Karatau-Mikhailovka (PIN collection 2784) (Jurassic to of Kazakhstan)

Where: Chimkent, Kazakhstan (42.9° N, 70.0° E: paleocoordinates 41.6° N, 73.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karabastau Formation, Callovian to Callovian (166.1 - 157.3 Ma)

Environment/lithology: lacustrine - large; lithified siltstone

• "dark shales" Grimaldi & Engel, 2005

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Reposited in the PIN

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 113439: authorized by Matthew Clapham, entered by Jered Karr on 31.07.2011, edited by Matthew Clapham, Andrew Spencer-Lee and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Diptera - Trichoceridae
Tanyochoreta (Trichokara) minuta17 Krzemińska et al. 2009 winter crane fly
PIN 2784/475
Tanyochoreta (Trichokara) composita17 Krzemińska et al. 2009 winter crane fly
PIN 2784/280, 285
Zherikhinina zherikhini17 Krzemińska et al. 2009 winter crane fly
PIN 2784/346
 Diptera - Elliidae
Polyanka minuta19 Krzemińska et al. 1993 fly
PIN 2784/388, 407, 414
 Diptera - Ptychopteridae
Eoptychopterina angularis n. sp.25 Lukashevich 1993 phantom crane fly
PIN 2784/74
Nedoptychoptera karatavica n. gen. n. sp.27 Lukashevich 1998 phantom crane fly
PIN 2784/261
 Diptera - Ansorgiidae
"Ansorgia praedicta n. gen. n. sp." = Ansorgius praedicta18
"Ansorgia praedicta n. gen. n. sp." = Ansorgius praedicta18 Krzemiński and Lukashevich 1993 fly
PIN 2784/114, 410, 428
 Diptera -
"? Xylomya shcherbakovi n. sp." = Xylomyia shcherbakovi32
"? Xylomya shcherbakovi n. sp." = Xylomyia shcherbakovi32 Mostovski 1999 fly
PIN 2784/131
 Diptera - Psychodidae
Praemacrochile ansorgei28 Lukashevich and Krzemiński 2009 moth fly
PIN 2784/151
 Diptera - Nemestrinidae
Protonemestrius rohdendorfi n. sp.31 Mostovski 1998 tangle-veined fly
PIN 2784/73
Archinemestrius litigiosus n. sp.31 Mostovski 1998 tangle-veined fly
PIN 2784/79
Archinemestrius karatavicus31 Rohdendorf 1968 tangle-veined fly
PIN 2784/621
 Diptera - Rhagionempididae
Rhagionempis antennata16 Rohdendorf 1938 fly
PIN 2784/88
 Diptera - Archisargidae
Archisargidae indet.29, Calosargus (Calosargus) niger n. sp.30, Parvisargus malus n. sp.29, ? Sharasargus spiniger n. sp.29, "? Prosolva karataviensis n. sp." = Mesosolva karataviensis29
Archisargidae indet.29 Rohdendorf 1962 fly
PIN 2784/357
Calosargus (Calosargus) niger n. sp.30 Mostovski 1997 fly
PIN 2784/77
Parvisargus malus n. sp.29 Mostovski 1996 fly
PIN 2784/430
? Sharasargus spiniger n. sp.29 Mostovski 1996 fly
PIN 2784/344
"? Prosolva karataviensis n. sp." = Mesosolva karataviensis29 Mostovski 1996 fly
PIN 2784/292
 Diptera - Protorhyphidae
Archirhyphus asiatcus26 Rohdendorf 1964 fly
PIN 2784/211, 598
 Dicondylia - Protomyrmeleontidae
Protomyrmeleon sp.34 Geinitz 1888 winged insect
PIN 2784/69
Protomyrmeleon grandis n. sp.34 Nel et al. 2005 winged insect
PIN 2784/67
Protomyrmeleon kazakhstanensis n. sp.34 Nel et al. 2005 winged insect
PIN 2784/70-71(pc)
 Odonata - Paragonophlebiidae
Paragonophlebia inexpectata n. gen. n. sp.33
Paragonophlebia inexpectata n. gen. n. sp.33 Nel 2009 dragonfly
PIN 2784/08
 Plecoptera - Perlariopseidae
Karanemoura abrupta47 Sinitshenkova 1987 stonefly
PIN 2784/208
 Plecoptera - Baleyopterygidae
Udopteryx expansa46 Sinichenkova 1985 stonefly
PIN 2784/243
 Blattodea - Liberiblattinidae
Hydrokhoohydra aquabella52 Vršanský 2019 cockroach
PIN 2784/789, 982
 Blattodea - Blattulidae
