Dominican amber (USNM Brodzinsky Lopez-Pena coll) (Miocene to of Dominican Republic)

Where: Dominican Republic (19.6° N, 70.8° W: paleocoordinates 19.4° N, 69.6° W)

• coordinate based on political unit

• outcrop-level geographic resolution

When: Burdigalian to Burdigalian (20.4 - 13.8 Ma)

• Age controversial, with the youngest proposed age of 20-15 mya based on foraminifera and the oldest as 45-30 mya based on coccoliths. Most of the amber is secondarily deposited in turbiditic sandstones.

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; amber

Size class: macrofossils

Preservation: soft parts, original chitin, amber

Reposited in the USNM

Primary reference: S. R. Davis and M. S. Engel. 2007. A new ambrosia beetle in Miocene amber of the Dominican Republic (Coleoptera: Curculionidae: Platypodinae). Alavesia 1:121-124 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 122473: authorized by Matthew Clapham, entered by Matthew Clapham on 29.12.2011, edited by Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Myriapoda
 Polydesmida - Pyrgodesmidae
Docodesmus brodzinskyi n. sp.32 Shear 1981 millipede
In J. Brodzinsky collection, Santo Domingo
 Siphonophorida - Siphonophoridae
Siphonocybe sp.32 Pocock 1903 millipede
In J. Brodzinsky collection, Santo Domingo
Arachnida
 Opiliones - Samoidae
Pellobunus proavus n. sp.5 Cokendolpher 1987 harvestman
USNM 5230
Insecta
 Diptera - Scatopsidae
"Rhegmoclemina (Neorhegmoclemina) sp." = Neorhegmoclemina6
"Rhegmoclemina (Neorhegmoclemina) sp." = Neorhegmoclemina6 Cook 1955 minute black scavenger fly
 Diptera - Tipulidae
Brachypremna brodzinskii n. sp.25 Krzemiński 1996 crane fly
USNM 8272
 Diptera - Limoniidae
Helius (Helius) oosterbroeki n. sp.24 Kopeć et al. 2016 crane fly
USNM Brodzinsky/Lopez-Penha 10189
Helius (Helius) neali n. sp.24 Kopeć et al. 2016 crane fly
USNM Brodzinsky/Lopez-Penha 10938
Elephantomyia (Elephantomyia) grata24 Podenas and Poinar 2001 crane fly
USNM Brodzinsky/Lopez-Penha 5546
 Diptera - Ceratopogonidae
Nannohelea "sp. A"38 Grogan and Wirth 1980 biting midge
USNM 9256
Baeodasymyia dominicana35 Szadziewski and Grogan 1994 biting midge
USNM 5540
Brachypogon (Isohelea) sp.38 Kieffer 1917 biting midge
USNM 2580, 2965, 3751, 5346, 5556, 5611, 7359, 7424, 685, 7691, 7781, 8618, 8666, 8681, 9301, 9501, 9514, 9712, 9735, 9862, 9947, 9958, 10083, 10180, 10373,10529, 10728, 10730, 10771, 10888
Brachypogon (Isohelea) prominuloides n. sp.38 Szadziewski and Grogan 1998 biting midge
USNM 9568
Brachypogon (Isohelea) dominicanus38 Szadziewski and Grogan 1998 biting midge
USNM 2693, 7642, 10729, 10732, 9137a, 9848, 10731, 10734
Stilobezzia (Stilobezzia) antilleana n. sp.38 Szadziewski and Grogan 1998 biting midge
USNM 9501a, USNM 10921,
Stilobezzia (Acanthohelea) "sp. A"38 Kieffer 1917 biting midge
USNM 11320
Stilobezzia (Acanthohelea) dominicana38 Szadziewski and Grogan 1998 biting midge
