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Haptophyta (haptophyte alga)

Synonymy list
YearName and author
1986Haptophyta Hibberd ex Cavalier-Smith
1997Haptophyta Young and Bown
2005Haptophyta Adl et al. p. 401 figs. Table 1
2015Haptophyta Ruggiero et al. pp. Table 2

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RankNameAuthor
kingdomChromistaCavalier-Smith 1981
—Hacrobia
RankNameAuthor
phylumHaptophytaHibberd ex Cavalier-Smith 1986
phylumHaptophytaHibberd ex Cavalier-Smith 1986

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Haptophyta Hibberd ex Cavalier-Smith 1986 [haptophyte alga]
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G. Aspidorhabdus Hay and Towe 1962
G. †Camuralithus de Kaenel and Villa 1996
Cl. Coccolithophyceae Rothmaler 1951
+
G. †Athenagalea Hattner and Wise 1980
G. †Biantholithus Bramlette and Martini 1964
Or. Braarudosphaerales Aubry 2013
+
Fm. †Polycyclolithaceae Forchheimer 1972
+
G. †Assipetra Roth 1973
+
†Assipetra infracretacea Thierstein 1973
†Assipetra terebrodentarius Applegate and Bralower 1987
G. †Eprolithus Stover 1966
+
†Eprolithus antiquus Perch-Nielsen 1979
†Eprolithus apertior Black 1973
†Eprolithus floralis Stradner 1962
†Eprolithus moratus Stover 1966
†Eprolithus octopetalus Varol 1992
†Eprolithus rarus Varol 1992
Invalid names: Polycyclolilthus Forchheimer 1968 [synonym]
G. †Farhania Varol 1992
+
†Farhania varolii Jakubowski 1986
G. †Hayesites Manivit 1971
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†Hayesites albiensis Manivit 1971
†Hayesites irregularis Thierstein et al. 1972
G. †Hexalithus Gardet 1955
+
†Hexalithus gardetiae Bukry 1969
†Hexalithus noeliae Loeblich and Tappan 1966
G. †Lithastrinus Stradner 1962
+
†Lithastrinus grillii Stradner 1962
†Lithastrinus septenarius Forchheimer 1972
G. †Micula Vekshina 1959 [coccolithophore]
+
†Micula adumbrata Burnett 1998
†Micula clypeata Lees and Bown 2005
†Micula concava Stradner 1960
†Micula cubiformis Forchheimer 1972
†Micula decussata Vekshina 1959 [coccolithophore]
†Micula murus Martini 1961
†Micula praemurus Bukry 1973
†Micula premolisilvae Lees and Bown 2005
†Micula prinsii Perch-Nielsen 1979
†Micula staurophora Gardet 1955
†Micula swastica Stradner and Steinmetz 1984
Invalid names: Nannotetraster Martini and Stradner 1960 [synonym]
G. †Polycostella Thierstein 1971
+
†Polycostella beckmanii Thierstein 1971
†Polycostella senaria Thierstein 1971
G. †Quadrum Prins and Perch-Nielsen 1977
+
†Quadrum bengalensis Burnett 1998
†Quadrum eneabrachium Varol 1992
†Quadrum eptabrachium Varol 1992
†Quadrum gartneri Prins and Perch-Nielsen 1977
†Quadrum giganteum Varol 1992
†Quadrum intermedium Varol 1992
†Quadrum octobrachium Varol 1992
†Quadrum svabenickae Burnett 1998
G. †Radiolithus Stover 1966
+
†Radiolithus orbiculatus Forchheimer 1972
†Radiolithus planus Stover 1966
G. †Rucinolithus Stover 1966
+
†Rucinolithus magnus Bukry 1975
†Rucinolithus windleyae Rutledge and Bown 1996
†Rucinolithus wisei Thierstein 1971
G. †Tegulalithus Crux 1986
+
†Tegulalithus tessellatus Stradner 1968
G. †Uniplanarius Hattner and Wise 1980
+
†Uniplanarius clarkei Lees and Bown 2005
†Uniplanarius gothicus Deflandre 1959
†Uniplanarius sissinghii Perch-Nielsen 1986
†Uniplanarius trifidus Stradner 1961
Fm. Ceratolithaceae Norris 1965
+
G. Amaurolithus Gartner and Bukry 1975
+
Amaurolithus primus Bukry and Percival 1971
G. Angulolithina Bukry 1973
G. Ceratolithus Kamptner 1950
G. Florisphaera Okada and Honjo 1973
G. †Nicklithus Raffi et al. 1998
+
Nicklithus amplificus Bukry and Percival 1971
G. †Orthorhabdus Bramlette and Wilcoxon 1967
Or. †Coccolithales Schwarz 1932
+
Fm. Coccolithaceae Poche 1913
+
G. †Bicolumnus Wei and Wise 1990
G. †Birkelundia Perch-Nielsen 1971
G. †Bramletteius Gartner 1969
G. †Campylosphaera Kamptner 1963
G. †Chiasmolithus Hay et al. 1966
+
†Chiasmolithus bidens Bramlette and Sullivan 1961
†Chiasmolithus consuetus Bramlette and Sullivan 1961
†Chiasmolithus danicus Hay and Mohler 1967
†Chiasmolithus eograndis Perch-Nielsen 1971
†Chiasmolithus expansus Bramlette and Sullivan 1961
†Chiasmolithus grandis Bramlette and Riedel 1954
†Chiasmolithus modestus Perch-Nielsen 1971
†Chiasmolithus oamaruensis Deflandre 1954
†Chiasmolithus solitus Bramlette and Sullivan 1961
G. †Clausicoccus Prins 1979
G. Coronocyclus Hay et al. 1966
G. †Ericsonia Black 1964
