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Graptolithina (graptolite)
Taxonomy
Graptolithina was named by Bronn (1849). It is extant. It was considered monophyletic by Mitchell et al. (2013).
It was reranked as the class Graptolithina by Mitchell et al. (2013); it was reranked as the subclass Graptolithina by Maletz (2014), Maletz (2017), Toro and Herrera Sánchez (2019) and Lo Valvo et al. (2020).
It was assigned to Hemichordata by Bulman (1970), Toro (1993), Toro (1995) and Sepkoski (2002); and to Pterobranchia by Mitchell et al. (2013), Maletz (2014), Maletz (2017), Toro and Herrera Sánchez (2019) and Lo Valvo et al. (2020).
It was reranked as the class Graptolithina by Mitchell et al. (2013); it was reranked as the subclass Graptolithina by Maletz (2014), Maletz (2017), Toro and Herrera Sánchez (2019) and Lo Valvo et al. (2020).
It was assigned to Hemichordata by Bulman (1970), Toro (1993), Toro (1995) and Sepkoski (2002); and to Pterobranchia by Mitchell et al. (2013), Maletz (2014), Maletz (2017), Toro and Herrera Sánchez (2019) and Lo Valvo et al. (2020).
Subtaxa
Acanthastida, Ascograptus, Camaroidea, Ceramograptus, Crinocaulis, Crustoidea, Cyclograptidae, Dendroidea, Diplograptoidea, Dithecodendridae, Dithecodendrum, Dithecoidea, Graptoblasti, Graptolithus, Graptoloida, Graptoloidea, Graptovermida, Leveillites, Limpidograptus, Palmatophycus, Pilograptus, Rhabdopleuridae, Siryngotaenia, Stolonoidea, Thamnograptus, Trigonograptus, Tuboidea, Wimanicrustidae
Synonyms
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Synonymy list
Year | Name and author |
---|---|
1768 | Graptolithus Linnaeus |
1849 | Graptolithina Bronn |
1873 | Trigonograptus Lapworth |
1968 | Trigonograptus Whittington |
1970 | Graptolithina Bulman p. V17 |
1993 | Graptolithina Toro p. 71 |
1995 | Graptolithina Toro p. 377 |
2002 | Graptolithina Sepkoski |
2002 | Limpidograptus Sepkoski |
2002 | Thamnograptus Sepkoski |
2013 | Graptolithina Mitchell et al. |
2014 | Graptolithina Maletz p. 486 |
2017 | Graptolithina Maletz p. 114 |
2019 | Graptolithina Toro and Herrera Sánchez p. 502 |
2020 | Graptolithina Lo Valvo et al. p. 538 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subcl. Graptolithina Bronn 1849 [graptolite]
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G. †Ascograptus
Or. †Camaroidea
+
Fm. †Cysticamaridae Bulman 1955
+
G. †Cysticamara
G. †Flexicollicamara
G. †Graptocamara
G. †Tubicamara
G. †Syringotaenia
G. †Ceramograptus
G. †Crinocaulis
Or. †Crustoidea Bulman 1970
+
G. †Ruedemannicrusta Kozlowski 1962
G. †Urbanekicrusta
G. †Xenocyathus
Invalid names: Hormograptidae Bulman 1970 [empty]
Fm. †Cyclograptidae Bulman 1938
+
G. †Calycotubus
G. †Camarotubus Mierzejewski 2001
G. †Conitubus
G. †Cyclograptus
G. †Discograptus
G. †Galeograptus
G. †Kozlowskitubus
G. †Marsipograptus Ruedemann 1936
+
†Marsipograptus bullatus Ruedemann 1936
†Marsipograptus percensis Ruedemann 1936
G. †Parvitubus
G. †Reticulograptus
G. †Rhipidodendrum
G. †Tubidendrum
Or. †Dendroidea Nicholson 1872
+
Fm. †Acanthograptidae Bulman 1938
+
G. †Callograptus
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Subg. †Callograptus (Pseudocallograptus) Skevington 1963
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†Callograptus (Pseudocallograptus) salteri Hall 1865
†Callograptus cylindricus Liu 1987
†Callograptus elegans Hall 1865
†Callograptus praesalteri Rickards et al. 1990
Invalid names: Odontocaulis Lapworth 1881 [synonym]
G. †Coremagraptus
G. †Dictyonema Hall 1851
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†Dictyonema hamiltoniae Hall 1865
†Dictyonema macgillivrayi Hall 1899
†Dictyonema pulchellum Hall 1899
†Dictyonema ramosissimum Ruedemann 1947
†Dictyonema splendens Billings 1874