Pseudomantina nigroalba51 Vršanský 2021 cockroach
PIN 2784/393
 Dermaptera - Semenoviolidae
Semenovioloides capitatus n. gen. n. sp.49
Semenovioloides capitatus n. gen. n. sp.49 Vishnyakova 1980 earwig
PIN 2784/2243
 Dermaptera - Protodiplatyidae
Protodiplatys fortis49 Martynov 1925 earwig
PIN 2784/2249
 Dermaptera - Dermapteridae
"Dermapteron incertae" = Dermapteron incerta49
"Dermapteron incertae" = Dermapteron incerta49 Martynov 1925 earwig
PIN 2784/2240
 Dermaptera - Turanoviidae
Turanovia incompleta n. gen. n. sp.49
Turanovia incompleta n. gen. n. sp.49 Vishnyakova 1980 earwig
PIN 2784/2250(2252)
 Orthoptera - Phasmomimidae
Jurophasmomima punctata n. gen. n. sp.6
Jurophasmomima punctata n. gen. n. sp.6 Gorochov 1988 grasshopper
PIN 2784/9
 Orthoptera - Prophalangopsidae
Karatailus micropterus n. gen. n. sp.7, ? Aboilus amplus n. sp.7, Aboilus zebra n. sp.7
Karatailus micropterus n. gen. n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/2275
? Aboilus amplus n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/2360, 59
Aboilus zebra n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/34
 Orthoptera - Protogryllidae
"Protogryllus karatavicus" = Falsispeculum karatavicus
"Protogryllus karatavicus" = Falsispeculum karatavicus Sharov 1968 grasshopper
PIN 2784/3
 Hymenoptera - Sepulcidae
Xyelula hybrida n. gen. n. sp.39
Xyelula hybrida n. gen. n. sp.39 Rasnitsyn 1969 stem sawfly
PIN 2784/1161
 Hymenoptera - Karatavitidae
Karatavites medius n. sp.39 Rasnitsyn 1969 wood wasp
PIN 2784/1169
 Hymenoptera - Protosiricidae
Protosirex xyelopterus n. gen. n. sp.39
Protosirex xyelopterus n. gen. n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1164
 Hymenoptera - Megalyridae
Leptocleistogaster pallida n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1175
Leptocleistogaster alboscuta41 Rasnitsyn 1975 wasp
PIN 2784/1192
Mesaulacinus sculpturatus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1193
 Hymenoptera - Brachycleistogastridae
Brachycleistogaster jurassica n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1174, 1194
Brachycleistogaster nigripes n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1176
 Hymenoptera - Kuafuidae
Leptogastrella leptogastra n. gen. n. sp.41
Leptogastrella leptogastra n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1204
 Hymenoptera - Bethylonymidae
Bethylonymellus bethyloides n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1195
Bethylonymellus pallens n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1195
Bethylonymellus cervicalis n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1189
 Hymenoptera - Praeaulacidae
Praeaulacites pachygaster n. gen. n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1185, 1202, 1208
Praeaulacon tenuis n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1182
Praeaulacus obscurus n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1191
Praeaulacus leptogaster n. sp.40 Rasnitsyn 1972 wasp
PIN 2784/1203
 Hymenoptera - Ephialtitidae
Sessiliventer oppositus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1207
Leptephialtites angustus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1199
Leptephialtites linearis n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1201
Leptephialtites stephanocephalus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1209
Leptephialtites picturatus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1200
