PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Haptophyta (haptophyte alga)

Taxonomy

Haptophyta was named by Hibberd ex Cavalier-Smith (1986). It is extant.

It was assigned to Chromista by Young and Bown (1997); and to Chromalveolata by Adl et al. (2005).

Synonymy list

Year	Name and author
1986	Haptophyta Hibberd ex Cavalier-Smith
1997	Haptophyta Young and Bown
2005	Haptophyta Adl et al. p. 401 figs. Table 1

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Rank	Name	Author
—	Life
—	Eukaryota	()
—	Chromalveolata

Rank	Name	Author
phylum	Haptophyta	Hibberd ex Cavalier-Smith 1986
phylum	Haptophyta	Hibberd ex Cavalier-Smith 1986

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Haptophyta Hibberd ex Cavalier-Smith 1986 [haptophyte alga]

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G. Aspidorhabdus Hay and Towe 1962

G. †Camuralithus de Kaenel and Villa 1996

Cl. Coccolithophyceae Rothmaler 1951

G. †Athenagalea Hattner and Wise 1980

G. †Biantholithus Bramlette and Martini 1964

Or. Braarudosphaerales Aubry 2013

Fm. †Polycyclolithaceae Forchheimer 1972

G. †Assipetra Roth 1973

†Assipetra infracretacea Thierstein 1973

†Assipetra terebrodentarius Applegate and Bralower 1987

G. †Eprolithus Stover 1966

†Eprolithus antiquus Perch-Nielsen 1979

†Eprolithus apertior Black 1973

†Eprolithus floralis Stradner 1962

†Eprolithus moratus Stover 1966

†Eprolithus octopetalus Varol 1992

†Eprolithus rarus Varol 1992

Invalid names: Polycyclolilthus Forchheimer 1968 [synonym]

G. †Farhania Varol 1992

†Farhania varolii Jakubowski 1986

G. †Hayesites Manivit 1971

†Hayesites albiensis Manivit 1971

†Hayesites irregularis Thierstein et al. 1972

G. †Hexalithus Gardet 1955

†Hexalithus gardetiae Bukry 1969

†Hexalithus noeliae Loeblich and Tappan 1966

G. †Lithastrinus Stradner 1962

†Lithastrinus grillii Stradner 1962

†Lithastrinus pentabrachius Varol 1992

†Lithastrinus quadricuspis Farhan 1997

†Lithastrinus septenarius Forchheimer 1972

G. †Micula Vekshina 1959 [coccolithophore]

†Micula adumbrata Burnett 1998

†Micula clypeata Lees and Bown 2005

†Micula concava Stradner 1960

†Micula cubiformis Forchheimer 1972

†Micula decussata Vekshina 1959 [coccolithophore]

†Micula murus Martini 1961

†Micula praemurus Bukry 1973

†Micula premolisilvae Lees and Bown 2005

†Micula prinsii Perch-Nielsen 1979

†Micula staurophora Gardet 1955

†Micula swastica Stradner and Steinmetz 1984

Invalid names: Nannotetraster Martini and Stradner 1960 [synonym]