USNM 2932, USNM 7615b, USNM 8657, USNM 9425a, USNM 3099a
"Culicoides mammalicolus n. sp." = Culicoides mammalicola38 Szadziewski and Grogan 1998 biting midge
USNM 3742
"Culicoides (Oecacta) ambericus n. sp." = Culicoides ambericus38 Szadziewski and Grogan 1998 biting midge
USNM 2905, USNM 8968a
"Culicoides (Oecacta) antilleanus n. sp." = Culicoides antilleanus38 Szadziewski and Grogan 1998 biting midge
USNM 9137.b
Culicoides (Oecacta) sp.38 Poey 1853 biting midge
USNM 2771a, 2852, 2884, 2903, 5454a, 7639, 8328, 8707, 10900a
Palpomyia wirthorum n. sp.36 Szadziewski and Grogan 1997 biting midge
USNM 10073 b
Dasyhelea sp.37 Kieffer 1913 biting midge
USNM 2772, 2870, 3076, 3098, 5559, 5611e,c, 7514, 736, 7832, 8271, 9074, 9262, 9490, 9552, 9731, 9758, 10089, 10206, 10378, 10702, 10742, 10771, 10855, 10872, 10907
"Dasyhelea (Sebessia) dominicana" = Dasyhelea dominicana37 Szadziewski and Grogan 1998 biting midge
USNM 10910, 2771b
"Dasyhelea (Sebessia) antilleana n. sp." = Dasyhelea antilleana37 Szadziewski and Grogan 1998 biting midge
USNM 8178
Forcipomyia sp.37 Meigen 1818 biting midge
Forcipomyia (Lasiohelea) sp.37 Kieffer 1921 biting midge
USNM 3061, 7611, 8229, 8484, 9679, 9712, 10774, 10795, 10894
Forcipomyia (Lasiohelea) americana n. sp.37 Szadziewski and Grogan 1998 biting midge
USNM 7905
Forcipomyia (Lasiohelea) woodruffi37 Szadziewski and Grogan 1998 biting midge
USNM 8460,10881
Forcipomyia (Synthyridomyia) tertiaricola n. sp.37 Szadziewski and Grogan 1998 biting midge
USNM 10039
Forcipomyia (Lepidohelea) sp.37 Kieffer 1917 biting midge
USNM 2652b, 2826, 3099b, 3709, 7609, 7768, 7871, 9413b, 10651
Forcipomyia (Lepidohelea) domibicolor37 Szadziewski and Grogan 1998 biting midge
USNM 3099b, 10651, 10868, 10876, 10908
Forcipomyia (Lepidohelea) antilleana n. sp.37 Szadziewski and Grogan 1998 biting midge
USNM 10915, 9202, 10866, 10894, 10897, 2826, 7609
Forcipomyia (Lepidohelea) lepidosuccinea n. sp.37 Szadziewski and Grogan 1998 biting midge
USNM 2932b
Forcipomyia (Trichohelea) sp.37 Goetghebuer 1920 biting midge
USNM 8968b
Forcipomyia (Thyridomyia) sp.37 Saunders 1925 biting midge
USNM 2678, 10015
Forcipomyia (Thyridomyia) frutetosuccinea n. sp.37 Szadziewski and Grogan 1998 biting midge
USNM 10549
Forcipomyia (Forcipomyia) grimaldii37 Szadziewski and Grogan 1998 biting midge
USNM 2799, 7437, 10879, 10892
Atrichopogon (Atrichopogon) sp.37 Kieffer 1906 biting midge
USNM 2652a
"Atrichopogon (Atrichopogon) dominicanus n. sp." = Atrichopogon dominicanus37 Szadziewski and Grogan 1998 biting midge
USNM 7875
 Diptera - Tabanidae
Stenotabanus woodruffi n. sp.12 Fairchild 1989 horse fly
10543
 Diptera - Phoridae
Apocephalus succineus3 Brown 2000 scuttle fly
USNM-IP 3804
 Diptera - Drosophilidae
Drosophilinae "indet. A"20, "Drosophila (Hirtodrosophila) paleothoracis" = Hirtodrosophila paleothoracis20, Drosophila (Drosophila) succini n. sp.20