Leptephialtites caudatus n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1206
Parephialtites reductus n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1177
Stephanogaster magna n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1205
Stephanogaster longipalpa41 Rasnitsyn 1975 wasp
PIN 2784/1184, 1178
Symphytopterus minutus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1211
Symphytopterus robustus41 Rasnitsyn 1975 wasp
PIN 2784/1183
Symphytopterus macrophthalmus n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1180
Symphyogaster cylindrica n. gen. n. sp.41 Rasnitsyn 1975 wasp
PIN 2784/1198
 Hymenoptera - Peleserphidae
Arkadiserphus leleji n. gen. n. sp.43
Arkadiserphus leleji n. gen. n. sp.43 Rasnitsyn 2021 wasp
PIN 2784/1225
 Hymenoptera - Mesoserphidae
Campturoserphus obscurus n. gen. n. sp.42, Auliserphus niger n. sp.42
Campturoserphus obscurus n. gen. n. sp.42 Rasnitsyn 1986 wasp
PIN 2784/1236
Auliserphus niger n. sp.42 Rasnitsyn 1986 wasp
PIN 2784/1227
 Hymenoptera - Anaxyelidae
Brachysyntexis nova n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1159
Urosyntexis depressa n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1160
Sphenosyntexis pallicornis n. sp.39 Rasnitsyn 1969 winged insect
PIN 2784/1162
 Hymenoptera - Xyelidae
Sirecomima xiphophora n. gen. n. sp.39, Microxyelecia brachycera n. gen. n. sp.39
Sirecomima xiphophora n. gen. n. sp.39 Rasnitsyn 1969 sawfly
PIN 2784/1173
Microxyelecia brachycera n. gen. n. sp.39 Rasnitsyn 1969 sawfly
PIN 2784/1172
 Hymenoptera -
"Mesolyda depressa n. sp." = Prolyda depressa39
"Mesolyda depressa n. sp." = Prolyda depressa39 Rasnitsyn 1969 sawfly
PIN 2784/1163
 Neuroptera - Kalligrammatidae
Kalligrammula karatensis n. sp.24 Liu et al. 2015 lacewing
PIN 2784/1032
Meioneurites (Parameioneurites) spectabilis n. sp.5 Engel 2005 lacewing
PIN 2784/1069
 Neuroptera - Psychopsidae
Propsychops karatavicus n. gen. n. sp.38 Panfilov 1980 silky lacewing
PIN 2784/1060
Cretapsychops skywalkeri15 Chemakos and Khramov 2022 silky lacewing
PIN 2784/1021
 Neuroptera -
Chrysoleonites intactus n. sp.38, "Nymphoides latus n. gen. n. sp." = Osmylites latus38
Chrysoleonites intactus n. sp.38 Panfilov 1980 lacewing
PIN 2784/1048
"Nymphoides latus n. gen. n. sp." = Osmylites latus38 Panfilov 1980 lacewing
PIN 2784/1046
 Neuroptera - Chrysopidae
Mesypochrysa reducta n. sp.38 Panfilov 1980 green lacewing
PIN 2784/1059
 Neuroptera - Osmylidae
Ensiosmylus acutus n. gen. n. sp.13
Ensiosmylus acutus n. gen. n. sp.13 Khramov 2014 lance lacewing
PIN 2784/1020
 Neuroptera - Berothidae
Berothone protea14, Mesithone maculata n. gen. n. sp.38, Mesithone magna n. sp.38
Berothone protea14 Panfilov 1980 beaded lacewing
PIN 2784/1061
Mesithone maculata n. gen. n. sp.38 Panfilov 1980 beaded lacewing
PIN 2784/1058
Mesithone magna n. sp.38 Panfilov 1980 beaded lacewing
PIN 2784/1025
 Neuroptera - Mantispidae
"Longicollum benmaddoxi n. gen. n. sp." = Longipronotum benmaddoxi12, Promantispa similis n. gen. n. sp.38
"Longicollum benmaddoxi n. gen. n. sp." = Longipronotum benmaddoxi12 Jepson et al. 2018 mantidfly
PIN 2784/1056
Promantispa similis n. gen. n. sp.38 Panfilov 1980 mantidfly
PIN 2784/1080
 Coleoptera - Trachypachidae
Karatoma agilis n. gen. n. sp.1
Karatoma agilis n. gen. n. sp.1 Ponomarenko 1977 false ground beetle
PIN 2784/1528
 Coleoptera - Carabidae
Mesorabus elongatus n. gen. n. sp.1 Ponomarenko 1977 ground beetle
