Huangbanjigou, Chaomidian Village (CNU collection) (Cretaceous of China)

Where: Liaoning, China (41.6° N, 120.8° E)

• Paleocoordinates: 41.7° N, 128.6° E (Wright 2013)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Yixian Formation (Jehol Group), Late/Upper Barremian (130.0 - 125.5 Ma)

• The Yixian Formation can be subdivided into five units, in ascending order, Lujiatun unit, Lower Lava, Jianshangou unit, Upper Lava, Dawangzhangzi unit, Jingangshan unit, and Huanghuashan unit. Most of the Jehol fossils from the Yixian Formation are preserved in four beds, that is, the Lujiatun bed (recalculated age is 124.0 ± 1.0/1.3 Ma, He et al., 2006), the Jianshangou bed (125.43 ± 0.50/0.88 Ma, Swisher et al., 1999), the Dawangzhangzi bed, and Jinggangshan bed (~122 Ma, Smith et al., 1995).U-Pb dating suggests that the Huangbanjigou section ranges from 124.9 ± 1.7 Ma to 122.8 ± 1.6 Ma (Yang et al., 2007). Chang et al. (2017) obtained Ar/Ar ages of 125.8 ± 1.0 Ma and 126.0 ± 0.8 Ma from the Lujiatun Unit, the lowest part of the Yixian Formation, below the lacustrine shales.

•He et al. (2004) reported a 40Ar/39Ar age of 120.3 ± 0.7 Ma for a tuff layer in the lower part of the Jiufotang Formation, and Chang et al. (2009) obtained a 40Ar/39Ar age of 122.1 ± 0.3 Ma for a tuff layer from the lowermost part of the Jiufotang Formation. Yu et al. (2021) dated an ash bed just above the Yixian/Jiufotang formation boundary to 122.0 ± 0.9 Ma and three tuff beds from the Xiaotaizi section to 118.9 ± 0.8 Ma, 118.8 ± 0.6 Ma, and 118.6 ± 1.1 Ma. These ages suggest that the Jiufotang Formation in Jianchang Basin was deposited at ca. 122.0–118.9 Ma.

•Olierrok et al. (2019) used a selection of U-Pb and 40Ar-39Ar geochronology to place the Barremian–Aptian boundary (the base of the Chron M0r) between 123.8 and 121.8 Ma. The GTS 2020 uses an age of 121.4 Ma for the base of the Aptian, and multiproxy orbital tuning gives an age of 120.2 Ma for the Barremian-Aptian boundary (Leandro et al., 2022). Therefore, the Yixian Formation should be entirely Late Barremian, with the Jiufotang Formation deposited in the Early Aptian (or latest Barremian at its base).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, tuffaceous mudstone

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Capital Normal University, Beijing

Primary reference: T. T. Wang, D. Ren, J. H. Liang and C. K. Shih. 2007. New Mesozoic cockroaches (Blattaria: Blattulidae) from Jehol Biota of western Liaoning in China. Annales Zoologici 57:483-495 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111710: authorized by Matthew Clapham, entered by Matthew Clapham on 28.06.2011, edited by Jered Karr

Creative Commons license: CC0 (CC0)