G. †Gephyrorhabdus Hill 1976
G. †Holodiscolithus Roth 1970
+
†Holodiscolithus solidus Deflandre and Fert 1954
G. †Hughesius Varol 1989
G. †Praeprinsius Varol and Jakubowski 1989
G. †Solidopons Theodoridis 1984
Invalid names: Coccolithacea Ellis and Lohman 1973 [empty], Coccolithophorales [objective synonym], Hymenomonadaceae Senn 1900 [empty], Pleurochrysidaceae Fresnel and Billard 1991 [empty]
Or. †Coccosphaerales Haeckel 1894
+
Fm. Calcidiscaceae Young and Bown 1997
+
G. Calcidiscus Kamptner 1950
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Calcidiscus leptoporus Murray and Blackman 1898
Calcidiscus macintyrei Bukry and Bramlette 1969
G. †Cryptococcolithus Gartner 1992
G. Hayaster Bukry 1973
G. †Hayella Gartner 1969
G. Oolithotus Cohen and Reinhardt 1968
G. †Daktylethra Gartner and Bukry 1969
Or. †Discoasterales Hay 1977
+
Fm. †Discoasteraceae Tan Sin Hok 1927
+
G. †Catinaster Martini and Bramlette 1963
Fm. †Fasciculithaceae Hay and Mohler 1967
+
G. †Fasciculithus Bramlette and Sullivan 1961
G. †Lithoptychius Aubry 2011
Fm. †Heliolithaceae Hay and Mohler 1967
+
G. †Bomolithus Roth 1973
G. †Heliolithus Bramlette and Sullivan 1961
Fm. †Ellipsagelosphaeraceae Noël 1965
+
Subfm. †Actinosphaeroideae Noël 1965
+
G. †Calolithus Noël 1965
G. †Ellipsolithus Sullivan 1964
G. Gladiolithus Jordan and Chamberlain 1992
G. †Ilselithina Stradner and Adamiker 1966
Or. Isochrysidales Pascher 1910
+
Fm. Gephyrocapsaceae Heimdal 1997
+
G. Crenalithus Roth 1973
Fm. Noelaerhabdaceae Jerkovic 1970
+
G. Apertapetra Hay et al. 1966
G. Cepekiella Roth 1970
G. †Cribrocentrum Perch-Nielsen 1971
G. Dictyococcites Black 1967
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†Dictyococcites scrippsae Bukry and Percival 1973
G. Emiliania Hay and Mohler 1967
G. Gephyrocapsa Kamptner 1943
G. †Pseudoemiliania Gartner 1969
Fm. †Prinsiacea Hay and Mohler 1967
+
G. †Toweius Hay and Mohler 1973
+
†Toweius africanus Perch-Nielsen 1981
†Toweius callosus Perch-Nielsen 1971
†Toweius eminens Bramlette and Sullivan 1961
†Toweius occultatus Locker 1967
†Toweius petalosus Ellis and Lohman 1973
†Toweius rotundus Perch-Nielsen 1978
†Toweius serotinus Bybell and Self-Trail 1995
Fm. †Prinsiaceae Hay and Mohler 1967
+
G. †Futyania Varol 1989
G. †Girgisia Varol 1989
G. Hornibrookina Edwards 1973
+
†Hornibrookina teuriensis Edwards 1973
†Hornibrookina weimerae Self-Trail 2011
G. †Neobiscutum Varol 1989
G. †Prinsius Hay and Mohler 1967
+
†Prinsius dimorphosus Perch-Nielsen 1977
†Prinsius martinii Haq 1971
†Prinsius tenuiculus Perch-Nielsen 1984
Invalid names: Isochrysidaceae Pascher 1910 [empty]
G. †Lanternithus Stradner 1962
+
†Lanternithus minutus Stradner 1962
G. †Minylitha Bukry 1973
G. Neosphaera Lecal-Schlauder 1949
G. †Pedinocyclus Bukry and Bramlette 1971
Fm. †Rhomboasteraceae Bown 2005
+
G. †Rhomboaster Bramlette and Sullivan 1961
G. †Tribrachiatus Shamrai 1963
G. Tetralithoides Theodoridis 1984
G. †Transversopontis Hay et al. 1966
Or. Zygodiscales Young and Bown 1997
+
Fm. Pontosphaeraceae Lemmermann 1908
+
G. Calciopilleus Müller 1974
G. Scyphosphaera Lohmann 1902
Fm. Zygodiscaceae Hay and Mohler 1967
+
G. †Isthmolithus Deflandre and Fert 1954
+
†Isthmolithus recurvus Deflandre 1954
G. †Lophodolithus Deflandre and Fert 1954
+
†Lophodolithus mochlophorus Deflandre 1954
G. †Nannotetrina Achuthan and Stradner 1969
G. †Neococcolithes Sujkowski 1931
+
†Neococcolithes dubius Deflandre and Fert 1954
G. †Zygodiscus Bramlette and Sullivan 1961
Invalid names: Helicosphaeraceae Black 1971 [empty]
Invalid names: Phaeocystales Medlin 2000 [empty], Prymnesiales Papenfuss 1955 [empty], Prymnesiophyceae Hibberd 1976 [synonym]
G. Conococcolithus Hay and Mohler 1967
G. Corannulus Stradner 1962
G. Craspedolithus Kamptner 1963
G. Crassapontosphaera Boudreaux and Hay 1969
G. †Crassidiscus Okada 1990
G. †Cricosphaera Braarud 1960
G. Cyclolithella Loeblich and Tappan 1963
G. †Cycloperfolithus Lehotayova and Priewalder 1978
+
G. †Agalmatoaster Klumpp 1953
+
G. †Trochoaster Klumpp 1953
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†Trochoaster simplex Klumpp 1953
G. †Duplexipodorhabdus Varol and Girgis 1994
G. Ellipsodiscoaster Boudreaux and Hay 1969
G. †Helicosphaera Kamptner 1954
+
†Helicosphaera burkei Black 1971
†Helicosphaera carteri Wallich 1877
†Helicosphaera cateri Wallich 1877
†Helicosphaera compacta Bramlette and Wilcoxon 1967
