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Eucycloscala
Taxonomy
Eucycloscala was named by Cossmann (1895) [Sepkoski's age data: Tr Anis T Dani Sepkoski's reference number: 1130]. It is not extant. It is the type genus of Eucycloscalidae.
It was assigned to Liotiinae by Szabo (1981), Batten (1985); to Archaeogastropoda by Sepkoski (2002); to Eucyclini by Kaim (2004); to Eucycloscalidae by Bandel (2010), Nützel and Kaim (2014); and to Eucyclidae by Ketwetsuriya et al. (2020).
It was assigned to Liotiinae by Szabo (1981), Batten (1985); to Archaeogastropoda by Sepkoski (2002); to Eucyclini by Kaim (2004); to Eucycloscalidae by Bandel (2010), Nützel and Kaim (2014); and to Eucyclidae by Ketwetsuriya et al. (2020).
Species
E. acanticum, E. asiatica, E. baltzeri, E. basistriata, E. binodosa, E. brunhuberi, E. circumnodosa, E. damesi, E. exigua, E. hanwangensis, E. izabellae, E. leo, E. margaritata, E. medionodosa, E. nitida, E. obliquicostata, E. pascoensis, E. praetor (syn. Turbo ornatus, Turbo phillipsii, Littorina praetor, Littorina ornata, Ooliticia phillipsii, Littorina phillipsii), E. semicancellata, E. spinosa, E. spinulosa, E. subbisertus, E. supranodosa, E. ultima
Species lacking formal opinion data
Synonymy list
| Year | Name and author |
|---|---|
| 1895 | Eucycloscala Cossmann |
| 1981 | Eucycloscala Szabo p. 60 |
| 1985 | Eucycloscala Batten p. 35 |
| 2002 | Eucycloscala Sepkoski |
| 2004 | Eucycloscala Kaim p. 17 |
| 2010 | Eucycloscala Bandel p. 437 |
| 2014 | Eucycloscala Nützel and Kaim p. 414 |
| 2020 | Eucycloscala Ketwetsuriya et al. p. 63 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Eucycloscala Cossmann 1895
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†Eucycloscala acanticum Uhlig 1881
†Eucycloscala asiatica Batten 1985
†Eucycloscala baltzeri Kittl 1892
†Eucycloscala basistriata Anderson 1975
†Eucycloscala binodosa Münster 1841
†Eucycloscala brunhuberi Schnittmann 1966
†Eucycloscala circumnodosa Kittl 1894
†Eucycloscala damesi Kittl 1892
†Eucycloscala exigua Healey 1908
†Eucycloscala hanwangensis Pan 1982
†Eucycloscala izabellae Kaim 2004
†Eucycloscala leo d'Orbigny 1847
†Eucycloscala margaritata Kittl 1900
†Eucycloscala medionodosa Batten 1985
†Eucycloscala nitida Dominici 2024
†Eucycloscala obliquicostata Yin and Yochelson 1983
†Eucycloscala pascoensis Haas 1953
†Eucycloscala praetor Goldfuss 1844
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Invalid names: Littorina ornata Jeffreys 1865 [synonym], Littorina phillipsii Morris and Lycett 1884 [synonym], Littorina praetor Goldfuss 1909 [synonym], Ooliticia phillipsii Morris and Lycett 1851 [synonym], Turbo ornatus Quenstedt 1853 [synonym], Turbo phillipsii Morris and Lycett 1851 [synonym]
†Eucycloscala semicancellata Kittl 1900
†Eucycloscala spinosa Klipstein 1843
†Eucycloscala spinulosa Klipstein 1843
†Eucycloscala subbisertus d'Orbigny 1849
†Eucycloscala supranodosa Klipstein 1843
†Eucycloscala ultima Ravn 1933
†Scalaria binodosa spinulosa Laube 1868
Diagnosis
No diagnoses are available
Measurements
| part | N | mean | minimum | maximum | median | s.d. | |
|---|---|---|---|---|---|---|---|
| height | 2 | 8.98 | 5.20 | 15.5 | 10.3 | 7.28 | |
| shell height | 28 | 5.86 | 0.800 | 19.4 | 7.15 | 4.88 | |
| shell diameter | 28 | 3.86 | 3.20 | 4.80 | 3.80 | 0.683 |
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| Source: superf = superfamily, c = class | |||||
| References: Kiessling 2004, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Kungurian to the top of the Selandian or 283.30000 to 59.24000 Ma
Minimum age of oldest fossil (stem group age): 274.4 Ma
Minimum age of oldest fossil (stem group age): 274.4 Ma
Collections (121 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Kungurian | Malaysia (Perak) | E. asiatica, E. medionodosa (62131) | |
| Wordian | Thailand (Ratchaburi) | E. sp. (40954) | |
| Capitanian | Japan | E. sp. (109667) | |
