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Chelonoidis hesterna

Reptilia - Testudines - Testudinidae

Taxonomy
Geochelone (Chelonoides) hesterna was named by Auffenberg (1971). Its type specimen is UCMP 40200, a skeleton (complete shell with skull, lower jaw, and most postcrania), and it is not a trace fossil. Its type locality is San Nicolas, which is in a Miocene terrestrial conglomerate/claystone in Colombia.

It was recombined as Geochelone hesterna by Crumly (1982), Riff et al. (2010), Sanchez-Villagra and Scheyer (2010); it was recombined as Chelonoidis hesterna by Auffenberg (1974), Franz and Franz (2009), de la Fuente et al. (2014), de la Fuente et al. (2018).

Synonymy list
YearName and author
1971Geochelone (Chelonoides) hesterna Auffenberg p. 106
1974Chelonoidis hesterna Auffenberg
1982Geochelone hesterna Crumly p. 215
2009Chelonoidis hesterna Franz and Franz
2010Geochelone hesterna Riff et al.
2010Geochelone hesterna Sanchez-Villagra and Scheyer
2014Chelonoidis hesterna de la Fuente et al.
2018Chelonoidis hesterna de la Fuente et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
familyTestudinidaeBatsch 1788
tribeGeochelonini
genusChelonoidis()
specieshesterna()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Chelonoidis hesterna Auffenberg 1971
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Diagnosis
ReferenceDiagnosis
M. S. de la Fuente et al. 2018A moderate-sized turtle, member of the clade Testudinidae based on the following characters: alternating pattern of the neural/costal associations; well-developed plastral buttresses that are in clear contact with the costal bones; coincidence of costo-peripheral sutures with pleuro–marginal sulci; fusion of twelfth marginal scutes; presence of a well-developed epiplastral lip; presence of a median maxillary ridge in the palate. Member of Chelonoidis based on the following characters: absence of cervical scute; first neural hexagonal with short postero-lateral sides; contact between pleural 3 and marginal 6. Differs from other members of Chelonoidis on the short and wide gular scutes and the presence of a circular deep pit in the premaxillae in ventral view. It is diagnosed as a member of the Ch. carbonaria group due to the shape of the shell, being narrow and high, with parallel edges when viewed from above, with a narrow posterior plastron lobe. It differs from Ch. carbonaria in the following characters: a smoother shell, without lateral incurving at the bridge; a proportional shorter anterior plastral lobe; gular scutes not reaching the entoplastron, humeral scutes expanded on the epiplastra; entoplastron longer than wide and a humeral scute that is longer along midline than the femoral scute. Chelonoidis hesterna differs from Ch. denticulata in possessing a smaller, wide axillary scar and an inguinal scale that is conspicuous in ventral view. The fibula and humerus are more curved in Ch. hesterna than in Ch. denticulata.