Taxonomic list

Insecta
 Blattaria - Blattulidae
Macaroblattula ellipsoides n. gen. n. sp. Wang et al. 2007 cockroach
Habroblattula drepanoides n. gen. n. sp.30 Wang et al. 2007 cockroach
CNU-B-LB2006369-1, 305, 378, 386, 389, 392, 301-304, 369-2, 390-1, 390-2, 306, 391
Elisama cuboides n. sp. Wang et al. 2007 cockroach
Elisama extenuata Ren 1995 cockroach
3 forewings, 13 complete specimens
 Hemiptera - Vetanthocoridae
Curticerus venustus n. gen. n. sp.33, Crassicerus furtivus n. gen. n. sp.33, Pustulithoracalis gloriosus n. gen. n. sp.33, Vetanthocoris decorus n. gen. n. sp.33, Vetanthocoris longispicus33, Curvicaudus ciliatus n. gen. n. sp.33, Byssoidecerus levigatus n. gen. n. sp.33, Collivetanthocoris rapax n. gen. n. sp.33, Mecopodus xanthos n. gen. n. sp.33
Curticerus venustus n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006083, 084
Crassicerus furtivus n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006032
Pustulithoracalis gloriosus n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006034, 037
Vetanthocoris decorus n. gen. n. sp.33 Yao et al. 2006 true bug
CHU-HE-LB2006058 (holotype)
Vetanthocoris longispicus33 Yao et al. 2006 true bug
CNU-HE-LB2006044,092,093,094
Curvicaudus ciliatus n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006023,024,022,029,030,033,0364
Byssoidecerus levigatus n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006025, 026, 035
Collivetanthocoris rapax n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006057, 050, 067
Mecopodus xanthos n. gen. n. sp.33 Yao et al. 2006 true bug
CNU-HE-LB2006031, 027
 Hemiptera - Torrirostratidae
Torrirostratus pilosus n. gen. n. sp.37, Flexicorpus acutirostratus n. gen. n. sp.37
Torrirostratus pilosus n. gen. n. sp.37 Yao et al. 2014 true bug
CNU-HE-LB2006506, 371/404, 318/373, 362, 364, 367, 368, 369, 370, 372; 2010044, 136, 137, 138, 139, 140, 141, 182, 183
Flexicorpus acutirostratus n. gen. n. sp.37 Yao et al. 2014 true bug
CNU-HE-LB2006435
 Hemiptera - Saldidae
Propritergum opimum n. gen. n. sp.40 Zhang et al. 2013 shore bug
CNU-HET-LB2006653
Mesolygaeus laiyangensis40 Ping 1928 shore bug
CNU-HET-LB2006657-658 (part and counterpart),200647
Luculentsalda maculosa n. gen. n. sp.41 Zhang et al. 2013 shore bug
CNU-HET-LB2006648, 651, 650
 Hemiptera - Naucoridae
Exilcrus cameriferus n. gen. n. sp.42, Miroculus laticephalus n. gen. n. sp.42
Exilcrus cameriferus n. gen. n. sp.42 Zhang et al. 2011 creeping water bug
Miroculus laticephalus n. gen. n. sp.42 Zhang et al. 2011 creeping water bug
 Hemiptera - Ochteridae
Floricaudus multilocellus n. gen.39, "Angulochterus quatrimaculatus n. gen. n. sp." = Angulochterus quadrimaculatus39, Pristinochterus zhangi n. gen. n. sp.35, Pristinochterus ovatus n. sp.39
Floricaudus multilocellus n. gen.39 Yao et al. 2011 velvety shore bug
"Angulochterus quatrimaculatus n. gen. n. sp." = Angulochterus quadrimaculatus39 Yao et al. 2011 velvety shore bug
Pristinochterus zhangi n. gen. n. sp.35 Yao et al. 2007 velvety shore bug
CNU-HE-LB2006301-302
Pristinochterus ovatus n. sp.39 Yao et al. 2011 velvety shore bug
CNU-HE-LB2009096
 Hemiptera - Primipentatomidae