G. †Ptiograptus
G. †Aretograptus Obut and Sobolevskaya 1962
G. †Cactograptus
G. †Chaunograptus
G. †Coelograptus
Fm. †Dendrograptidae Roemer 1897
+
G. †Airograptus Ruedemann 1916
G. †Aspidograptus
+
†Aspidograptus implicatus Hopkinson 1875
†Aspidograptus smithtonensis Rickards et al. 1990
Invalid names: Clematograptus Hopkinson 1875 [replaced]
G. †Dendrograptus Hall 1858
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†Dendrograptus gracilis Hall 1865
†Dendrograptus hallianus Prout 1851
†Dendrograptus kindlei Ruedemann 1933
G. †Desmograptus
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†Desmograptus cancellatus Hopkinson 1875
†Desmograptus quiltyi Rickards et al. 1990
Invalid names: Rhizograptus Spencer 1878 [synonym]
G. †Graptolodendrum
G. †Licnograptus
G. †Ophigraptus
G. †Ptilograptus
Invalid names: Chaunograptidae Bulman 1955 [empty]
Fm. †Dithecodendridae Obut 1964
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G. †Archaeolafoea Chapman 1919
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†Archaeolafoea longicornis Chapman 1919
†Archaeolafoea monegettae Chapman 1919
G. †Bulmanidendrum Obut 1974
G. †Dalyia Walcott 1919
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†Dalyia annularoides Resser and Howell 1938
†Dalyia pennsylvanica Resser and Howell 1938
†Dalyia racemata Walcott 1919
G. †Karasidendrum Sennikov 1998
G. †Ovetograptus Sdzuy 1974
G. †Protodendrum Sennikov 1998
G. †Siberiodendrum Obut 1964
G. †Sibiriograptus Obut 1964
G. †Sotograptus Sdzuy 1974
G. †Tarnagraptus Sdzuy 1974
G. †Dithecodendrum
Unr. †Graptoloida Maletz et al. 2009 [graptoloid]
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Unr. †Eugraptoloida Maletz et al. 2009
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Invalid names: Panreclinata Maletz et al. 2009 [empty]
Or. †Graptoloidea Lapworth 1875
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G. †Apiograptus
G. †Arienigraptus
G. †Arnheimograptus
Subor. †Axonophora Frech 1897
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Infraor. †Diplograptina Lapworth 1880
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Superfm. †Climacograptoidea Frech 1897 [climacograptid]
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Fm. †Climacograptidae Frech 1807
G. †Undulograptus Bouček 1973
Invalid names: Climacograptacea Mitchell et al. 2007 [replaced]
Fm. †Dicranograptidae Lapworth 1873
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G. †Dicranograptus Hall 1865
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Invalid names: Cladograpsus Emmons 1855 [replaced], Diceratograptus [synonym]
Subfm. †Nemagraptinae Maletz 2014
Superfm. †Dicranograptoidea Lapworth 1873
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Invalid names: Dicranograptacea Storch et al. 2011 [replaced]
Fm. †Diplograptidae
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G. †Amplexograptus
G. †Anticostia Stewart and Mitchell 1997
G. †Cephalograptus
Subfm. †Diplograptinae Lapworth 1873
Subfm. †Orthograptinae Mitchell 1987
G. †Rectograptus Pribyl 1949
Invalid names: Orthograptidae Mitchell 1987 [synonym]
Fm. †Lasiograptidae Lapworth 1879 [lasiograptid]
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G. †Arachniograptus Ross and Berry 1963
G. †Archiretiolites Eisenack 1935
G. †Brevigraptus Mitchell 1988
G. †Gymnograptus
G. †Hallograptus Lapworth 1876
G. †Neurograptus Elles and Wood 1908
G. †Nymphograptus Elles and Wood 1908
G. †Orthoretiolites Whittington 1954
G. †Paraplegmatograptus Mu and Lin 1984
G. †Phormograptus Whittington 1955
G. †Pipiograptus Whittington 1955
G. †Plegmatograptus Elles and Wood 1908
G. †Sunigraptus Mu 1993
G. †Tysanograptus Elles and Wood 1908
G. †Yangzigraptus Mu 1983