Drosophilinae "indet. A"20 Rondani 1856 fruit fly
USNM 8512
"Drosophila (Hirtodrosophila) paleothoracis" = Hirtodrosophila paleothoracis20 Grimaldi 1987 fruit fly
USNM 10664
Drosophila (Drosophila) succini n. sp.20 Grimaldi 1987 fruit fly
USNM 8512
 Diptera - Tanypezidae
Neotanypeza dominicana n. sp.26 Lonsdale and Apigian 2010 fly
USNM 7457
 Diptera - Tephritidae
Problepharoneura antiqua n. gen. n. sp.27
Problepharoneura antiqua n. gen. n. sp.27 Norrbom and Condon 1999 fruit fly
USNM ENT 00028651
 Diptera - Milichiidae
Milichiella hennigi n. sp.2 Brake 2006 fly
USNM Ent 00028676
 Diptera - Bombyliidae
Anthrax succini n. sp.19
Anthrax succini n. sp.19 Greenwalt and Evenhuis 2015 bee fly
USNM 508762
 Diptera - Scenopinidae
Metatrichia pria46 Yeates and Grimaldi 1993 window fly
USNM 12495
 Hemiptera - Miridae
Diphleps yenli30 Santiago-Blay and Poinar 1993 plant bug
USNM 12535
 Hemiptera - Aradidae
Calisiopsis brodzinskyorum n. sp.15 Froeschner 1992 flat bug
USNM Brodzinsky/Lopez coll.
 Hemiptera - Aetalionidae
Aetalionini indet.10 Spinola 1850 treehopper
 Hemiptera - Cicadellidae
Cicadellini indet.11 Latreille 1825 leafhopper
BLPC 7629
Jassoqualus hispaniolensis n. sp.11 Dietrich and Vega 1995 leafhopper
BPLC #5618
"Krisna garciamarquezi n. sp." = Archiokrisna garciamarquezi11 Dietrich and Vega 1995 leafhopper
BPLC #8649
Xestocephalus sp.11 Baker 1915 leafhopper
BLPC 2512, 2675-2677, 2679, 2681, 2682, 2684-2686, 2690-2692, 2861, 5078, 5533, 5567, 5603, 5626, 7076, 7086, 7096, 7402, 7419, 7479, 7630, 7632, 7633, 7635, 7638, 7640, 7641, 7649, 7651, 7657, 7658
Dikraneurini indet.11 McAtee 1926 leafhopper
BLPC 7653
? Agallia sp.11 Curtis 1833 leafhopper
BLPC 8653
 Coleoptera - Lucanidae
Syndesus ambericus n. sp.45 Woodruff 2009 stag beetle
USNM #502873
 Coleoptera - Lycidae
Cessator brodzinskyi n. sp.13 Ferreira and Ivie 2017 net-winged beetle
 Coleoptera - Curculionidae
Mesocordylus longiscapus n. sp.9 Davis and Engel 2009 snout beetle
USNM 506758
Dryophthorus acarophilus n. sp.7 Davis and Engel 2006 snout beetle
USNM 502865
Stenommatus pulvereus n. sp.7 Davis and Engel 2006 snout beetle
USNM 502681, 505478
Diaprepes anticus n. sp.14 Franz and Zhang 2017 snout beetle
USNM 505325
Tropirhinus palpebratus n. sp.14 Franz and Zhang 2017 snout beetle
USNM 505319
Proeces longirostrum n. sp.8 Davis and Engel 2007 snout beetle
USNM 502728 (Woodruff 5501)
Caulophilus ashei8 Davis and Engel 2006 snout beetle
USNM 506622 (Woodruff 11495)
Caulophilus bennetti n. sp.8 Davis and Engel 2007 snout beetle
USNM 505352 (Woodruff 9801)
Caulophilus falini n. sp.8 Davis and Engel 2007 snout beetle
USNM 505815 (Woodruff 10265)
Caulophilus swensoni n. sp.8 Davis and Engel 2007 snout beetle
USNM 502877 (Woodruff 5660), 502666 (Woodruff 5439)
Paralicus abnormis n. sp.8 Davis and Engel 2007 snout beetle
USNM 505350 (Woodruff 9799)