PIN 2784/1525
Cordorabus notatus1 Ponomarenko 1977 ground beetle
PIN 2784/1320
Protorabus planus1 Ponomarenko 1977 ground beetle
PIN 2784/1299
Ovrabites ovalis1 Ponomarenko 1977 ground beetle
PIN 2784/1326
 Coleoptera - Coptoclavidae
Charonoscapha ovata1 Ponomarenko 1977 beetle
PIN 2784/1925
 Coleoptera - Elateridae
Litholacon conicicollis n. sp.4 Dolin 1980 click beetle
PIN 2784/1371
Desmatus lapidarius3 Dolin 1975 click beetle
PIN 2784/1377
Plesiorhaphes scaber n. gen. n. sp.4 Dolin 1980 click beetle
PIN 2784/1383
Lithomerus cockerelli4 Dolin 1980 click beetle
PIN 2784/1366, 1389
Acheonus gracilis n. sp.4 Dolin 1980 click beetle
PIN 2784/1404
Lithoptychus incertus4 Dolin 1980 click beetle
PIN 2784/1373, 1390, 1398
Lapidostenus longicornis n. sp.4 Dolin 1980 click beetle
PIN 2784/1393
Lapidostenus insignis n. sp.4 Dolin 1980 click beetle
PIN 2784/1379
Negastrioides globicollis4 Dolin 1980 click beetle
PIN 2784/1375
Codemus alatus4 Dolin 1980 click beetle
PIN 2784/1368, 1391
Codemus carinatus4 Dolin 1980 click beetle
PIN 2784/1405, 1372, 1399
Codemus zherichini4 Dolin 1980 click beetle
PIN 2784/1365, 1367
Codemus micros4 Dolin 1980 click beetle
PIN 2784/1384
Codemus sharovi4 Dolin 1980 click beetle
PIN 2784/1378
Hypnomorphus rasnitzyni n. sp.4 Dolin 1980 click beetle
PIN 2784/1362
Lithosomus longicollis n. sp.4 Dolin 1980 click beetle
PIN 2784/1363
Platyelater reflexicollis4 Dolin 1980 click beetle
PIN 2784/1387
Lapidiconides innatus n. sp.4 Dolin 1980 click beetle
PIN 2784/1376(1400)
 Coleoptera - Cerophytidae
"Idiomerus latissimus n. sp." = Necromera latissima4, "Idiomerus brevicornis n. sp." = Necromera brevicornis4
"Idiomerus latissimus n. sp." = Necromera latissima4 Dolin 1980 beetle
PIN 2784/1380
"Idiomerus brevicornis n. sp." = Necromera brevicornis4 Dolin 1980 beetle
PIN 2784/1374
 Coleoptera - Hybosoridae
Protohybosorus karatavicus n. gen. n. sp.35
Protohybosorus karatavicus n. gen. n. sp.35 Nikolajev 2010 scavenger scarab beetle
PIN 2784/1656
 Coleoptera - Caridae
Nanophydes ovatus23 Arnoldi 1977 weevil
PIN 2784/1496
 Coleoptera - Obrieniidae
Kararhynchus jurassicus n. sp.21 Legalov 2012 weevil
PIN 2784/1602
 Coleoptera - Anthribidae
Juranthribus thompsoni n. gen.20 Legalov 2011 fungus weevil
PIN 2784/1657
Protoscelis jurassica22 Medvedev 1968 fungus weevil
PIN 2784/1426,1441,1446
Protoscelis longicornis22 Medvedev 1968 fungus weevil
PIN 2784/1426,1428,1429,1436
Protoscelis parvula22 Medvedev 1968 fungus weevil
PIN 2784/1434,1435,1437,1438,1442,1444,1445
Protoscelis nitidicornis22 Medvedev 1968 fungus weevil
PIN 2784/1430,1440
Protoscelis medvedevi22 Legalov 2013 fungus weevil
PIN 2784/1433,1461
 Coleoptera - Nemonychidae
"Distenorrhinus (Distenorrhinus) pallidirostris" = Distenorrhinus (Distenorrhinus) pallidirostris11, "Distenorrhinus (Distenorrhinus) rotundicollis n. sp." = Distenorrhinus (Distenorrhinus) pallidirostris11, Distenorrhinus (Distenorrhinus) antennatus11, "Pseudobrenthorrhinus tenuicornis n. sp." = Eccoptarthrus tenuicornis10, Astenorrhinus major9, "Ampliceps furcitibia" = Ampliceps furcitibia9, "Medmetrioxenoides prodromus n. gen. n. sp." = Ampliceps furcitibia8, Oxycorynoides (Oxycorynoides) similis9, Probelus (Probelus) longitarsus9, Probelus (Probelus) curvispinus9, Probelus (Probelus) handlirschi n. sp.9, Probelus (Probelus) nikolaevi n. sp.9, Probelus (Probelus) scudderi n. sp.9, "Belonotaris (Arnoldibelus) rohdendorfi n. sp." = Belonotaris rohdendorfi9, Belonotaris (Arnoldibelus) heeri9, Belonotaris (Arnoldibelus) karatavicus9, "Belonotaris (Arnoldibelus) gratshevi" = Belonotaris gratshevi9, Archaeorrhynchus carpenteri n. sp.9, "Brenthorrhinoides pubescens n. sp." = Paleocartus pubescens10