G. †Yinograptus Mu 1962
Infraor. †Neograptina Storch et al. 2011
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Fm. †Normalograptidae Storch and Serpagli 1993
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G. †Cystograptus
G. †Hedrograptus Obut 1949
G. †Metaclimacograptus Bulman and Rickards 1968
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Invalid names: Lithuanograptus Paskevicius 1976 [synonym]
G. †Normalograptus Legrand 1987
G. †Pronormalograptus Chen et al. 2015
G. †Pseudoglyptograptus Bulman and Rickards 1968
G. †Rhaphidograptus
Superfm. †Retiolitoidea Lapworth 1873
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Fm. †Neodiplograptidae Melchin et al. 2011
Fm. †Retiolitidae Lapworth 1873
G. †Crinitograptus Rickards 1995
Subor. †Dichograptina Lapworth 1873
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Fm. †Dichograptidae Lapworth 1873
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Subfm. †Didymograpti Bulman 1970
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G. †Atopograptus
G. †Azygograptus Nicholson and Lapworth 1875
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Invalid names: Nicholsonograptus Boucek and Pribyl 1951 [synonym], Pseudazygograptus [synonym]
G. †Janograptus
G. †Maeandrograptus
G. †Oncograptus
G. †Parazygograptus
G. †Skiagraptus
G. †Didymograptus McCoy 1851
+
Subg. †Didymograptus (Didymograptellus) Cooper and Fortey 1982
Subg. †Didymograptus (Didymograptus) M'Coy 1851
†Didymograptus balticus Tullberg 1880
†Didymograptus demissus Törnquist 1901
†Didymograptus ensjoensis Monsen 1937
†Didymograptus euodus Lapworth 1875
†Didymograptus extensus Hall 1858
†Didymograptus filiformis Tullberg 1880
†Didymograptus holmi Törnquist 1901
†Didymograptus latus Hall 1907
†Didymograptus murchisoni Beck 1839
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Invalid names: Didymograptus pseudogeminus Bouček 1973 [synonym]
†Didymograptus nicholsoni Lapworth 1875
†Didymograptus pacificus Ruedemann 1947
†Didymograptus paraindentus Berry 1960
†Didymograptus paraindentus Berry 1960
†Didymograptus protobalticus Monsen 1937
†Didymograptus pusillus Tullberg 1880
†Didymograptus spinulosus Perner 1895
†Didymograptus suecicus Tullberg 1880
†Didymograptus turgidus Lee 1974
†Didymograptus undulatus Törnquist 1901
†Didymograptus uniformis Elles and Wood 1901
Invalid names: Cladograpsus Geinitz 1852 [synonym]
Subfm. †Goniograpti Bulman 1970
+
G. †Anomalograptus
G. †Brachiograptus
G. †Oslograptus
G. †Stellatograptus
G. †Triaenograptus
G. †Yushanograptus
G. †Zygograptus
G. †Loganograptus Hall 1868
G. †Orthodichograptus Thomas 1972
Subfm. †Schizograpti Bulman 1970
+
G. †Anthograptus Tornquist 1904
G. †Holograptus
G. †Mimograptus
G. †Schizograptus
G. †Trochograptus
Invalid names: Dichograpti Bulman 1970 [empty], Dichograptinae Lapworth 1873 [empty], Tetragrapti Bulman 1970 [empty]
Fm. †Phyllograptidae Kiessling 2009
+
G. †Harrisgraptus VandenBerg 2019
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†Harrisgraptus eocaduceus Harris 1933
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Invalid names: Harrisgraptus hemicyclus Harris 1933 [synonym]
G. †Phyllograptus Hall 1858
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†Phyllograptus anna Hall 1865
†Phyllograptus ilicifolius Hall 1858
†Phyllograptus loringi White 1874
G. †Pseudophyllograptus Cooper and Fortey 1982
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†Pseudophyllograptus angustifolius Hall 1857
†Pseudophyllograptus densus Tornquist 1879
G. †Tetragraptus Salter 1863
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†Tetragraptus acclinans Keble 1920
†Tetragraptus akzharensis Tzaj 1968
†Tetragraptus amii Elles and Wood 1902
†Tetragraptus bigsbyi Hall 1865