G. †Polycostella Thierstein 1971

†Polycostella beckmanii Thierstein 1971

†Polycostella senaria Thierstein 1971

G. †Quadrum Prins and Perch-Nielsen 1977

†Quadrum bengalensis Burnett 1998

†Quadrum eneabrachium Varol 1992

†Quadrum eptabrachium Varol 1992

†Quadrum gartneri Prins and Perch-Nielsen 1977

†Quadrum giganteum Varol 1992

†Quadrum intermedium Varol 1992

†Quadrum octobrachium Varol 1992

†Quadrum svabenickae Burnett 1998

G. †Radiolithus Stover 1966

†Radiolithus hollandicus Varol 1992

†Radiolithus orbiculatus Forchheimer 1972

†Radiolithus planus Stover 1966

G. †Rucinolithus Stover 1966

†Rucinolithus magnus Bukry 1975

†Rucinolithus windleyae Rutledge and Bown 1996

†Rucinolithus wisei Thierstein 1971

G. †Tegulalithus Crux 1986

†Tegulalithus septentrionalis Stradner 1963

†Tegulalithus tessellatus Stradner 1968

G. †Uniplanarius Hattner and Wise 1980

†Uniplanarius clarkei Lees and Bown 2005

†Uniplanarius gothicus Deflandre 1959

†Uniplanarius sissinghii Perch-Nielsen 1986

†Uniplanarius trifidus Stradner 1961

Fm. Ceratolithaceae Norris 1965

G. Amaurolithus Gartner and Bukry 1975

Amaurolithus primus Bukry and Percival 1971

G. Angulolithina Bukry 1973

G. Ceratolithus Kamptner 1950

G. Florisphaera Okada and Honjo 1973

G. †Nicklithus Raffi et al. 1998

Nicklithus amplificus Bukry and Percival 1971

G. †Orthorhabdus Bramlette and Wilcoxon 1967

Or. †Coccolithales Schwarz 1932

Fm. Coccolithaceae Poche 1913

G. †Bicolumnus Wei and Wise 1990

G. †Birkelundia Perch-Nielsen 1971

G. †Bramletteius Gartner 1969

G. †Campylosphaera Kamptner 1963

G. †Chiasmolithus Hay et al. 1966

†Chiasmolithus bidens Bramlette and Sullivan 1961

†Chiasmolithus consuetus Bramlette and Sullivan 1961

†Chiasmolithus danicus Hay and Mohler 1967

†Chiasmolithus eograndis Perch-Nielsen 1971

†Chiasmolithus expansus Bramlette and Sullivan 1961

†Chiasmolithus grandis Bramlette and Riedel 1954

†Chiasmolithus modestus Perch-Nielsen 1971

†Chiasmolithus oamaruensis Deflandre 1954

†Chiasmolithus solitus Bramlette and Sullivan 1961

G. †Clausicoccus Prins 1979

G. Coronocyclus Hay et al. 1966

G. †Ericsonia Black 1964

G. †Gephyrorhabdus Hill 1976

G. †Holodiscolithus Roth 1970

†Holodiscolithus solidus Deflandre and Fert 1954

G. †Hughesius Varol 1989

G. †Praeprinsius Varol and Jakubowski 1989

G. †Solidopons Theodoridis 1984

Invalid names: Hymenomonadaceae Senn 1900 [empty], Pleurochrysidaceae Fresnel and Billard 1991 [empty]

G. †Daktylethra Gartner and Bukry 1969

Or. †Discoasterales Hay 1977

Fm. †Fasciculithaceae Hay and Mohler 1967

G. †Fasciculithus Bramlette and Sullivan 1961

G. †Lithoptychius Aubry 2011

Fm. †Heliolithaceae Hay and Mohler 1967

G. †Bomolithus Roth 1973

G. †Heliolithus Bramlette and Sullivan 1961

Invalid names: Discoasteraceae Tan Sin Hok 1927 [empty]