Dryotribus amplioculus n. sp.8 Davis and Engel 2007 snout beetle
USNM 505329 (Woodruff 9778)
Micromimus orcus n. sp.8 Davis and Engel 2007 snout beetle
USNM 505348 (Woodruff 9797), 505324 (Woodruff 9773)
Stenotrupis breviscapus n. sp.8 Davis and Engel 2007 snout beetle
USNM 505344 (Woodruff 9793), 504857 (Woodruff 9304), 505354 (Woodruff 9803)
Cossonus hinojosai n. sp.8 Davis and Engel 2007 snout beetle
USNM 505580 (Woodruff 10029)
Cenocephalus quasiexquisitus n. sp. Davis and Engel 2007 ambrosia beetle
USNM 505815
 Coleoptera - Chrysomelidae
Cryptocephalus kheelorum n. sp.4 Chamorro and Bukejs 2015 leaf beetle
Delocrania sp.31 Guerin-Meneville 1844 tortoise beetle
"precise origin unavailable", "may be deposited at NMNH"
 Coleoptera - Tenebrionidae
Nesocyrtosoma minisculum n. sp.23 Hopp and Ivie 2009 darkling beetle
 Lepidoptera - Riodinidae
Voltinia dramba n. sp.22
Voltinia dramba n. sp.22 Hall et al. 2004 butterfly
 Trichoptera - Pseudoneureclipsidae
Antillopsyche oliveri44 Wichard 1985 caddisfly
 Trichoptera - Hydroptilidae
Ochrotrichia aliceae n. sp.43 Wichard 2000 microcaddisfly
USNM 9981 ENT-I-17891
Ochrotrichia (Ochrotrichia) chaulioda n. sp.40 Wells and Wichard 1989 microcaddisfly
Ochrotrichia (Ochrotrichia) denaia n. sp.40 Wells and Wichard 1989 microcaddisfly
Leucotrichia adela n. sp.40 Wells and Wichard 1989 microcaddisfly
 Trichoptera - Glossosomatidae
Cubanoptila mederi n. sp.41 Wichard 1989 caddisfly
 Trichoptera - Leptoceridae
Setodes resinacapta n. sp.42 Wichard 1995 long-horned caddisfly
USNM 5023 ENT-I-7049
Setodes aureoinclusa n. sp.42 Wichard 1995 long-horned caddisfly
USNM 9096 ENT-I-10680
 Hymenoptera - Scolebythidae
"Dominibythus inopinatus n. gen. n. sp." = Pristapenesia inopinata29
"Dominibythus inopinatus n. gen. n. sp." = Pristapenesia inopinata29 Prentice and Poinar 1996 wasp
USNM 7594
 Hymenoptera - Crabronidae
Trypoxylon (Trypoxylon) electrum n. sp.1 Antropov 1995 apoid wasp
USNM 8429
 Hymenoptera -
Scelioninae indet.39 Thomson 1858 wasp
USNM 504043, 506135
Macroteleia sp.39 Westwood 1835 wasp
USNM 505551
Duta sp.39 Nixon 1933 wasp
USNM 503993
Spiniteleia sp.39 Masner 1980 wasp
USNM 504039
Gryon sp.39 Haliday 1833 wasp
USNM 501977, 502177, 503169, 503174, 503967, 503972, 504024, 504108, 504092, 505932(x2)
Paridris yumai n. sp.39 Talamas 2015 wasp
USNM 503175 (ENT00977548), 504379, 504846, 505333, 506231
Odontacolus sp.39 Kieffer 1910 wasp
USNM 502761, 503341, 504084, 504087
Idris sp.39 Foerster 1856 wasp
Triteleia sp.39 Kieffer 1906 wasp
USNM 502381, 503983, 503988
Embidobia sp.39 Ashmead 1896 wasp
USNM 501905, 504068
Leptoteleia sp.39 Kieffer 1908 wasp
USNM 504061
Opisthacantha sp.39 Ashmead 1893 wasp
USNM 501975, 503753, 504090
Calotelea sp.39 Westwood 1837 wasp
Psilanteris sp.39 Kieffer 1916 wasp
USNM 505541
Telenomus sp.39 Haliday 1833 wasp
USNM 502995(x4), 503179, 503344, 503436, 504018, 504359(x2), 504521, 505745