Oropentatoma epichara n. gen. n. sp.32, Primipentatoma peregrina n. gen. n. sp.32, Primipentatoma peregrina32, Quadrocoris radius n. gen. n. sp.32, Breviscutum lunatum n. gen. n. sp.32
Oropentatoma epichara n. gen. n. sp.32 Yao et al. 2013 true bug
CNU-HE-LB2006768(769), 770
Primipentatoma peregrina n. gen. n. sp.32 Yao et al. 2013 true bug
CNU-HE-LB2006001-6,9,11-15,17-20,96,97,764,766,772,773,775,777
Primipentatoma peregrina32 Yao et al. 2013 true bug
CNU-HE-LB2009019-027
Quadrocoris radius n. gen. n. sp.32 Yao et al. 2013 true bug
CNU-HE-LB2009017
Breviscutum lunatum n. gen. n. sp.32 Yao et al. 2013 true bug
CNU-HE-LB2006007
 Hemiptera - Cydnidae
Orienicydnus hongi n. gen. n. sp.34, Cilicydnus robustispinus n. gen. n. sp.34
Orienicydnus hongi n. gen. n. sp.34 Yao et al. 2007 burrowing bug
CNU-HE-NN2006230
Cilicydnus robustispinus n. gen. n. sp.34 Yao et al. 2007 burrowing bug
CNU-HE-NN2006297(298), 299
 Hemiptera - Kobdocoridae
Clavaticoris zhengi n. gen. n. sp.38, Halonatusivena nervosus n. sp.5, Halonatusivena shii n. gen. n. sp.5, Venicoris solaris n. gen. n. sp.38
Clavaticoris zhengi n. gen. n. sp.38 Yao et al. 2012 true bug
CNU-HE-LB2009016
Halonatusivena nervosus n. sp.5 Du et al. 2016 true bug
CNU-HE-LB2006356, 176, 218, 355
Halonatusivena shii n. gen. n. sp.5 Du et al. 2016 true bug
CNU-HE-LB2006354, 335-340, 342-344, 346, 349-353, 382, 384
Venicoris solaris n. gen. n. sp.38 Yao et al. 2012 true bug
CNU-HE-LB2006526, 508, 509, 511-513, 515-525, 527-624, 626, 628, 630, 632, 634, 636, 638, 640, 642
 Hemiptera - Pachymeridiidae
Bellicoris mirabilis n. gen. n. sp.36, Beipiaocoris multifurcus n. gen. n. sp.36
Bellicoris mirabilis n. gen. n. sp.36 Yao et al. 2008 true bug
CNU-HE-LB2006267, 197, 205, 263-296
Beipiaocoris multifurcus n. gen. n. sp.36 Yao et al. 2008 true bug
CNU-HE-LB2006193, 177-192, 194-196, 198-209, 211-262
 Hemiptera - Yuripopovinidae
Changirostrus maculatus n. gen. n. sp.6, Dehiscensicoris sanctus n. gen. n. sp.6, Crassiantenninus minutus n. gen. n. sp.6
Changirostrus maculatus n. gen. n. sp.6 Du et al. 2016 true bug
CNU-HE-LB2006365, 363/388
Dehiscensicoris sanctus n. gen. n. sp.6 Du et al. 2016 true bug
CNU-HE-LB2006152, 99/101/103/105/107/109/111/113/115/117/119/121/123/127-129/131-136/139-145/147-153/156/157/159-165/167/169-174/320-322
Crassiantenninus minutus n. gen. n. sp.6 Du et al. 2016 true bug
CNU-HE-LB2006130, 125/135/146/154/155/323
 Coleoptera - Lasiosynidae
Lasiosyne integera n. sp.11 Guo et al. 2022 beetle
CNU-COL-LB-20069861
 Coleoptera - Caridae
"Leptocar polychaetus n. gen. n. sp." = Montsecanomalus zherikhini15, Cretonanophyes punctatus n. sp.15, Abrocar macilentus n. sp.15
"Leptocar polychaetus n. gen. n. sp." = Montsecanomalus zherikhini15 Liu and Ren 2006 weevil
CNU-C-LB2006101
Cretonanophyes punctatus n. sp.15 Liu and Ren 2007 weevil
CNU-LB-2006103
Abrocar macilentus n. sp.15 Liu and Ren 2007 weevil
CNU-C-LB2006102
 Coleoptera - Mordellidae
Bellimordella robusta n. sp.17 Liu et al. 2008 tumbling flower beetle
CNU-C-LB2006204
Bellimordella longispina n. sp.17 Liu et al. 2008 tumbling flower beetle
CNU-C-LB2006203
Bellimordella capitulifera n. gen. n. sp.17 Liu et al. 2008 tumbling flower beetle
