Karatau-Mikhailovka (PIN collection 2784) (Jurassic to of Kazakhstan)

Where: Chimkent, Kazakhstan (42.9° N, 70.0° E: paleocoordinates 41.6° N, 73.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karabastau Formation, Callovian to Callovian (166.1 - 157.3 Ma)

Environment/lithology: lacustrine - large; lithified siltstone

• "dark shales" Grimaldi & Engel, 2005

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Reposited in the PIN

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 113439: authorized by Matthew Clapham, entered by Jered Karr on 31.07.2011, edited by Matthew Clapham, Andrew Spencer-Lee and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Dicondylia - Protomyrmeleontidae
Protomyrmeleon sp.34 Geinitz 1888 winged insect
PIN 2784/69
Protomyrmeleon grandis n. sp.34 Nel et al. 2005 winged insect
PIN 2784/67
Protomyrmeleon kazakhstanensis n. sp.34 Nel et al. 2005 winged insect
PIN 2784/70-71(pc)
 Odonata - Paragonophlebiidae
Paragonophlebia inexpectata n. gen. n. sp.33
Paragonophlebia inexpectata n. gen. n. sp.33 Nel 2009 dragonfly
PIN 2784/08
 Hymenoptera - Anaxyelidae
Sphenosyntexis pallicornis n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1162
Brachysyntexis nova n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1159
Urosyntexis depressa n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1160
 Hymenoptera - Xyelidae
Microxyelecia brachycera n. gen. n. sp.39, Sirecomima xiphophora n. gen. n. sp.39
Microxyelecia brachycera n. gen. n. sp.39 Rasnitsyn 1969 sawfly
PIN 2784/1172
Sirecomima xiphophora n. gen. n. sp.39 Rasnitsyn 1969 sawfly
PIN 2784/1173
 Hymenoptera - Karatavitidae
Karatavites medius n. sp.39 Rasnitsyn 1969 wood wasp
PIN 2784/1169
 Hymenoptera - Mesoserphidae
Campturoserphus obscurus n. gen. n. sp.42, Auliserphus niger n. sp.42
Campturoserphus obscurus n. gen. n. sp.42 Rasnitsyn 1986 wasp
PIN 2784/1236
Auliserphus niger n. sp.42 Rasnitsyn 1986 wasp
PIN 2784/1227
 Hymenoptera - Peleserphidae
Arkadiserphus leleji n. gen. n. sp.43
Arkadiserphus leleji n. gen. n. sp.43 Rasnitsyn 2021 wasp
PIN 2784/1225
 Hymenoptera - Kuafuidae
Leptogastrella leptogastra n. gen. n. sp.41
Leptogastrella leptogastra n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1204
 Hymenoptera - Brachycleistogastridae
Brachycleistogaster nigripes n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1176
Brachycleistogaster jurassica n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1174, 1194
 Hymenoptera - Bethylonymidae
Bethylonymellus bethyloides n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1195
Bethylonymellus pallens n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1195
Bethylonymellus cervicalis n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1189
 Hymenoptera - Megalyridae
Leptocleistogaster alboscuta41 Rasnitsyn 1975 wasp
PIN 2784/1192
Leptocleistogaster pallida n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1175
Mesaulacinus sculpturatus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1193
 Hymenoptera - Praeaulacidae
Praeaulacon tenuis n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1182
Praeaulacus obscurus n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1191
Praeaulacus leptogaster n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1203
Praeaulacites pachygaster n. gen. n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1185, 1202, 1208
 Hymenoptera - Ephialtitidae
