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Gigantochersina ammon
Taxonomy
Testudo ammon was named by Andrews (1903). Its type specimen is CGM 8130, a partial skeleton (nearly complete carapace and plastron), and it is a 3D body fossil. Its type locality is NNW of Birket Qarun, which is in an Eocene/Oligocene fluvial horizon in the Jebel Qatrani Formation of Egypt.
It was recombined as Geochelone (Geochelone) ammon by Auffenberg (1974); it was recombined as Geochelone ammon by Crumly (1982); it was recombined as Gigantochersina ammon by Holroyd and Parham (2003), Vlachos and Rabi (2018), Georgalis et al. (2021).
It was recombined as Geochelone (Geochelone) ammon by Auffenberg (1974); it was recombined as Geochelone ammon by Crumly (1982); it was recombined as Gigantochersina ammon by Holroyd and Parham (2003), Vlachos and Rabi (2018), Georgalis et al. (2021).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1903 | Testudo ammon Andrews |
| 1906 | Testudo ammon Andrews p. 278 |
| 1906 | Testudo beadnelli Andrews p. 285 |
| 1974 | Geochelone (Geochelone) ammon Auffenberg p. 154 |
| 1974 | Geochelone (Geochelone) beadnelli Auffenberg p. 154 |
| 1982 | Geochelone ammon Crumly p. 215 |
| 1982 | Geochelone beadnelli Crumly p. 215 |
| 2003 | Gigantochersina ammon Holroyd and Parham p. 688 |
| 2018 | Gigantochersina ammon Vlachos and Rabi p. 663 |
| 2021 | Gigantochersina ammon Georgalis et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Gigantochersina ammon Andrews 1903
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Invalid names: Testudo beadnelli Andrews 1906 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| G. L. Georgalis et al. 2021 | Gigantochersina ammon can be diagnosed as a member of Testudinidae and Geochelona by the full list of shell characters provided for these clades above. Gigantochersina ammon can be differentiated from other testudinids by its large size (carapace length around 90 cm), highly arched carapace, wide nuchal without posterior constriction, presence of seven neurals, neural I hexagonal, presence of a cervical scute, absence of contact of marginals IV with pleurals II, rather prominent gular protrusion of the shell (at least in male individuals), paired gulars, humero-pectoral sulcus in contact with the posterior margin of the entoplastron, narrow pectorals, rather elongated epiplastra, presence of a prominent epiplastral excavation, humeral equal or shorter than gulars, posterior plastral lobe with subparallel lateral margins across their whole layout (with the exception of a well-developed change of curvature in the region where they contact with the femoro-anal sulci [i.e., prominent notch at the femoral-anal sulcus in males and not so prominent in females]), short xiphiplastra, presence of a wide and shallow anal notch, and plastral bridge small (up to equal to half the length of the plastron). |
Measurements
No measurements are available
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| Source: f = family, c = class, subp = subphylum | |||||
| References: Ernst and Barbour 1989, Carroll 1988, Hendy et al. 2009 | |||||
Collections (3 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Eocene - Early/Lower Oligocene | Egypt (Fayum) | Testudo ammon, Testudo beadnelli (65960) | |
| Rupelian | Egypt (Fayum) | Gigantochersina ammon (166979) | |
| Zanclean | Kenya | Testudo ammon (22234) |