†Helicosphaera intermedia Martini 1965
†Helicosphaera lophota Bramlette and Sullivan 1961
†Helicosphaera neolophota Barron et al. 1984
†Helicosphaera obliqua Bramlette and Wilcoxon 1967
†Helicosphaera pacifica Müller and Brönnimann 1974
†Helicosphaera papillata Bukry and Bramlette 1969
†Helicosphaera parallela Bramlette and Wilcoxon 1967
†Helicosphaera reticulata Bramlette and Wilcoxon 1967
†Helicosphaera rhomba Bukry 1971
†Helicosphaera sellii Bukry and Bramlette 1969
†Helicosphaera seminulum Bramlette and Sullivan 1961
+
†Helicosphaera seminulum seminulum Bramlette and Sullivan 1961
†Helicosphaera seminulum lophota Bramlette and Sullivan 1961
†Helicosphaera stalis Theodoridis 1984
†Helicosphaera truncata Bramlette and Wilcoxon 1967
Invalid names: Helicopontosphaera Hay and Mohler 1967 [synonym]
G. Heliorthus Brönnimann and Stradner 1960
+
†Heliorthus tenuis Stradner 1961
G. Koczyia Boudreaux and Hay 1969
G. Leptodiscus Bukry and Bramlette 1969
Fm. Lithostromationaceae Deflandre 1959
+
G. Imperiaster Martini 1970
G. Lithostromation Deflandre 1942
G. Nannocorbis Müller 1974
G. Orthozygus Bramlette and Wilcoxon 1967
G. †Pervilithus Crux 1981
G. †Phanulithus Wise and Wind 1977
G. Pyrocyclus Hay and Towe 1962
G. Reticulofenestra Hay et al. 1966
+
†Reticulofenestra dictyoda Stradner 1968
†Reticulofenestra reticulata Roth and Thierstein 1972
†Reticulofenestra rotaria Theodoridis 1984
†Reticulofenestra scrippsae Bukry and Percival 1973
G. Rhabdolithina Reinhardt 1967
G. Sphenaster Wilcoxon 1970
G. Striatococcolithus Bukry 1971
G. †Syracosphaera Lohmann 1902
+
†Syracosphaera heimii Lohmann 1919
Syracosphaera mediterranea Gaarder and Heirndal 1977
G. Umbilicosphaera Lohmann 1902
+
†Umbilicosphaera jafari Muller 1974
Invalid names: Geminilithella Backman 1980 [synonym]
G. Vermiculithina Bukry and Percival 1971
G. Wiseorhabdus Bukry 1981
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitep
Environment: oceanic, oligotrophic, marine, brackishp
Locomotion: passively mobilep
Life habit: planktonicp
Depth habitat: surfacep
Diet: "photoautotroph"p
Created: 2006-10-31 06:24:05
Modified: 2006-10-31 08:24:05
Source: p = phylum
Reference: Kiessling 2003

Age range

Maximum range based only on fossils: base of the Late/Upper Famennian to the top of the Holocene or 365.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 358.86 Ma

Collections (10697 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Late/Upper Famennian365.2 - 358.86USA (Missouri) Leptodiscus corrugatus (6027 6029)
Gzhelian - Asselian303.7 - 293.52Russian Federation Calcidiscus sp. (75611)
Middle Hettangian - Late/Upper Hettangian201.4 - 199.5Canada (Alberta) Crucirhabdus primulus, Mitrolithus elegans, Mitrolithus jansae, Parhabdolithus liasicus, Crepidolithus crassus, Tubirhabdus patulus (197630)
Hettangian - Sinemurian201.4 - 192.9Austria Schizosphaerella punctulata (38056 38057 38058 38059)
Late/Upper Sinemurian - Late/Upper Pliensbachian199.5 - 184.2Morocco Bussonius prinsii, Lotharingius frodoi, Lotharingius haufii, Lotharingius sigillatus, Similiscutum finchii, Calyculus sp., Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus liasicus, Schizosphaerella sp., Schizosphaerella spp. (205779) Calcivascularis jansae, Schizosphaerella sp. (205765) Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp. (205777) Calyculus sp., Crucirhabdus primulus, Mitrolithus lenticularis, Mitrolithus elegans, Schizosphaerella sp., Crepidolithus crassus, Tubirhabdus patulus (205769) Lotharingius frodoi, Lotharingius haufii, Similiscutum finchii, Similiscutum cruciulus, Mitrolithus elegans, Calcivascularis jansae, Parhabdolithus liasicus (205774) Lotharingius frodoi, Similiscutum cruciulus, Mitrolithus elegans, Calcivascularis jansae, Parhabdolithus robustus, Parhabdolithus liasicus, Schizosphaerella sp., Schizosphaerella spp. (205772) Lotharingius haufii, Mitrolithus elegans, Schizosphaerella sp. (205776) Lotharingius haufii, Similiscutum novum, Similiscutum finchii, Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp., Crepidolithus crassus (205778) Mitrolithus lenticularis, Schizosphaerella sp. (205764) Schizosphaerella sp. (205759 205760 205762 205763 205767 205768 205770 205775) Schizosphaerella sp., Schizosphaerella spp. (205761 205773) Similiscutum cruciulus, Calyculus sp., Crucirhabdus primulus, Calcivascularis jansae, Parhabdolithus robustus, Parhabdolithus liasicus, Schizosphaerella spp., Schizosphaerella sp., Crepidolithus crassus (205771)