Stephanogaster longipalpa41 Rasnitsyn 1975 wasp
PIN 2784/1184, 1178
Stephanogaster magna n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1205
Sessiliventer oppositus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1207
Parephialtites reductus n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1177
Leptephialtites caudatus n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1206
Leptephialtites stephanocephalus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1209
Leptephialtites angustus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1199
Leptephialtites linearis n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1201
Leptephialtites picturatus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1200
Symphyogaster cylindrica n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1198
Symphytopterus robustus41 Rasnitsyn 1975 wasp
PIN 2784/1183
Symphytopterus minutus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1211
Symphytopterus macrophthalmus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1180
 Hymenoptera - Sepulcidae
Xyelula hybrida n. gen. n. sp.39
Xyelula hybrida n. gen. n. sp.39 Rasnitsyn 1969 stem sawfly
PIN 2784/1161
 Hymenoptera - Protosiricidae
Protosirex xyelopterus n. gen. n. sp.39
Protosirex xyelopterus n. gen. n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1164
 Hymenoptera -
"Mesolyda depressa n. sp." = Prolyda depressa39
"Mesolyda depressa n. sp." = Prolyda depressa39 Rasnitsyn 1969 sawfly
PIN 2784/1163
 Mecoptera - Nannochoristidae
Itaphlebia sharovi37 Novokshonov 1997 scorpionfly
PIN 2784/2223
 Mecoptera - Bittacidae
Prohylobittacus sorosi n. gen. n. sp.36
Prohylobittacus sorosi n. gen. n. sp.36 Novokshonov 1993 hangingfly
PIN 2784/2219
 Neuroptera - Osmylidae
Ensiosmylus acutus n. gen. n. sp.13
Ensiosmylus acutus n. gen. n. sp.13 Khramov 2014 lance lacewing
PIN 2784/1020
 Neuroptera - Chrysopidae
Mesypochrysa reducta n. sp.38 Panfilov 1980 green lacewing
PIN 2784/1059
 Neuroptera - Psychopsidae
Propsychops karatavicus n. gen. n. sp.38 Panfilov 1980 silky lacewing
PIN 2784/1060
Cretapsychops skywalkeri15 Chemakos and Khramov 2022 silky lacewing
PIN 2784/1021
 Neuroptera - Kalligrammatidae
Kalligrammula karatensis n. sp.24 Liu et al. 2015 lacewing
PIN 2784/1032
Meioneurites (Parameioneurites) spectabilis n. sp.5 Engel 2005 lacewing
PIN 2784/1069
 Neuroptera -
Chrysoleonites intactus n. sp.38, "Nymphoides latus n. gen. n. sp." = Osmylites latus38
Chrysoleonites intactus n. sp.38 Panfilov 1980 lacewing
PIN 2784/1048
"Nymphoides latus n. gen. n. sp." = Osmylites latus38 Panfilov 1980 lacewing
PIN 2784/1046
 Neuroptera - Mantispidae
Promantispa similis n. gen. n. sp.38, "Longicollum benmaddoxi n. gen. n. sp." = Longipronotum benmaddoxi12
Promantispa similis n. gen. n. sp.38 Panfilov 1980 mantidfly
PIN 2784/1080
"Longicollum benmaddoxi n. gen. n. sp." = Longipronotum benmaddoxi12 Jepson et al. 2018 mantidfly
PIN 2784/1056
 Neuroptera - Berothidae
Mesithone maculata n. gen. n. sp.38, Mesithone magna n. sp.38, Berothone protea14
Mesithone maculata n. gen. n. sp.38 Panfilov 1980 beaded lacewing
PIN 2784/1058
Mesithone magna n. sp.38 Panfilov 1980 beaded lacewing
PIN 2784/1025
Berothone protea14 Panfilov 1980 beaded lacewing
PIN 2784/1061
 Coleoptera - Hybosoridae
Protohybosorus karatavicus n. gen. n. sp.35
Protohybosorus karatavicus n. gen. n. sp.35 Nikolajev 2010 scavenger scarab beetle
PIN 2784/1656
 Coleoptera - Cerophytidae
