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Testudo brontops

Osteichthyes - Testudines - Testudinidae

Taxonomy
Testudo brontops was named by Marsh (1890). Its type specimen is YPM VP 000608 (holotype), a shell, and it is a 3D body fossil. Its type locality is Indian Creek (Testudo brontops type locality), which is in a Chadronian terrestrial horizon in the Chadron Formation of South Dakota.

It was recombined as Testudo (Hesperotestudo) brontops by Williams (1950), Williams (1953); it was recombined as Geochelone (Hesperotestudo) brontops by Auffenberg (1963); it was recombined as Hesperotestudo brontops by Hutchison (1996).

Synonymy list
YearName and author
1890Testudo brontops Marsh p. 179 figs. Pl. 8
1902Testudo brontops Hay p. 451
1904Testudo brontops Hay
1930Testudo brontops Hay p. 103
1950Testudo (Hesperotestudo) brontops Williams p. 20
1953Testudo (Hesperotestudo) brontops Williams p. 544
1963Geochelone (Hesperotestudo) brontops Auffenberg p. 87
1996Hesperotestudo brontops Hutchison p. 345
2018Testudo brontops Vlachos

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
RankNameAuthor
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
familyTestudinidaeBatsch 1788
subfamilyTestudininae
tribeTestudininiBatsch 1788
genusTestudo
speciesbrontops

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Testudo brontops Marsh 1890
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Diagnosis
ReferenceDiagnosis
E. Vlachos 2018“Testudo” brontops can be diagnosed as a member of Testudinidae based on the coincidence between the pleuro-marginal sulci and the costo-peripheral suture, the fused marginals XII, the well-differentiated neural series, and the presence of a well-developed epiplastral lip. “Testudo” brontops can be differentiated from stem and basal testudinids (e.g., Hadrianus, Oligopherus) based on the 4<8<4<8 anterior neural formula. “Testudo” brontops can be differentiated from other testudinids with a cervical scute (e.g., Gopherus, Stylemys, Hesperotestudo) based on the medially convex humero-pectoral sulcus that overlaps the posterior part of the entoplastron.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialf
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: herbivoref
Diet 2: herbivoref
Reproduction: oviparousf
Created: 2005-08-26 07:16:55
Modified: 2005-09-09 16:16:46
Source: f = family, c = class, subp = subphylum
References: Hendy et al. 2009, Carroll 1988, Ernst and Barbour 1989

Age range: Chadronian or 37.00000 to 33.90000 Ma

Collections: one only


Time interval Ma Country or state Original ID and collection number
Chadronian37.0 - 33.9USA (South Dakota) Testudo brontops (type locality: 191398)