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Testudo hexagonata

Reptilia - Testudines - Testudinidae

Taxonomy
Testudo hexagonata was named by Cope (1893). Its type specimen is TMM 30967-1817, a partial shell (shell fragments), and it is a 3D body fossil. Its type locality is Rock Creek, which is in an Irvingtonian terrestrial horizon in the Tule Formation of Texas.

It was considered a nomen vanum by Reynoso and Montellano-Ballesteros (2004); it was recombined as Gopherus hexagonatus by Vlachos (2018) and De La Garza et al. (2018).

Synonyms
Synonymy list
YearName and author
1893Testudo hexagonata Cope
1902Testudo hexagonata Hay
1908Testudo campester Hay
1937Bysmachelys canyonensis Johnston p. 447
1966Gopherus huecoensis Strain
1982Gopherus canyonensis Bramble p. 854
1982Gopherus hexagonata Bramble p. 854
2004Gopherus canyonensis Reynoso and Montellano-Ballesteros p. 824
2018Gopherus hexagonatus De La Garza et al.
2018Gopherus hexagonatus Vlachos

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
RankNameAuthor
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
Pantestudinidae
familyTestudinidaeBatsch 1788
genusTestudo
specieshexagonata

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
E. D. Cope 1893The lip of the plastron is not produced and bifurcate, as in T. agassizii
(from S. E. California), but the posterior lobe is quite similar to that of
the latter species. The posterior lobe of the T. laticaudata is intermediate
in proportions between those of T. hexagonata and T. polyphemus.
Both of the extinct species may belong to the genus Xerobates,
Ag., but this can not be determined until the crania are discovered.
The specimens of this species are smaller than the single one which
represents the T. laticaudata. Though allied to the latter, the forms of
the lobes of the plastron display too many differences to permit me to
regard them as identical.
E. Vlachos 2018Gopherus hexagonatus can be diagnosed as a member of Gopherus based on the characters mentioned above. Gopherus hexagonatus differs from other Gopherus to the exclusion of Gopherus donlaloi in its larger size and from Gopherus donlaloi in the presence of hexagonal neurals III–VIII.