Pliensbachian192.9 - 184.2Spain (Jaén) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Biscutum depravatum, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus recondita, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201756) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201752) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Crucirhabdus primulus, Mitrolithus lenticularis, Mitrolithus elegans, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201755) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201753) Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Schizosphaerella punctulata, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201758) Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201759) Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Mitrolithus lenticularis, Calcivascularis jansae, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201754)
Spinatum192.9 - 184.2Morocco Lotharingius sigillatus (203735 203736)
Early/Lower Toarcian184.2 - 182.9Spain (Granada) Watznaueria colacicchii, Lotharingius velatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Similiscutum sp. 1, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199494)
Toarcian184.2 - 174.7Saudi Arabia Ellipsagelosphaera britannica, Tetralithus gothicus (45089) Ellipsagelosphaera britannica, Tetralithus goticus (45191)
Toarcian184.2 - 174.7Spain (Murcia) Discorhabdus striatus, Schizosphaerella punctulata, Crepidolithus crassus (201778) Lotharingius hauffii, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella punctulata (201774) Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201781) Lotharingius sigillatus, Biscutum intermedium, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201787) Lotharingius sigillatus, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201792) Lotharingius sigillatus, Lotharingius hauffii, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201772) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201776) Watznaueria colacicchii, Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201794) Watznaueria colacicchii, Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201779) Watznaueria colacicchii, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201793) Watznaueria colacicchii, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201775) Watznaueria colacicchii, Lotharingius sigillatus, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201788) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201764) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201780) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (201771) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201765 201766) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (201768) Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201777) Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Triscutum sullivanii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201773) Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201783) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201782) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Triscutum sullivanii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201789) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Biscutum intermedium, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201791) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (201790) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius velatus, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Triscutum sullivanii, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201770) Watznaueria colacicchii, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201769)