Fm. †Ellipsagelosphaeraceae Noël 1965

Subfm. †Actinosphaeroideae Noël 1965

G. †Calolithus Noël 1965

G. †Ellipsolithus Sullivan 1964

G. Gladiolithus Jordan and Chamberlain 1992

G. †Ilselithina Stradner and Adamiker 1966

Or. Isochrysidales Pascher 1910

Fm. Gephyrocapsaceae Heimdal 1997

G. Crenalithus Roth 1973

Fm. Noelaerhabdaceae Jerkovic 1970

G. Apertapetra Hay et al. 1966

G. Cepekiella Roth 1970

G. †Cribrocentrum Perch-Nielsen 1971

G. Dictyococcites Black 1967

†Dictyococcites scrippsae Bukry and Percival 1973

G. Emiliania Hay and Mohler 1967

G. Gephyrocapsa Kamptner 1943

G. †Pseudoemiliania Gartner 1969

Fm. †Prinsiaceae Hay and Mohler 1967

G. †Futyania Varol 1989

G. †Girgisia Varol 1989

G. Hornibrookina Edwards 1973

†Hornibrookina teuriensis Edwards 1973

†Hornibrookina weimerae Self-Trail 2011

G. †Neobiscutum Varol 1989

G. †Prinsius Hay and Mohler 1967

†Prinsius dimorphosus Perch-Nielsen 1977

†Prinsius martinii Haq 1971

†Prinsius tenuiculus Perch-Nielsen 1984

Invalid names: Isochrysidaceae Pascher 1910 [empty]

G. †Lanternithus Stradner 1962

†Lanternithus minutus Stradner 1962

G. †Minylitha Bukry 1973

G. Neosphaera Lecal-Schlauder 1949

G. †Pedinocyclus Bukry and Bramlette 1971

Fm. †Rhomboasteraceae Bown 2005

G. †Rhomboaster Bramlette and Sullivan 1961

G. †Tribrachiatus Shamrai 1963

G. Tetralithoides Theodoridis 1984

G. †Transversopontis Hay et al. 1966

Or. Zygodiscales Young and Bown 1997

Fm. Pontosphaeraceae Lemmermann 1908

G. Calciopilleus Müller 1974

G. Scyphosphaera Lohmann 1902

Fm. Zygodiscaceae Hay and Mohler 1967

G. †Isthmolithus Deflandre and Fert 1954

†Isthmolithus recurvus Deflandre 1954

G. †Lophodolithus Deflandre and Fert 1954

†Lophodolithus mochlophorus Deflandre 1954

G. †Nannotetrina Achuthan and Stradner 1969

G. †Neococcolithes Sujkowski 1931

†Neococcolithes dubius Deflandre and Fert 1954

†Neococcolithes protenus Black 1967

G. †Zygodiscus Bramlette and Sullivan 1961

Invalid names: Helicosphaeraceae Black 1971 [empty]

Invalid names: Phaeocystales Medlin 2000 [empty], Prymnesiales Papenfuss 1955 [empty], Prymnesiophyceae Hibberd 1976 [synonym]

G. Conococcolithus Hay and Mohler 1967

G. Corannulus Stradner 1962

G. Craspedolithus Kamptner 1963

G. Crassapontosphaera Boudreaux and Hay 1969

G. †Crassidiscus Okada 1990

G. †Cricosphaera Braarud 1960

G. Cyclolithella Loeblich and Tappan 1963

G. †Cycloperfolithus Lehotayova and Priewalder 1978

Fm. †Discoasteridae

G. †Agalmatoaster Klumpp 1953

†Agalmatoaster septemradiatus Klumpp 1953

G. †Trochoaster Klumpp 1953

†Trochoaster simplex Klumpp 1953

G. †Duplexipodorhabdus Varol and Girgis 1994

G. Ellipsodiscoaster Boudreaux and Hay 1969

G. †Helicosphaera Kamptner 1954

†Helicosphaera carteri Wallich 1877

†Helicosphaera cateri Wallich 1877

†Helicosphaera ethologa Bown 2005

†Helicosphaera lophota Bramlette and Sullivan 1961

†Helicosphaera neolophota Barron et al. 1984

†Helicosphaera orientalis Black 1971

†Helicosphaera stalis Theodoridis 1984

Invalid names: Helicopontosphaera Hay and Mohler 1967 [synonym]