†Tetragraptus decipiens Hall 1899
†Tetragraptus insuetus Keble and Benson 1929
†Tetragraptus phyllograptoides Strandmark 1902
†Tetragraptus pseudobigsbyi Skevington 1965
†Tetragraptus quadribrachiatus Hall 1858
†Tetragraptus reclinatus Elles and Wood 1902
Fm. †Pterograptidae Mu 1950
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G. †Didymograptellus Cooper and Fortey 1982
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†Didymograptellus bifidus Hall 1865
†Didymograptellus nitidus Hall 1858
†Didymograptellus protobifidus Elles 1933
G. †Pseudobryograptus Mu 1957
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†Pseudobryograptus incertus Harris and Thomas 1935
†Pseudobryograptus parallelus Mu 1957
G. †Xiphograptus
Fm. †Tetragraptidae Frech 1897
+
+
†Pseudotrigonograptus ensiformis Hall 1858
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Invalid names: Pseudotrigonograptus angustus Mu and Lee 1958 [synonym], Pseudotrigonograptus magnus Mu and Lee 1958 [synonym], Pseudotrigonograptus uniformis Mu and Lee 1958 [synonym]
Fm. †Didymograptidae Mu 1950
+
G. †Baltograptus Maletz 1994
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†Baltograptus deflexus Elles and Wood 1901
†Baltograptus extremus Maletz and Slovacek 2013
†Baltograptus geometricus Törnquist 1901
†Baltograptus jacksoni Rushton 2011
†Baltograptus kurcki Törnquist 1901
†Baltograptus minutus Törnquist 1879
†Baltograptus vacillans Tullberg 1880
G. †Expansograptus Bouček and Přibyl 1953
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†Didymograptus (Expansograptus) validus Törnquist 1901
†Expansograptus abnormis Hsu 1934
†Expansograptus hirundo Salter 1863
†Expansograptus rigoletto Maletz et al. 1991
Subor. †Didymograptina Lapworth 1875
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G. †Eisenackograptus
G. †Eoglyptograptus
G. †Euclimacograptus
G. †Exigraptus
Subor. †Glossograptina Jaanuson 1960
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Fm. †Glossograptidae Lapworth 1873
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G. †Lonchograptus
G. †Nanograptus
+
†Paraglossograptus tentaculatus Hall 1858
+
Invalid names: Paraglossograptus etheridgei Harris 1924 [synonym]
Fm. †Isograptidae Harris 1933
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G. †Isograptus Moberg 1892
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†Isograptus gibberulus Nicholson 1875
†Isograptus victoriae Harris 1933
Invalid names: Arienigraptidae Yu and Fang 1981 [synonym]
Invalid names: Bireclinata Maletz et al. 2009 [synonym]
Subor. †Graptodendroidina Mu and Lin 1981
+
Fm. †Anisograptidae Bullman 1950
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G. †Adelograptus Bulman 1941
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†Adelograptus antiquus Hall 1899
†Adelograptus brevibrachiatus Cho and Kim 2007
†Adelograptus clarki Hall 1899
†Adelograptus tenellus Linnarsson 1871
+
Invalid names: Adelograptus hunnebergensis Moberg 1892 [synonym]
G. †Ancoragraptus Jackson and Lenz 2003
G. †Anisograptus Ruedemann 1937
+
†Anisograptus compactus Cooper and Stewart 1979
†Anisograptus delicatulus Cooper and Stewart 1979
†Anisograptus matanensis Ruedemann 1937
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Invalid names: Anisograptus flexuosus Bulman 1941 [synonym], Anisograptus norvegicus Bulman 1954 [synonym], Anisograptus primigenius Bulman 1950 [synonym], Anisograptus richardsoni Bulman 1941 [synonym]
G. †Choristograptus
G. †Dictyograptus Hopkinson and Lapworth 1875
G. †Kiaerograptus
+
†Kiaerograptus bulmani Thomas 1973
†Kiaerograptus quasimodo Rushton 1981
†Kiaerograptus supremus Lindholm 1991
G. †Paratemnograptus
G. †Psigraptus Jackson 1967
+
†Psigraptus arcticus Jackson 1967