| Anisian | China (Guizhou) | E. binodosa (98055) | |
| Pelsonian | China (Guizhou) | Eucycloscala indet. (31610) | |
| Pelsonian - Illyrian | China (Guizhou) | E. binodosa, E. obliquicostata (44250) Eucycloscala indet., E. obliquicostata (31612) E. n. sp. A (31617) E. obliquicostata (31611 31628 44247) E. sp. (31632) E. sp., E. binodosa (31616) E. sp., E. binodosa, E. obliquicostata (31618 31619 31625 31631) E. sp., Eucycloscala indet. (31626) E. sp., Eucycloscala indet., E. binodosa, E. obliquicostata (31620) E. sp., Eucycloscala indet., E. obliquicostata (31624) E. sp., E. obliquicostata (31621 31622 31623 31627) E. spinulosa, E. obliquicostata (44252) | |
| Ladinian | Italy | Cyclonema circumnodosum (138255) | |
| Ladinian | Peru | E. exigua (206475) | |
| Longobardian | Italy | E. nitida (236003 236004 236005) | |
| Julian | Italy | E. binodosa (69650 69655 69658) E. binodosa, E. spinosa (137435) E. biserta (69659) E. biserta, E. binodosa (69668) E. biserta, E. sp. (69664 202422) E. ornata (69676) E. ornata, E. sp., E. damesi (34692 69661) E. sp. (69651 69666 69672 69679) E. sp., E. binodosa (69669) E. spinosa (163817) E. supranodosa, E. baltzeri (137437) Monodonta supranodosa (146388) Scalaria baltzeri (144927) Scalaria baltzeri, Trochus subbisertus, Scalaria supranodosa, Scalaria binodosa, Scalaria spinosa (144631) Scalaria damesi (144633) Trochus binodosus (137429) Trochus subbisertus, Scalaria supranodosa (144634) | |
| Carnian | Italy | E. binodosa (164208) E. ornata, E. binodosa (164191) | |
| Carnian | Hungary (Bakony) | E. binodosa, E. spinulosa, E. supranodosa, E. semicancellata, E. damesi (203210) E. damesi (203197) E. margaritata (203199) | |
| Carnian | Italy (South Tyrol) | Scalaria supranodosa (205149) | |
| Norian | Mexico (Sonora) | E. subbisertus (39172) | |
| Norian - Rhaetian | Peru | E. pascoensis (68098 68135 68138) | |
| Sevatian | Iran | E. epitoniformis (37959) | |
| Sevatian | USA (Idaho) | E. sp., E. binodosa (38377) | |
| Rhaetian | Peru | E. baltzeri, E. pascoensis (68096) E. pascoensis (68148) | |
| Angulata | Germany (Baden-Württemberg) | E. sp. (90442) | |
| Hettangian - Early/Lower Sinemurian | United Kingdom | E. sp. (118018) | |
| Obtusum | Germany | E. brunhuberi (151335) | |
| Late/Upper Pliensbachian | Germany (North Rhine-Westphalia) | E. dunkeri (225402) | |
| Margaritatus | United Kingdom | E. leo (118052) | |
| Spinatum | Germany | E. escheri, E. aedilis (159446) | |
| Spinatum | Germany (Bavaria) | E. escheri (25502) | |
| Early/Lower Aalenian | Germany | E. sp. (58216) | |
| Early/Lower Bajocian | Germany | E. praetor (150926 150927 150929 151069 151070) | |
| Humphresianum - Parkinsoni | Hungary | E. acanticum (91409) | |
| Early/Lower Bajocian - Late/Upper Oxfordian | Romania (Brasov) | Ooliticia phillipsii (81442) | |
| Late/Upper Bajocian | Poland | E. sp. 3 (63069 63071) | |
| Bathonian | Poland | Littorina praetor (91908) | |
| Late/Upper Bathonian | Poland | E. sp. (141288 141289 141290 141291 141292) | |
| Middle Bathonian | Poland | E. sp. (141265 141267 141269 141270 141277 141278 141279 141293) E. sp. 2, E. izabellae (63058) E. sp. 3 (63072) | |
| Middle Bathonian - Late/Upper Bathonian | Poland | E. sp. (141283) | |
| Bathonian - Callovian | Germany | E. aedilis (58398 58414) | |
| Callovian | Germany | E. aedilis (58203) | |
| Early/Lower Callovian | Germany | E. sp. (58212) | |
| Late/Upper Callovian | Poland | E. sp. 1 (63064) | |
| Late/Upper Oxfordian | Poland | E. augur (66972) | |
| Early/Lower Albian | United Kingdom (England) | E. mulleti (1864) | |
| Late/Upper Campanian | USA (Tennessee) | E. tuberculata (88357) | |
| Danian | Poland | E. crassilabris (27625) | |
| Danian | Denmark | E. ultima (137438 176994) | |
| Paleocene | Austria (Salzburg) | E. basistriata (84335) | |
| Paleocene | Germany | E. basistriata (3114) | |
| Selandian | Denmark | E. crassilabris (35263 35264 35265 35266 35267 51500) |