G. Heliorthus Brönnimann and Stradner 1960

†Heliorthus tenuis Stradner 1961

G. Koczyia Boudreaux and Hay 1969

G. Leptodiscus Bukry and Bramlette 1969

Fm. Lithostromationaceae Deflandre 1959

G. Imperiaster Martini 1970

G. Lithostromation Deflandre 1942

G. Nannocorbis Müller 1974

G. Orthozygus Bramlette and Wilcoxon 1967

G. †Pervilithus Crux 1981

G. †Phanulithus Wise and Wind 1977

G. Pyrocyclus Hay and Towe 1962

G. Reticulofenestra Hay et al. 1966

†Reticulofenestra bisecta Roth 1970

†Reticulofenestra bisecta bisecta Roth 1970

†Reticulofenestra daviesii Haq 1971

†Reticulofenestra dictyoda Stradner 1968

†Reticulofenestra filewiczii Dunkley 2009

†Reticulofenestra hampdenensis Edwards 1973

†Reticulofenestra minuta Roth 1970

†Reticulofenestra oamaruensis Stradner 1968

†Reticulofenestra pseudoumbilicus Gartner 1967

†Reticulofenestra reticulata Roth and Thierstein 1972

†Reticulofenestra rotaria Theodoridis 1984

†Reticulofenestra scrippsae Bukry and Percival 1973

†Reticulofenestra umbilicus Levin 1965

G. Rhabdolithina Reinhardt 1967

G. Sphenaster Wilcoxon 1970

G. Striatococcolithus Bukry 1971

G. †Syracosphaera Lohmann 1902

†Syracosphaera heimii Lohmann 1919

Syracosphaera mediterranea Gaarder and Heirndal 1977

G. Umbilicosphaera Lohmann 1902

†Umbilicosphaera jafari Muller 1974

Invalid names: Geminilithella Backman 1980 [synonym]

G. Vermiculithina Bukry and Percival 1971

G. Wiseorhabdus Bukry 1981

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

low Mg calcitep

Environment:

oceanic, oligotrophic, marine, brackishp

Locomotion:

passively mobilep

Life habit:

planktonicp

Depth habitat:

surfacep

Diet:

"photoautotroph"p

Created:

2006-10-31 06:24:05

Modified:

2006-10-31 08:24:05

Source: p = phylum

Reference: Kiessling 2003

Age range

Maximum range based only on fossils: base of the Late/Upper Famennian to the top of the Holocene or 365.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 358.86 Ma

