saltus (225245) Ptychagnostus indet., Goniagnostus scarabaeus, Aotagnostus culminosus (225220) Ptychagnostus punctuosus, Aotagnostus protentus, Myrmecomimus tribulis (225243) Ptychagnostus punctuosus, Onymagnostus seminula, Aotagnostus aotus (225218) Ptychagnostus punctuosus, Onymagnostus seminula, Aotagnostus culminosus, Aotagnostus protentus, Onymagnostus mundus, Myrmecomimus tribulis (225221)
Zhanaarykian503.0 - 500.5Kazakhstan Goniagnostus nathorsti (118099) Lejopyge armata, Ptychagnostus aculeatus, Pseudophalacroma ovale, Goniagnostus nathorsti, Goniagnostus spiniger, Lejopyge calva, Pseudophalacroma dubium, Myrmecomimus solitarius (9958) Lejopyge armata, Ptychagnostus aculeatus, Pseudophalacroma ovale, Goniagnostus spiniger, Lejopyge calva, Pseudophalacroma dubium, Myrmecomimus solitarius (118102) Pseudophalacroma ovale, Goniagnostus nathorsti, Pseudophalacroma dubium (9961) Pseudophalacroma ovale, Goniagnostus nathorsti, Pseudophalacroma dubium, Onymagnostus stenorrhachis, Zeteagnostus incautus, Onymagnostus hybridus (9962 118097) Ptychagnostus punctuosus, Ptychagnostus cuyanus, Ptychagnostus spinosus, Ptychagnostus acidusus (118096) Ptychagnostus punctuosus, Ptychagnostus cuyanus, Ptychagnostus spinosus, Ptychagnostus acidusus, Ptychagnostus aksayensis (9964) Ptychagnostus punctuosus, Ptychagnostus cuyanus, Ptychagnostus spinosus, Ptychagnostus acidusus, Ptychagnostus aksayensis, Ptychagnostus affinis (118095) Ptychagnostus spinosus, Ptychagnostus acidusus, Ptychagnostus aksayensis, Goniagnostus nathorsti, Zeteagnostus incautus (9963)
Boomerangian500.5 - 499.0Australia (Queenslandd) Ptychagnostus sp. P aff. nathorsti, Ptychagnostus sp. O, Ptychagnostus aculeatus, Pseudophalacroma sp. K, Lejopyge laevigata, Pseudophalacroma dubium (225252)
Boomerangian500.5 - 499.0Australia (Queensland) Ptychagnostus cassis (225231)
Boomerangian500.5 - 499.0Mexico Ptychagnostus sp., Ptychagnostus atavus, Onymagnostus hybridus (200299)
Boomerangian500.5 - 499.0New Zealand (Nelson) Pseudophalacroma sp., Goniagnostus spiniger (7897)
Guzhangian500.5 - 497.0Norway (Hedmark) Lejopyge laevigata (192873 192874)
Guzhangian500.5 - 497.0Antarctica Lejopyge sp., Lejopyge armata, Ptychagnostidae indet., Goniagnostus nathorsti, Lejopyge calva, Onymagnostus hybridus, Acidusus aculeatus (227789)
Guzhangian500.5 - 497.0China (Hunan) Goniagnostus fumicola (49179 126310 126315 126317 126318 126321 126322) Goniagnostus fumicola, Lejopyge sinensis (49178 126313 126314 126316) Goniagnostus nathorsti (128348) Goniagnostus spiniger (49168 49303 128361) Goniagnostus spiniger, Lejopyge laevigata (49169 49176) Goniagnostus spiniger, Lejopyge multifora (49301 126287) Lejopyge armata (49293 49294 49295 49296 49299 49300 128345 128346 128347 128350 128353 128357 128362 128363) Lejopyge armata, Goniagnostus fumicola, Lejopyge sinensis (128929) Lejopyge armata, Goniagnostus nathorsti (128343) Lejopyge armata, Goniagnostus spiniger (128356 128359) Lejopyge armata, Goniagnostus spiniger, Lejopyge laevigata (126304 128351 128354) Lejopyge armata, Goniagnostus venustus (128930) Lejopyge armata, Lejopyge laevigata (49292 49298 128365) Lejopyge armata, Lejopyge laevigata, Lejopyge calva (49166) Lejopyge armata, Ptychagnostus atavus (128344) Lejopyge laevigata (49170 49173 49174 128342 128366 128367) Lejopyge laevigata, Goniagnostus fumicola, Lejopyge sinensis (126312) Ptychagnostus aculeatus, Lejopyge laevigata (49172)
Guzhangian500.5 - 497.0Sweden (Jämtland) Goniagnostus nathorsti, Lejopyge laevigata (240069) Goniagnostus nathorsti, Lejopyge laevigata, Acidusus aculeatus (240068)
Guzhangian500.5 - 497.0China (Zhejiang) Lejopyge sinensis, Goniagnostus nodibundus (125542 125674)
Guzhangian500.5 - 497.0China (Sichuan) Lejopyge laevigata (127068 127070)
Ayusokkanian499.5 - 497.0Kazakhstan Lejopyge armata, Lejopyge laevigata, Goniagnostus fumicola (118177) Lejopyge armata, Ptychagnostus aculeatus (9943 9944 9945 9947 118166 118168) Lejopyge armata, Ptychagnostus aculeatus, Goniagnostus spiniger, Lejopyge calva (118164 118165) Lejopyge armata, Ptychagnostus aculeatus, Pseudophalacroma ovale, Goniagnostus spiniger, Lejopyge calva, Pseudophalacroma dubium (118158) Lejopyge armata, Ptychagnostus aculeatus, Pseudophalacroma ovale, Goniagnostus spiniger, Lejopyge laevigata, Lejopyge calva, Pseudophalacroma dubium, Myrmecomimus solitarius (9950) Lejopyge laevigata (9949) Lejopyge laevigata, Goniagnostus fumicola (118178 118181 118182 118186) Lejopyge laevigata, Lejopyge cos, Goniagnostus fumicola (118179 118180)
Dresbachian497.0 - 486.85Canada (Newfoundland and Labrador) Ptychagnostus aculeatus (9692) Ptychagnostus atavus (9703) Ptychagnostus atavus, Ptychagnostus fissus (9702)
Furongian497.0 - 486.85Norway (Telemark) Lejopyge armata (192871)
Merioneth497.0 - 486.85USA (Vermont) Ptychagnostus sp. (217)