Calliscelio sp.39 Ashmead 1893 wasp
USNM 504070, 504078, 504092, 504432
Aradophagus sp.39 Ashmead 1893 wasp
USNM 504808
Probaryconus sp.39 Kieffer 1908 wasp
Embioctonus sp.39 Masner 1980 wasp
USNM 504343
 Hymenoptera - Platygastridae
Parabaeus sp.39 Kieffer 1910 wasp
USNM 504900
Fidiobia sp.39 Ashmead 1894 wasp
Platygastrinae indet.39 Haliday 1833 wasp
USNM 502389, 503261, 504094, 504104, 504689, 505436
Allostemma sp.39 Masner and Huggert 1989 wasp
USNM 502077, 502310, 504115
Leptacis sp.39 Förster 1856 wasp
USNM 502309, 503693, 505564
Inostemma sp.39 Haliday 1833 wasp
USNM 505077
Platygaster sp.39 Latreille 1809 wasp
USNM 501972, 503056, 503901, 503969, 504268, 504359, 504537, 505070, 505208, 505406, 505995
 Hymenoptera - Geoscelionidae
Plaumannion fistulosum n. sp.39 Talamas 2015 wasp
USNM 502963 (ENT00903996)
 Hymenoptera - Argidae
Didymia davisi n. sp.33
Didymia davisi n. sp.33 Smith 1992 argid sawfly
NMNH 10500
 Blattodea -
Erucoblatta semicaeca n. gen. n. sp.16
Erucoblatta semicaeca n. gen. n. sp.16 Gorochov and Anisyutkin 2007 cockroach
USNM 503476
 Blattodea - Corydiidae
Holocompsa nigra n. sp.16 Gorochov and Anisyutkin 2007 cockroach
USNM 502411
Holocompsa abbreviata n. sp.16 Gorochov and Anisyutkin 2007 cockroach
USNM 504367
 Blattodea - Pseudophyllodromiidae
Euthlastoblatta sp.21 Hebard 1917 cockroach
Plectoptera sp.21 Saussure 1864 cockroach
Plectoptera electrina n. sp.16 Gorochov and Anisyutkin 2007 cockroach
USNM 504367
 Plecoptera - Perlidae
Dominiperla antigua n. gen. n. sp.34
Dominiperla antigua n. gen. n. sp.34 Stark and Lentz 1992 common stonefly
BPLC 5658
 Zoraptera - Zorotypidae
Zorotypus palaeus n. sp.28 Poinar 1988 winged insect
BPLC 7406
 Orthoptera - Mogoplistidae
Pseudarachnocephalus latiusculus n. sp.17 Gorochov 2010 scaly cricket
NMNH 502773 (Brodzinsky/Lopez-Pena coll)
Pseudarachnocephalus dominicanus n. gen. n. sp.17 Gorochov 2010 scaly cricket
NMNH 504396, 504366, 504387 (Brodzinsky/Lopez-Pena coll)
 Orthoptera - Gryllidae
Proanaxipha ? latoca17 Vickery and Poinar 1994 true crickets
NMNH 502986, 503253, 503257, 504373, 504397, 504436, 504456
? Grossoxipha feminea n. sp.17 Gorochov 2010 true crickets
NMNH 502292, 504395, 505727 (Brodzinsky/Lopez-Pena coll)
 Orthoptera - Trigonidiidae
Abanaxipha longispina17 Vickery and Poinar 1994 grasshopper
NMNH 502562, 504374 (Brodzinsky/Lopez-Pena coll)
? Cyrtoxipha illegibilis n. sp.17 Gorochov 2010 grasshopper
NMNH 506712 (Brodzinsky/Lopez-Pena coll)
? Cyrtoxipha electrina n. sp.17 Gorochov 2010 grasshopper
NMNH 504374 (Brodzinsky/Lopez-Pena coll)
 Orthoptera - Tettigoniidae
Miophlugis rostratus n. gen. n. sp.18
Miophlugis rostratus n. gen. n. sp.18 Gorochov 2010 katydid
NMNH 504426 (Brodzinsky/Lopez-Pena coll)
 Orthoptera - Tridactylidae
Archaeoellipes ? engeli18 Heads 2010 pygmy mole cricket
NMNH 503419 (Brodzinsky/Lopez-Pena coll)