Collections (10668 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Late/Upper Famennian	365.2 - 358.86	USA (Missouri)	Leptodiscus corrugatus (6027 6029)
Middle Hettangian - Late/Upper Hettangian	201.4 - 199.5	Canada (Alberta)	Tubirhabdus patulus, Crepidolithus crassus, Mitrolithus jansae, Mitrolithus elegans, Crucirhabdus primulus, Parhabdolithus liasicus (197630)
Hettangian - Sinemurian	201.4 - 192.9	Austria	Schizosphaerella punctulata (38056 38057 38058 38059)
Late/Upper Sinemurian - Late/Upper Pliensbachian	199.5 - 184.2	Morocco	Bussonius prinsii, Lotharingius frodoi, Lotharingius haufii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus liasicus, Schizosphaerella sp., Schizosphaerella spp., Similiscutum finchii, Calyculus sp. (205779) Calcivascularis jansae, Schizosphaerella sp. (205765) Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp. (205777) Crepidolithus crassus, Calcivascularis jansae, Crucirhabdus primulus, Parhabdolithus robustus, Parhabdolithus liasicus, Schizosphaerella spp., Schizosphaerella sp., Similiscutum cruciulus, Calyculus sp. (205771) Crepidolithus crassus, Lotharingius haufii, Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp., Similiscutum novum, Similiscutum finchii (205778) Lotharingius frodoi, Calcivascularis jansae, Mitrolithus elegans, Parhabdolithus robustus, Parhabdolithus liasicus, Schizosphaerella sp., Schizosphaerella spp., Similiscutum cruciulus (205772) Lotharingius frodoi, Lotharingius haufii, Calcivascularis jansae, Mitrolithus elegans, Parhabdolithus liasicus, Similiscutum finchii, Similiscutum cruciulus (205774) Lotharingius haufii, Mitrolithus elegans, Schizosphaerella sp. (205776) Mitrolithus lenticularis, Schizosphaerella sp. (205764) Schizosphaerella sp. (205759 205760 205762 205763 205767 205768 205770 205775) Schizosphaerella sp., Schizosphaerella spp. (205761 205773) Tubirhabdus patulus, Crepidolithus crassus, Mitrolithus lenticularis, Mitrolithus elegans, Crucirhabdus primulus, Schizosphaerella sp., Calyculus sp. (205769)
Pliensbachian	192.9 - 184.2	Spain (Jaén)	Tubirhabdus patulus, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp (201758) Tubirhabdus patulus, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Orthogonoides hamiltoniae, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus (201759) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp (201753) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Mitrolithus elegans, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus (201755) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp (201752) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum depravatum, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus recondita (201756) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus (201754)
Spinatum	192.9 - 184.2	Morocco	Lotharingius sigillatus (203735 203736)
Early/Lower Toarcian	184.2 - 182.9	Spain (Granada)	Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus striatus, Similiscutum sp. 1, Carinolithus superbus (199494)
Toarcian	184.2 - 174.7	Spain (Jaén)	Tubirhabdus patulus, Crepidolithus cavus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Orthogonoides hamiltoniae, Schizosphaerella punctulata, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp (201668) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum depravatum, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita (201673) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Calyculus sp (201675) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Orthogonoides hamiltoniae, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp (201670) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp (201671) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Orthogonoides hamiltoniae, Schizosphaerella punctulata, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp (201685) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp (201666) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus sp (201674) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp (201669) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Mitrolithus elegans, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp (201667) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Orthogonoides hamiltoniae, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus (201751) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp (201750) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp (201678) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp (201665) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp (201672) Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Schizosphaerella punctulata, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus sp (201676) Tubirhabdus patulus, Crepidolithus granulatus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Orthogonoides hamiltoniae, Schizosphaerella punctulata, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp (201677)
Toarcian	184.2 - 174.7	Saudi Arabia	Ellipsagelosphaera britannica, Tetralithus gothicus (45089) Ellipsagelosphaera britannica, Tetralithus goticus (45191)
Toarcian	184.2 - 174.7	Spain (Murcia)	Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201775) Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201779) Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Triscutum sullivanii, Carinolithus superbus (201770) Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (201765 201766) Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Schizosphaerella punctulata, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201776) Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (201772) Crepidolithus crassus, Lotharingius sigillatus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus superbus (201792) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus (201778) Lotharingius hauffii, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus superbus (201774) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201794) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Triscutum sullivanii, Carinolithus poulnabronei, Carinolithus superbus (201789) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (201777) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201793) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Triscutum sullivanii, Carinolithus poulnabronei, Carinolithus superbus (201773) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius umbriensis, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201783) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius barozii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (201764) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201791) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (201769) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201781) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201782) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (201780) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius sigillatus, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus striatus, Carinolithus superbus (201787) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus superbus (201788) Zeugrhabdotus erectus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus superbus (201771) Zeugrhabdotus erectus, Crepidolithus crassus, Lotharingius umbriensis, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (201768) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201790)
Middle Toarcian	182.9 - 180.4	Spain (Murcia)	Crepidolithus crassus, Bussonius prinsii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae (201809) Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae (201804) Crepidolithus crassus, Bussonius prinsii, Lotharingius umbriensis, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. A, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201796) Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201799) Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Schizosphaerella punctulata, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae (201813) Crepidolithus crassus, Lotharingius hauffii, Lotharingius velatus, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201807) Crepidolithus crassus, Lotharingius hauffii, Lotharingius velatus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus superbus (201806) Crepidolithus crassus, Lotharingius hauffii, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201805) Crepidolithus crassus, Lotharingius umbriensis, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201802) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (201797) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201798) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201812) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201800) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201803) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (201795) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Mitrolithus lenticularis, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae (201808) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201810) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Schizosphaerella punctulata, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201811) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius umbriensis, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (201801)
Levesquei	182.9 - 174.7	Germany	Bussonius leufuensis (59428) Crepidolithus crassus, Bussonius leufuensis, Lotharingius crucicentralis, Lotharingius velatus, Watznaueria contracta, Crucirhabdus primulus, Schizosphaerella sp., Retecapsa incompta, Discorhabdus criotus, Triscutum sullivanii, Triscutum tiziense, Carinolithus crassus, Carinolithus superbus (59430) Crepidolithus crassus, Lotharingius crucicentralis, Crucirhabdus primulus, Retecapsa incompta, Triscutum sullivanii, Carinolithus superbus (59434) Lotharingius velatus, Watznaueria contracta, Schizosphaerella sp., Retecapsa incompta, Discorhabdus criotus (59431)
Late/Upper Toarcian	180.4 - 174.7	Germany	Crepidolithus crassus, Schizosphaerella sp., Retecapsa incompta, Discorhabdus striatus, Carinolithus superbus (59454) Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Tubirhabdus patulus, Tubirhabdus patulus, Crepidolithus crassus, Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Lotharingius haufforum, Lotharingius sp. 4, Lotharingius barozii, Lotharingius contractus, Lotharingius sigillatus, Crucirhabdus magnus, Parhabdolithus cavus, Stradnerlithus humilis, Schizosphaerella punctulata, Biscutum criotum, Biscutum sp. 2, Biscutum sp. 4, Biscutum striatum, Biscutum intermedium, Axopodorhabdus depravatus, Palaeopontosphaera dubia, Discorhabdus novus, Triscutum beaminsterensene, Triscutum sullivanii, Ethmorhabdus crucifer, Axopodorhabdus atavus, Axopodorhabdus cylindratus, Carinolithus sceptrum, Carinolithus superbus, Proculithus charlottei, Calyculus cribrum, Catillus hommerilii (59453) Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Tubirhabdus patulus, Tubirhabdus patulus, Crepidolithus crassus, Vagalapilla quadriarculla, Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius barozii, Lotharingius contractus, Lotharingius contractus, Lotharingius sigillatus, Parhabdolithus cavus, Stradnerlithus humilis, Schizosphaerella punctulata, Biscutum striatum, Biscutum intermedium, Palaeopontosphaera dubia, Discorhabdus novus, Axopodorhabdus atavus, Axopodorhabdus cylindratus, Carinolithus superbus, Calyculus sp. 1, Calyculus sp. 2, Proculithus fistulatus, Calyculus cribrum, Catillus hommerilii (59451) Zeugrhabdotus choffatii, Zeugrhabdotus sp. 1, Zeugrhabdotus sp. 2, Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Vagalapilla quadriarculla, Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Lotharingius sp. 2, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius barozii, Lotharingius contractus, Lotharingius contractus, Lotharingius sigillatus, Crucirhabdus magnus, Parhabdolithus cavus, Schizosphaerella punctulata, Biscutum criotum, Biscutum striatum, Biscutum intermedium, Axopodorhabdus depravatus, Palaeopontosphaera sp. 1, Palaeopontosphaera dubia, Discorhabdus novus, Triscutum pricatillus, Axopodorhabdus atavus, Axopodorhabdus cylindratus, Carinolithus sp. 1, Carinolithus sceptrum, Carinolithus superbus, Calyculus sp. 2, Proculithus fistulatus, Proculithus charlottei (59449)