"Idiomerus brevicornis n. sp." = Necromera brevicornis4, "Idiomerus latissimus n. sp." = Necromera latissima4
"Idiomerus brevicornis n. sp." = Necromera brevicornis4 Dolin 1980 beetle
PIN 2784/1374
"Idiomerus latissimus n. sp." = Necromera latissima4 Dolin 1980 beetle
PIN 2784/1380
 Coleoptera - Elateridae
Litholacon conicicollis n. sp.4 Dolin 1980 click beetle
PIN 2784/1371
Desmatus lapidarius3 Dolin 1975 click beetle
PIN 2784/1377
Plesiorhaphes scaber n. gen. n. sp.4 Dolin 1980 click beetle
PIN 2784/1383
Lithomerus cockerelli4 Dolin 1980 click beetle
PIN 2784/1366, 1389
Acheonus gracilis n. sp.4 Dolin 1980 click beetle
PIN 2784/1404
Lithosomus longicollis n. sp.4 Dolin 1980 click beetle
PIN 2784/1363
Lapidiconides innatus n. sp.4 Dolin 1980 click beetle
PIN 2784/1376(1400)
Platyelater reflexicollis4 Dolin 1980 click beetle
PIN 2784/1387
Lithoptychus incertus4 Dolin 1980 click beetle
PIN 2784/1373, 1390, 1398
Codemus sharovi4 Dolin 1980 click beetle
PIN 2784/1378
Codemus micros4 Dolin 1980 click beetle
PIN 2784/1384
Codemus alatus4 Dolin 1980 click beetle
PIN 2784/1368, 1391
Codemus carinatus4 Dolin 1980 click beetle
PIN 2784/1405, 1372, 1399
Codemus zherichini4 Dolin 1980 click beetle
PIN 2784/1365, 1367
Negastrioides globicollis4 Dolin 1980 click beetle
PIN 2784/1375
Lapidostenus longicornis n. sp.4 Dolin 1980 click beetle
PIN 2784/1393
Lapidostenus insignis n. sp.4 Dolin 1980 click beetle
PIN 2784/1379
Hypnomorphus rasnitzyni n. sp.4 Dolin 1980 click beetle
PIN 2784/1362
 Coleoptera - Anthribidae
Juranthribus thompsoni n. gen.20 Legalov 2011 fungus weevil
PIN 2784/1657
Protoscelis parvula22 Medvedev 1968 fungus weevil
PIN 2784/1434,1435,1437,1438,1442,1444,1445
Protoscelis medvedevi22 Legalov 2013 fungus weevil
PIN 2784/1433,1461
Protoscelis longicornis22 Medvedev 1968 fungus weevil
PIN 2784/1426,1428,1429,1436
Protoscelis nitidicornis22 Medvedev 1968 fungus weevil
PIN 2784/1430,1440
Protoscelis jurassica22 Medvedev 1968 fungus weevil
PIN 2784/1426,1441,1446
 Coleoptera - Caridae
Nanophydes ovatus23 Arnoldi 1977 weevil
PIN 2784/1496
 Coleoptera - Obrieniidae
Kararhynchus jurassicus n. sp.21 Legalov 2012 weevil
PIN 2784/1602
 Coleoptera - Nemonychidae
Oxycorynoides (Oxycorynoides) similis9, Probelus (Probelus) handlirschi n. sp.9, Probelus (Probelus) longitarsus9, Probelus (Probelus) nikolaevi n. sp.9, Probelus (Probelus) scudderi n. sp.9, Probelus (Probelus) curvispinus9, "Belonotaris (Arnoldibelus) rohdendorfi n. sp." = Belonotaris rohdendorfi9, Belonotaris (Arnoldibelus) karatavicus9, Belonotaris (Arnoldibelus) heeri9, "Belonotaris (Arnoldibelus) gratshevi" = Belonotaris gratshevi9, "Ampliceps furcitibia" = Ampliceps furcitibia9, "Medmetrioxenoides prodromus n. gen. n. sp." = Ampliceps furcitibia8, Archaeorrhynchus carpenteri n. sp.9, "Brenthorrhinoides pubescens n. sp." = Paleocartus pubescens10, "Pseudobrenthorrhinus tenuicornis n. sp." = Eccoptarthrus tenuicornis10, Astenorrhinus major9, Distenorrhinus (Distenorrhinus) antennatus11, "Distenorrhinus (Distenorrhinus) pallidirostris" = Distenorrhinus (Distenorrhinus) pallidirostris11, "Distenorrhinus (Distenorrhinus) rotundicollis n. sp." = Distenorrhinus (Distenorrhinus) pallidirostris11
Oxycorynoides (Oxycorynoides) similis9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1465, 1466, 1469, 1470, 1473, 1475-1479, 1481, 1485, 1490, 1492, 1494, 1497, 1498, 1500, 1504, 1507, 1509-1514, 1519-1523, 1913