CNU-C-LB2006202
Mirimordella gracilicruralis n. gen. n. sp.14 Liu et al. 2007 tumbling flower beetle
CNU-C-LB2006201
 Coleoptera - Pleocomidae
Pleocoma dolichophylla n. sp.20 Nikolajev and Ren 2012 rain beetle
CNU-COL-LB2009725
 Coleoptera - Glaphyridae
Glaphyrus ancestralis n. sp.19, Mesohypna lopatini n. gen. n. sp.21
Glaphyrus ancestralis n. sp.19 Nikolajev and Ren 2011 bumble bee scarab beetle
CNU-COL-LB2009604, 662
Mesohypna lopatini n. gen. n. sp.21 Nikolajev and Ren 2013 bumble bee scarab beetle
CNU-COL-LB2009710
 Coleoptera - Staphylinidae
Cretoquedius infractus n. sp.25 Solodovnikov and Yue 2013 rove beetle
CNU-COL-LB200951
 Coleoptera - Hydrophilidae
Hydroyixia elongata n. gen. n. sp.8, Hydroyixia latissima n. sp.8, Alegorius sp.8, Alegorius yixianus n. gen. n. sp.8
Hydroyixia elongata n. gen. n. sp.8 Fikáček et al. 2014 water scavenger beetle
CNU-2010005, 2009075
Hydroyixia latissima n. sp.8 Fikáček et al. 2014 water scavenger beetle
CNU-COL-LB2009074, 107, 2010014
Alegorius sp.8 Fikáček et al. 2014 water scavenger beetle
CNU-2009081
Alegorius yixianus n. gen. n. sp.8 Fikáček et al. 2014 water scavenger beetle
CNU-2009079, 078, 080, 077, 2010024, 025
 Coleoptera - Elateridae
Paralithomerus parallelus n. sp.4 Chang et al. 2008 click beetle
CNU-C-LB2006872
Paralithomerus exquisitus n. gen. n. sp.4 Chang et al. 2008 click beetle
CNU-C-LB2006874
Curtelater wui n. gen. n. sp.2 Chang and Ren 2008 click beetle
CNU-C-LB2006830
Cryptocoelus gianteus n. sp.3 Chang et al. 2007 click beetle
CNU-C-LB2006851
Bilineariselater foveatus n. gen. n. sp.2 Chang and Ren 2008 click beetle
CNU-C-LB2006801
 Coleoptera - Eucnemidae
Apoclion antennatus n. sp.1 Chang et al. 2010 false click beetle
Paradesmatus dilatatus n. sp.1 Chang et al. 2010 false click beetle
Anoixis complanus n. gen. n. sp.1 Chang et al. 2010 false click beetle
 Coleoptera - Artematopodidae
Forticatinius elegans n. gen. n. sp.26
Forticatinius elegans n. gen. n. sp.26 Tan and Ren 2007 soft bodied plant beetle
CNU-C-LB2006012
 Coleoptera - Ommatidae
"Tetraphalerus latus n. sp." = Allophalerus latus28, "Brochocoleus angustus n. sp." = Pareuryomma angustum29, "Brochocoleus sulcatus n. sp." = Odontomma sulcatum29, "Tetraphalerus curtinervis n. sp." = Monticupes curtinervis28
"Tetraphalerus latus n. sp." = Allophalerus latus28 Tan et al. 2007 beetle
CNU-C-LB2006005
"Brochocoleus angustus n. sp." = Pareuryomma angustum29 Tan et al. 2007 beetle
CNU-C-LB2006009, 055
"Brochocoleus sulcatus n. sp." = Odontomma sulcatum29 Tan et al. 2007 beetle
CNU-C-LB2006011
"Tetraphalerus curtinervis n. sp." = Monticupes curtinervis28 Tan et al. 2007 beetle
CNU-C-LB2006006
 Coleoptera - Cupedidae
"Priacma clavata n. sp." = Apriacma clavata27, "Priacma renaria n. sp." = Apriacma renaria27
"Priacma clavata n. sp." = Apriacma clavata27 Tan et al. 2006 reticulated beetle
CNU-C-LB2006001
"Priacma renaria n. sp." = Apriacma renaria27 Tan et al. 2006 reticulated beetle
CNU-C-LB2006002
 Coleoptera - Dytiscidae
Sinoporus lineatus n. gen. n. sp.22, Liadroporus elegans n. gen. n. sp.22, Liadytiscus cretaceus n. gen. n. sp.22, Liadytiscus longitibialis n. sp.22, Liadytiscus latus n. sp.22, Mesoderus ventralis n. sp.22, Mesoderus magnus n. gen. n. sp.22