Toarcian184.2 - 174.7Spain (Jaén) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Axopodorhabdus atavus, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Diductius constans, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201750) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Diductius constans, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201678) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Tubirhabdus patulus (201675) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201751) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Axopodorhabdus atavus, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus cavus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201668) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Tubirhabdus patulus (201673) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Axopodorhabdus atavus, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201670) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Axopodorhabdus atavus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201677) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus sp, Axopodorhabdus atavus, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Diductius constans, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201674) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201666) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201671) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Diductius constans, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201669) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp, Mitrolithus lenticularis, Mitrolithus elegans, Calcivascularis jansae, Diductius constans, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201667) Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Axopodorhabdus atavus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201685) Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Axopodorhabdus atavus, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Diductius constans, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201676) Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201665) Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Axopodorhabdus atavus, Crucirhabdus primulus, Mitrolithus lenticularis, Calcivascularis jansae, Parhabdolithus distinctus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Tubirhabdus patulus (201672)
Middle Toarcian182.9 - 180.4Spain (Murcia) Bussonius prinsii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Crepidolithus crassus (201809) Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201810) Lotharingius hauffii, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Mitrolithus lenticularis, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201808) Lotharingius hauffii, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201811) Lotharingius hauffii, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201805) Lotharingius sigillatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Mitrolithus lenticularis, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201803) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201799) Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201812) Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Schizosphaerella punctulata, Crepidolithus crassus (201813) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Mitrolithus lenticularis, Schizosphaerella punctulata, Crepidolithus crassus (201802) Lotharingius velatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201807) Lotharingius velatus, Lotharingius hauffii, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201806) Watznaueria colacicchii, Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Schizosphaerella punctulata, Crepidolithus crassus (201804) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201795) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Mitrolithus lenticularis, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201800) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Mitrolithus lenticularis, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201798) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201801) Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201797) Watznaueria sp. A, Bussonius prinsii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Mitrolithus lenticularis, Schizosphaerella punctulata, Crepidolithus crassus (201796)
Levesquei182.9 - 174.7Germany Bussonius leufuensis (59428) Lotharingius crucicentralis, Retecapsa incompta, Triscutum sullivanii, Carinolithus superbus, Crucirhabdus primulus, Crepidolithus crassus (59434) Watznaueria contracta, Bussonius leufuensis, Lotharingius velatus, Lotharingius crucicentralis, Retecapsa incompta, Discorhabdus criotus, Triscutum sullivanii, Triscutum tiziense, Carinolithus crassus, Carinolithus superbus, Crucirhabdus primulus, Schizosphaerella sp., Crepidolithus crassus (59430) Watznaueria contracta, Lotharingius velatus, Retecapsa incompta, Discorhabdus criotus, Schizosphaerella sp. (59431)