Probelus (Probelus) handlirschi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1451, 1455
Probelus (Probelus) longitarsus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1505
Probelus (Probelus) nikolaevi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1449
Probelus (Probelus) scudderi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1457
Probelus (Probelus) curvispinus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1448
"Belonotaris (Arnoldibelus) rohdendorfi n. sp." = Belonotaris rohdendorfi9 Legalov 2014 pine flower snout beetle
PIN 2784/1452; PIN 2784/1450 in Legalov, 2022
Belonotaris (Arnoldibelus) karatavicus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1454, 1459, 1499
Belonotaris (Arnoldibelus) heeri9 Legalov 2014 pine flower snout beetle
PIN 2784/1518
"Belonotaris (Arnoldibelus) gratshevi" = Belonotaris gratshevi9 Legalov 2010 pine flower snout beetle
PIN 2784/1450
"Ampliceps furcitibia" = Ampliceps furcitibia9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1460, 1462, 1467, 1468, 1472, 1487, 1488, 1491, 1495, 1501, 1503, 1506, 1508, 1515, 1516, 1524
"Medmetrioxenoides prodromus n. gen. n. sp." = Ampliceps furcitibia8 Arnoldi 1977 pine flower snout beetle
PIN 2784/1486
Archaeorrhynchus carpenteri n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1453, 1456
"Brenthorrhinoides pubescens n. sp." = Paleocartus pubescens10 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1464
"Pseudobrenthorrhinus tenuicornis n. sp." = Eccoptarthrus tenuicornis10 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1493(1517)
Astenorrhinus major9 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1458
Distenorrhinus (Distenorrhinus) antennatus11 Arnoldi 1977 pine flower snout beetle
PIN 2784/1488
"Distenorrhinus (Distenorrhinus) pallidirostris" = Distenorrhinus (Distenorrhinus) pallidirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1681, 1582
"Distenorrhinus (Distenorrhinus) rotundicollis n. sp." = Distenorrhinus (Distenorrhinus) pallidirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1483(1502), 1493, 1471, 1774
 Coleoptera - Carabidae
Mesorabus elongatus n. gen. n. sp.1 Ponomarenko 1977 ground beetle
PIN 2784/1525
Cordorabus notatus1 Ponomarenko 1977 ground beetle
PIN 2784/1320
Protorabus planus1 Ponomarenko 1977 ground beetle
PIN 2784/1299
Ovrabites ovalis1 Ponomarenko 1977 ground beetle
PIN 2784/1326
 Coleoptera - Coptoclavidae
Charonoscapha ovata1 Ponomarenko 1977 beetle
PIN 2784/1925
 Coleoptera - Trachypachidae
Karatoma agilis n. gen. n. sp.1
Karatoma agilis n. gen. n. sp.1 Ponomarenko 1977 false ground beetle
PIN 2784/1528
 Blattaria - Skokidae
Skok svaba50 Vršanský 2007 jumping cockroach
PIN 2784/684
 Psocodea - Electrentomidae
Paramesopsocus adibi n. sp.2 Azar et al. 2008 book louse
PIN 2784/2096, 1998, 2001, 2340; 2339 ("Mesopsocidae in Rasnitsyn, 2002" according to Mockford et al., 2013)
 Hemiptera - Tettigarctidae
Sunotettigarcta kudryashevae n. sp.44 Shcherbakov 2008 hairy cicada
PIN 2784/2163
 Thysanoptera - Liassothripidae
Liassothrips crassipes45 Martynov 1927 thrips
PIN 2784/2207-2209, 2268, 2269
 Permopsocida - Archipsyllidae
Archipsylla similis n. sp.48 Vishnyakova 1976 winged insect
PIN 2784/2341, 2343
 Plecoptera - Baleyopterygidae
Udopteryx expansa46 Sinichenkova 1985 stonefly
PIN 2784/243
 Plecoptera - Perlariopseidae
Karanemoura abrupta47 Sinitshenkova 1987 stonefly
PIN 2784/208
 Orthoptera - Phasmomimidae