"Distenorrhinus (Distenorrhinus) pallidirostris" = Distenorrhinus (Distenorrhinus) pallidirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1681, 1582
"Distenorrhinus (Distenorrhinus) rotundicollis n. sp." = Distenorrhinus (Distenorrhinus) pallidirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1483(1502), 1493, 1471, 1774
Distenorrhinus (Distenorrhinus) antennatus11 Arnoldi 1977 pine flower snout beetle
PIN 2784/1488
"Pseudobrenthorrhinus tenuicornis n. sp." = Eccoptarthrus tenuicornis10 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1493(1517)
Astenorrhinus major9 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1458
"Ampliceps furcitibia" = Ampliceps furcitibia9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1460, 1462, 1467, 1468, 1472, 1487, 1488, 1491, 1495, 1501, 1503, 1506, 1508, 1515, 1516, 1524
"Medmetrioxenoides prodromus n. gen. n. sp." = Ampliceps furcitibia8 Arnoldi 1977 pine flower snout beetle
PIN 2784/1486
Oxycorynoides (Oxycorynoides) similis9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1465, 1466, 1469, 1470, 1473, 1475-1479, 1481, 1485, 1490, 1492, 1494, 1497, 1498, 1500, 1504, 1507, 1509-1514, 1519-1523, 1913
Probelus (Probelus) longitarsus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1505
Probelus (Probelus) curvispinus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1448
Probelus (Probelus) handlirschi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1451, 1455
Probelus (Probelus) nikolaevi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1449
Probelus (Probelus) scudderi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1457
"Belonotaris (Arnoldibelus) rohdendorfi n. sp." = Belonotaris rohdendorfi9 Legalov 2014 pine flower snout beetle
PIN 2784/1452; PIN 2784/1450 in Legalov, 2022
Belonotaris (Arnoldibelus) heeri9 Legalov 2014 pine flower snout beetle
PIN 2784/1518
Belonotaris (Arnoldibelus) karatavicus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1454, 1459, 1499
"Belonotaris (Arnoldibelus) gratshevi" = Belonotaris gratshevi9 Legalov 2010 pine flower snout beetle
PIN 2784/1450
Archaeorrhynchus carpenteri n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1453, 1456
"Brenthorrhinoides pubescens n. sp." = Paleocartus pubescens10 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1464
 Mecoptera - Bittacidae
Prohylobittacus sorosi n. gen. n. sp.36
Prohylobittacus sorosi n. gen. n. sp.36 Novokshonov 1993 hangingfly
PIN 2784/2219
 Mecoptera - Nannochoristidae
Itaphlebia sharovi37 Novokshonov 1997 scorpionfly
PIN 2784/2223
 Psocodea - Electrentomidae
Paramesopsocus adibi n. sp.2 Azar et al. 2008 book louse
PIN 2784/2096, 1998, 2001, 2340; 2339 ("Mesopsocidae in Rasnitsyn, 2002" according to Mockford et al., 2013)
 Permopsocida - Archipsyllidae
Archipsylla similis n. sp.48 Vishnyakova 1976 winged insect
PIN 2784/2341, 2343
 Hemiptera - Tettigarctidae
Sunotettigarcta kudryashevae n. sp.44 Shcherbakov 2008 hairy cicada
PIN 2784/2163
 Thysanoptera - Liassothripidae
Liassothrips crassipes45 Martynov 1927 thrips
PIN 2784/2207-2209, 2268, 2269
 Blattaria - Skokidae
Skok svaba50 Vršanský 2007 jumping cockroach
PIN 2784/684