Late/Upper Toarcian180.4 - 174.7Spain (Granada) Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius crucicentralis, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202107) Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202110) Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202114) Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202103) Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202117) Discorhabdus striatus, Schizosphaerella punctulata, Crepidolithus crassus (202106) Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Schizosphaerella punctulata, Crepidolithus crassus (202108) Lotharingius sigillatus, Lotharingius barozii, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199177) Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202097) Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199496) Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202100) Lotharingius sigillatus, Lotharingius velatus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202098) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Biscutum dubium, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202101) Lotharingius velatus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202111) Lotharingius velatus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202116) Lotharingius velatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (199235) Schizosphaerella punctulata, Crepidolithus crassus (202113) Watznaueria colacicchii, Lotharingius sigillatus, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199198) Watznaueria colacicchii, Lotharingius sigillatus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202115) Watznaueria colacicchii, Lotharingius sigillatus, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202105) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199487) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199248) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (199253) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Triscutum sullivanii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199495) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Triscutum sullivanii, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199165) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199169) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius barozii, Lotharingius crucicentralis, Biscutum depravatum, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201880) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201948) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199164) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199493) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199229) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199195) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius velatus, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199168) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199187) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Triscutum sullivanii, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199171) Watznaueria colacicchii, Watznaueria sp. B, Lotharingius crucicentralis, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199492) Watznaueria colacicchii, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199173) Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199166) Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum depravatum, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (199167) Watznaueria sp. A, Bussonius prinsii, Lotharingius sigillatus, Discorhabdus striatus, Discorhabdus criotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202099) Watznaueria sp. A, Discorhabdus striatus, Similiscutum novum, Schizosphaerella punctulata, Crepidolithus crassus (202104) Watznaueria sp. A, Lotharingius crucicentralis, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202109) Watznaueria sp. A, Lotharingius sigillatus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202112) Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199247) Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum depravatum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (199181) Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Biscutum