Sinoporus lineatus n. gen. n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009104
Liadroporus elegans n. gen. n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009030
Liadytiscus cretaceus n. gen. n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009028
Liadytiscus longitibialis n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009026
Liadytiscus latus n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009031
Mesoderus ventralis n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009029
Mesoderus magnus n. gen. n. sp.22 Prokin and Ren 2010 predaceous diving beetle
CNU-COL-LB2009033
 Neuroptera - Kalligrammatidae
Oregramma illecebrosa n. sp.31 Yang et al. 2014 lacewing
CNU-NEU-LB2009031
 Neuroptera - Mesochrysopidae
Longicellochrysa yixiana n. gen. n. sp.23
Longicellochrysa yixiana n. gen. n. sp.23 Ren et al. 2010 lacewing
CNU-NEU-LJ2009012 (locality given as Chaomidian village)
 Hymenoptera - Pelecinidae
Eopelecinus huangi n. sp.13, Eopelecinus tumidus n. sp.13, Shoushida infera n. sp.10, Shoushida regilla n. gen. n. sp.16, Azygopelecinus clavatus n. gen. n. sp.7, Megapelecinus nashi n. sp.24, Megapelecinus changi n. gen. n. sp.24, Sinopelecinus viriosus7, Abropelecinus annulatus n. gen. n. sp.7
Eopelecinus huangi n. sp.13 Liu et al. 2011 wasp
Eopelecinus tumidus n. sp.13 Liu et al. 2011 wasp
Shoushida infera n. sp.10 Guo et al. 2016 wasp
CNU-HYM-LB-2006105
Shoushida regilla n. gen. n. sp.16 Liu et al. 2009 wasp
CNU-HYM-LB2006074
Azygopelecinus clavatus n. gen. n. sp.7 Feng et al. 2010 wasp
Megapelecinus nashi n. sp.24 Shih et al. 2010 wasp
Megapelecinus changi n. gen. n. sp.24 Shih et al. 2010 wasp
Sinopelecinus viriosus7 Zhang et al. 2002 wasp
Abropelecinus annulatus n. gen. n. sp.7 Feng et al. 2010 wasp
 Hymenoptera - Anaxyelidae
Hemisyntexis lepida n. gen. n. sp.12 Li et al. 2024 winged insect
CNU-HYM-LB2024501
Eosyntexis conflata n. sp.12 Li et al. 2024 winged insect
CNU-HYM-LB2024502
Parasyntexis khasurtensis12 Kopylov 2019 winged insect
CNU-HYM-LB2024503
 Orthoptera - Prophalangopsidae
Allaboilus hani n. sp.9, "Parahagla sibirica" = Parahagla sibirica9, "Grammohagla latibasis n. sp." = Parahagla sibirica18, "Trachohagla jeholia n. gen. n. sp." = Parahagla sibirica18, "Chifengia angustata n. sp." = Parahagla sibirica18, "Chifengia lata n. sp." = Parahagla sibirica18, Ashanga hongi9
Allaboilus hani n. sp.9 Gu et al. 2010 hump-winged cricket
CNU-ORT-LB2009049
"Parahagla sibirica" = Parahagla sibirica9 Sharov 1968 hump-winged cricket
CNU-ORT-LB2006020, 054, 059, 103, 105, 106, 2008079, 081, 2009050-52, 079
"Grammohagla latibasis n. sp." = Parahagla sibirica18 Sharov 1968 hump-winged cricket
CNU-O-LB2006007
"Trachohagla jeholia n. gen. n. sp." = Parahagla sibirica18 Sharov 1968 hump-winged cricket
CNU-O-LB2006008
"Chifengia angustata n. sp." = Parahagla sibirica18 Sharov 1968 hump-winged cricket
CNU-O-LB2006006
"Chifengia lata n. sp." = Parahagla sibirica18 Sharov 1968 hump-winged cricket
CNU-O-LB2006009
Ashanga hongi9 Meng and Ren 2006 hump-winged cricket