Jurophasmomima punctata n. gen. n. sp.6
Jurophasmomima punctata n. gen. n. sp.6 Gorochov 1988 grasshopper
PIN 2784/9
 Orthoptera - Protogryllidae
"Protogryllus karatavicus" = Falsispeculum karatavicus
"Protogryllus karatavicus" = Falsispeculum karatavicus Sharov 1968 grasshopper
PIN 2784/3
 Orthoptera - Prophalangopsidae
Karatailus micropterus n. gen. n. sp.7, Aboilus zebra n. sp.7, ? Aboilus amplus n. sp.7
Karatailus micropterus n. gen. n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/2275
Aboilus zebra n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/34
? Aboilus amplus n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/2360, 59
 Dermaptera - Protodiplatyidae
Protodiplatys fortis49 Martynov 1925 earwig
PIN 2784/2249
 Dermaptera - Dermapteridae
"Dermapteron incertae" = Dermapteron incerta49
"Dermapteron incertae" = Dermapteron incerta49 Martynov 1925 earwig
PIN 2784/2240
 Dermaptera - Turanoviidae
Turanovia incompleta n. gen. n. sp.49
Turanovia incompleta n. gen. n. sp.49 Vishnyakova 1980 earwig
PIN 2784/2250(2252)
 Dermaptera - Semenoviolidae
Semenovioloides capitatus n. gen. n. sp.49
Semenovioloides capitatus n. gen. n. sp.49 Vishnyakova 1980 earwig
PIN 2784/2243
 Blattodea - Blattulidae
Pseudomantina nigroalba51 Vršanský 2021 cockroach
PIN 2784/393
 Blattodea - Liberiblattinidae
Hydrokhoohydra aquabella52 Vršanský 2019 cockroach
PIN 2784/789, 982
 Diptera - Protorhyphidae
Archirhyphus asiatcus26 Rohdendorf 1964 fly
PIN 2784/211, 598
 Diptera - Archisargidae
Archisargidae indet.29, Parvisargus malus n. sp.29, "? Prosolva karataviensis n. sp." = Mesosolva karataviensis29, Calosargus (Calosargus) niger n. sp.30, ? Sharasargus spiniger n. sp.29
Archisargidae indet.29 Rohdendorf 1962 fly
PIN 2784/357
Parvisargus malus n. sp.29 Mostovski 1996 fly
PIN 2784/430
"? Prosolva karataviensis n. sp." = Mesosolva karataviensis29 Mostovski 1996 fly
PIN 2784/292
Calosargus (Calosargus) niger n. sp.30 Mostovski 1997 fly
PIN 2784/77
? Sharasargus spiniger n. sp.29 Mostovski 1996 fly
PIN 2784/344
 Diptera - Nemestrinidae
Protonemestrius rohdendorfi n. sp.31 Mostovski 1998 tangle-veined fly
PIN 2784/73
Archinemestrius karatavicus31 Rohdendorf 1968 tangle-veined fly
PIN 2784/621
Archinemestrius litigiosus n. sp.31 Mostovski 1998 tangle-veined fly
PIN 2784/79
 Diptera - Rhagionempididae
Rhagionempis antennata16 Rohdendorf 1938 fly
PIN 2784/88
 Diptera - Trichoceridae
Tanyochoreta (Trichokara) composita17 Krzemińska et al. 2009 winter crane fly
PIN 2784/280, 285
Tanyochoreta (Trichokara) minuta17 Krzemińska et al. 2009 winter crane fly
PIN 2784/475
Zherikhinina zherikhini17 Krzemińska et al. 2009 winter crane fly
PIN 2784/346
 Diptera - Ptychopteridae
Eoptychopterina angularis n. sp.25 Lukashevich 1993 phantom crane fly
PIN 2784/74
Nedoptychoptera karatavica n. gen. n. sp.27 Lukashevich 1998 phantom crane fly
PIN 2784/261
 Diptera - Ansorgiidae
"Ansorgia praedicta n. gen. n. sp." = Ansorgius praedicta18
"Ansorgia praedicta n. gen. n. sp." = Ansorgius praedicta18 Krzemiński and Lukashevich 1993 fly
PIN 2784/114, 410, 428
 Diptera - Elliidae
Polyanka minuta19 Krzemińska et al. 1993 fly
PIN 2784/388, 407, 414
 Diptera -
"? Xylomya shcherbakovi n. sp." = Xylomyia shcherbakovi32
"? Xylomya shcherbakovi n. sp." = Xylomyia shcherbakovi32 Mostovski 1999 fly
PIN 2784/131
 Diptera - Psychodidae
Praemacrochile ansorgei28 Lukashevich and Krzemiński 2009 moth fly
PIN 2784/151