dubium, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199490) Watznaueria sp. B, Bussonius prinsii, Lotharingius sigillatus, Lotharingius velatus, Lotharingius hauffii, Lotharingius crucicentralis, Biscutum depravatum, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199172) Watznaueria sp. B, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Schizosphaerella punctulata (202102) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius barozii, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus, Tubirhabdus patulus (199488) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (199231) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199491)
Late/Upper Toarcian180.4 - 174.7Germany Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius contractus, Lotharingius barozii, Biscutum striatum, Biscutum intermedium, Palaeopontosphaera dubia, Discorhabdus novus, Carinolithus superbus, Calyculus sp. 1, Calyculus sp. 2, Calyculus cribrum, Proculithus fistulatus, Catillus hommerilii, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Parhabdolithus cavus, Stradnerlithus humilis, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Tubirhabdus patulus, Tubirhabdus patulus, Vagalapilla quadriarculla (59451) Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Lotharingius haufforum, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius barozii, Biscutum criotum, Biscutum sp. 2, Biscutum sp. 4, Biscutum striatum, Axopodorhabdus depravatus, Biscutum intermedium, Palaeopontosphaera dubia, Discorhabdus novus, Triscutum beaminsterensene, Triscutum sullivanii, Carinolithus sceptrum, Carinolithus superbus, Calyculus cribrum, Proculithus charlottei, Catillus hommerilii, Ethmorhabdus crucifer, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Crucirhabdus magnus, Parhabdolithus cavus, Stradnerlithus humilis, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Tubirhabdus patulus, Tubirhabdus patulus (59453) Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Lotharingius sp. 2, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius contractus, Lotharingius barozii, Biscutum criotum, Biscutum striatum, Axopodorhabdus depravatus, Biscutum intermedium, Palaeopontosphaera sp. 1, Palaeopontosphaera dubia, Discorhabdus novus, Triscutum pricatillus, Carinolithus sp. 1, Carinolithus sceptrum, Carinolithus superbus, Calyculus sp. 2, Proculithus fistulatus, Proculithus charlottei, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Crucirhabdus magnus, Parhabdolithus cavus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus choffatii, Zeugrhabdotus sp. 1, Zeugrhabdotus sp. 2, Zeugrhabdotus erectus, Tubirhabdus patulus, Vagalapilla quadriarculla (59449) Retecapsa incompta, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella sp., Crepidolithus crassus (59454)
Early/Lower Aalenian174.7 - 170.9Germany Bussonius prinsii, Calyculus sp. (59458) Bussonius prinsii, Retecapsa incompta, Discorhabdus striatus, Carinolithus superbus, Calyculus sp., Schizosphaerella sp., Crepidolithus crassus (59457)
Bajocian170.9 - 168.2Spain Watznaueria communis, Watznaueria contracta, Watznaueria britannica, Cyclagelosphaera margerelii (214379)
Bajocian170.9 - 168.2Saudi Arabia Cyclagelosphaera margereli (45268) Ellipsagelosphaera britannica, Schizosphaerella punctulata, Tetralithus gothicus (45265)
Bathonian168.2 - 165.3Saudi Arabia Diazomatolithus lehmani, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Tetralithus gothicus (45519) Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Hexalithus magharensis, Schizosphaerella punctulata, Tetralithus gothicus (45464)
Callovian165.3 - 161.5Saudi Arabia Cyclagelosphaera margereli, Tetralithus gothicus (45558) Diazomatolithus lehmani, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Tetralithus gothicus (45547) Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Tetralithus gothicus (45549) Ellipsagelosphaera communis, Cyclagelosphaera margereli (45555) Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli (45540) Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Polypodorhabdus escaigii, Ethmorhabdus anglicus, Stephanolithion bigoti, Zeugrhabdotus erectus (47890) Zygodiscus sp., Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Discorhabdus sp., Ethmorhabdus gallicus, Stephanolithion bigoti, Zeugrhabdotus erectus, Tetralithus gothicus (45541)