Late/Upper Toarcian	180.4 - 174.7	Spain (Granada)	Crepidolithus crassus, Bussonius prinsii, Lotharingius sigillatus, Watznaueria sp. A, Schizosphaerella punctulata, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (202099) Crepidolithus crassus, Lotharingius barozii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus superbus (201880) Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (202100) Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199181) Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199231) Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius velatus, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199235) Crepidolithus crassus, Lotharingius crucicentralis, Watznaueria sp. A, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (202109) Crepidolithus crassus, Lotharingius sigillatus, Lotharingius velatus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (202098) Crepidolithus crassus, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei (202105) Crepidolithus crassus, Lotharingius sigillatus, Watznaueria sp. A, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei (202112) Crepidolithus crassus, Lotharingius velatus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (202111) Crepidolithus crassus, Lotharingius velatus, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (202116) Crepidolithus crassus, Schizosphaerella punctulata (202113) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei (202117) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus (202106) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei (202110) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (202103) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii (202108) Crepidolithus crassus, Watznaueria sp. A, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum (202104) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (199169) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (202107) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199187) Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199172) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius barozii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199493) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (199168) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp (202097) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199496) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei (199248) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199487) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. B, Schizosphaerella punctulata, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (199173) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (199491) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. B, Schizosphaerella punctulata, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199492) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus superbus (199164) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (201948) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199195) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (199198) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus (202115) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Triscutum sullivanii, Carinolithus poulnabronei, Carinolithus superbus (199495) Tubirhabdus patulus, Crepidolithus crassus, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (202101) Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei (202114) Watznaueria sp. B, Schizosphaerella punctulata, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei (202102) Zeugrhabdotus erectus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus (199167) Zeugrhabdotus erectus, Crepidolithus crassus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (199253) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus (199166) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Bussonius prinsii, Lotharingius umbriensis, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199247) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius barozii, Lotharingius crucicentralis, Lotharingius sigillatus, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199177) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius barozii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp (199488) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Triscutum sullivanii, Carinolithus superbus (199165) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus sp. cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus (199490) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Lotharingius velatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Schizosphaerella punctulata, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Triscutum sullivanii, Carinolithus superbus (199171) Zeugrhabdotus erectus, Tubirhabdus patulus, Crepidolithus crassus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Schizosphaerella punctulata, Biscutum intermedium, Biscutum dubium, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus (199229)
Early/Lower Aalenian	174.7 - 170.9	Germany	Bussonius prinsii, Calyculus sp. (59458) Crepidolithus crassus, Bussonius prinsii, Schizosphaerella sp., Retecapsa incompta, Discorhabdus striatus, Carinolithus superbus, Calyculus sp. (59457)
Bajocian	170.9 - 168.2	Spain	Watznaueria communis, Watznaueria contracta, Watznaueria britannica, Cyclagelosphaera margerelii (214379)
Bajocian	170.9 - 168.2	Saudi Arabia	Cyclagelosphaera margereli (45268) Ellipsagelosphaera britannica, Schizosphaerella punctulata, Tetralithus gothicus (45265)
Bathonian	168.2 - 165.3	Saudi Arabia	Diazomatolithus lehmani, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Tetralithus gothicus (45519) Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Schizosphaerella punctulata, Hexalithus magharensis, Tetralithus gothicus (45464)
Callovian	165.3 - 161.5	Saudi Arabia	Cyclagelosphaera margereli, Tetralithus gothicus (45558) Diazomatolithus lehmani, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Tetralithus gothicus (45547) Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Tetralithus gothicus (45549) Ellipsagelosphaera communis, Cyclagelosphaera margereli (45555) Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli (45540) Zeugrhabdotus erectus, Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Stephanolithion bigoti, Discorhabdus sp., Ethmorhabdus gallicus, Zygodiscus sp., Tetralithus gothicus (45541) Zeugrhabdotus erectus, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Stephanolithion bigoti, Polypodorhabdus escaigii, Ethmorhabdus anglicus (47890)
Middle Oxfordian	161.5 - 154.8	Switzerland	Crepidolithus perforatus, Vekshinella quadriarculla, Ellipsagelosphaera britannica, Cyclagelosphaera margereti, Stephanolithion bigoti, Discorhabdus patulus, Ethmorhabdus gallicus, Axopodorhabdus cylindratus, Hexapodorhabdus cuvillieri (58568)