†Psigraptus jacksoni Rickards and Stait 1984
†Psigraptus lenzi Jackson 1967
+
Invalid names: Psigraptus quadribrachiatus Zhao and Zhang 1985 [synonym]
†Psigraptus magnus Zhao and Zhang 1986
Invalid names: Diphygraptus Zhao and Zhang 1985 [synonym], Holopsigraptus Zhao and Zhang 1985 [synonym], Muenzigraptus Zhao and Zhang 1985 [synonym], Neoclonograptus Zhao and Zhang 1985 [synonym], Yukonograptus Lin 1981 [synonym]
G. †Radiograptus
G. †Rhabdinopora Eichwald 1860
+
†Rhabdinopora enigma Cooper and Stewart 1979
†Rhabdinopora flabelliformis Eichwald 1840
†Rhabdinopora umbellata Liu 1987
Invalid names: Heterograptus Lin 1986 [synonym]
G. †Staurograptus
+
†Staurograptus dichotomus Emmons 1855
+
Invalid names: Staurograptus apertus Jackson and Lenz 1962 [synonym], Staurograptus tenuis Jackson 1974 [synonym]
†Staurograptus mojotorensis Monteros and Moya 2011
Invalid names: Aletograptus Obut and Sobolevskaya 1962 [synonym]
G. †Triograptus
Invalid names: Nephelograptidae Boucek 1957 [synonym]
Fm. †Mastigograptidae Bates and Urbanek 2002
+
G. †Mastigograptus Ruedemann 1908
+
†Mastigograptus serialis Chapman and Thomas 1936
†Mastigograptus tenuiramosus Walcott 1883
G. †Micrograptus Eisenack 1974
G. †Hirsutograptus
G. †Joamgsjamotes
G. †Metabrograptus
Subor. †Monograptina Lapworth 1880
+
Fm. †Cryptograptidae Boucek 1933
+
G. †Neocolonograptus Urbanek 1997
+
†Neocolonograptus lochkovensis Přibyl 1940
†Neocolonograptus parultimus Jaeger 1975
†Neocolonograptus ultimus Perner 1899
G. †Neolobograptus
G. †Parapetalolithis Koren' and Rickards 1996
G. †Pararetiograptus
G. †Pseudoisograptus
G. †Rivagraptus
+
†Rivagraptus bellulus Törnquist 1890
†Rivagraptus kayi Churkin and Carter 1970
†Rivagraptus supernus Fu and Song 1986
Subor. †Sinograpta Maletz 2009
+
Fm. †Sinograptidae Mu 1957
+
G. †Allograptus
G. †Holmograptus
+
†Holmograptus bovis Williams and Stevens 1988
†Holmograptus spinosus Ruedemann 1904
G. †Sinograptus Mu 1957
+
†Sinograptus rastritoides Mu 1957
†Sinograptus typicalis Mu 1957
+
Invalid names: Sinograptus aequalis Mu 1957 [synonym], Sinograptus declinatus Yu and Fang 1983 [synonym]
G. †Tylograptus Mu 1957
+
†Tylograptus globiformis Mu 1957
†Tylograptus intermedius Mu 1957
+
Invalid names: Tylograptus grandis Hsu and Chao 1976 [synonym], Tylograptus multidentatus Hsu and Chao 1976 [synonym]
Subor. †Sinograptina Maletz 2009
+
Fm. †Sigmagraptidae Cooper and Fortey 1982
+
G. †Eotetragraptus Bouček and Přibyl 1952
G. †Praegoniograptus Rickards and Chapman 1991
G. †Sigmagraptus Ruedemann 1904
+
G. †Sinoreteograptus
G. †Stromatograptus
G. †Urbanekograptus
Invalid names: Bryograptidae Obut 1957 [empty], Dichograptacea Lapworth 1873 [empty], Virgellina Fortey and Cooper 1986 [empty]
G. †Leveillites
G. †Pilograptus
Fm. †Rhabdopleuridae Harmer 1905
+
G. †Calyxhydra Kozlowski 1959
G. †Chitinodendron Eisenack 1938
G. Cylindrotheca Rabenhorst 1848
G. †Epigraptus
G. †Fasciculitubus
G. †Graptovermis
G. †Haplograptus Ruedemann 1933
+
†Haplograptus wisconsinensis Ruedemann 1933
†Haplograptus wisconsinensis montanensis Lochman 1964
G. †Kystodendron
G. †Rhabdopleurites
G. †Rhabdopleuroides
G. †Rhabdotubus
G. Rhapdopleura Allman 1869
G. †Stolonodendrum
G. †Siryngotaenia
Fm. †Wimanicrustidae Bulman 1970
+
G. †Bulmanicrusta Kozlowski 1962
G. †Ellesicrusta
G. †Holmicrusta
G. †Hormograptus
G. †Lapworthicrusta
G. †Wimanicrusta
Invalid names: Diplograptoidea Lapworth 1880 [empty], Dithecoidea Obut 1964 [empty], Graptolithus Linnaeus 1768 [invalid subgroup], Graptovermida Kozlowski 1949 [empty], Limpidograptus [nomen vanum], Stolonoidea Kozlowski 1938 [empty], Thamnograptus [nomen vanum], Trigonograptus Lapworth 1873 [nomen vanum], Tuboidea Kozlowski 1938 [empty]
Diagnosis
No diagnoses are available