Hakel (Lewis collection) (Cretaceous of Lebanon)

Also known as Haqel, Haqil

Where: Lebanon (34.2° N, 35.8° E: paleocoordinates 13.0° N, 30.4° E)

• coordinate estimated from map

• local area-level geographic resolution

When: Metoicoceras geslinianum ammonoid zone, Late/Upper Cenomanian (99.6 - 93.5 Ma)

• The precise age of Hâqel and Hjoûla varies according to different authors. ln his comprehensive sedimentological study Hückel (1970) divided the Cenomanian Limestones into seven units and concluded that Hâqel (unite V) and Hjoûla (unite IV) were deposited during the later Lower Cenomanian. Owing to the occurrence of the ammonite Allocrioceras annulatum as a member of the Metoicoceras geslinianum Zone, Wippich & Lehmann (2004) confirmed the conclusions of Hemleben (1977) and dated Hâqel and Hjoûla as Upper Cenomanian (Fig. 2). Dalla Vecchia et al. (2002) considered Hâqel and Hjoûla as lower Upper Cenomanian, because Nammoûra (another fossil-rich locality a few kilometres in the South of Hjoûla and slightly younger than Hâqel and Hjoûla) is dated as Middle Cenomanian.

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; lithified carbonate

Size class: macrofossils

Preservation: soft parts

Collected by E.R. Lewis; reposited in the BMNH

Collection methods: quarrying

• Specimens initially in collection of Prof. E.R. Lewis, Syrian Protestant College, Lebanon. Later transferred to the museum of Mr. Robert Damon at Weymouth, and many specimens later acquired by BMNH.

Primary reference: H. Woodward. 1879. Contributions to the knowledge of fossil Crustacea. Quarterly Journal of the Geological Society London 35:549-556 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 216596: authorized by Matthew Clapham, entered by Matthew Clapham on 20.12.2020, edited by Josh Mordaunt

Creative Commons license: CC BY (attribution)

Taxonomic list

Osteichthyes
  -
Pseudoberyx grandis n. sp.1 Davis 1887
Actinopteri
 Pycnodontiformes - Pycnodontidae
Palaeobalistum libanicum2 Gorjanović-Kramberger 1895
Coccodus armatus1 Pictet 1850
 Ichthyodectiformes - Ichthyodectidae
"Chirocentrites libanicus" = Ichthyodectes libanicus1
"Chirocentrites libanicus" = Ichthyodectes libanicus1 Pictet and Humbert 1866
 Albuliformes - Albulidae
"Clupea lewisii n. sp." = Lebonichthys lewisi1
"Clupea lewisii n. sp." = Lebonichthys lewisi1 Davis 1887 bonefish
 Acanthomorphata - Aipichthyidae
Aipichthys velifer3, "Aipichthys minor" = Aipichthys minor3, "Platax brevis n. sp." = Aipichthys minor1
Aipichthys velifer3 Woodward 1901
"Aipichthys minor" = Aipichthys minor3 Pictet 1850
"Platax brevis n. sp." = Aipichthys minor1 Pictet 1850
 Beryciformes - Hoplopterygiidae
"Pycnosterinx lewisii n. sp." = Hoplopteryx lewisi1
"Pycnosterinx lewisii n. sp." = Hoplopteryx lewisi1 Davis 1887
 Aulopiformes - Aulopidae
"Pseudoberyx longispina n. sp." = Nematonotus longispinus1
"Pseudoberyx longispina n. sp." = Nematonotus longispinus1 Davis 1887
 Aulopiformes - Apateopholidae
"Rhinellus laniatus n. sp." = Apateopholis laniatus1, "Rhinellus longirostris n. sp." = Apateopholis laniatus1
"Rhinellus laniatus n. sp." = Apateopholis laniatus1 Davis 1887
"Rhinellus longirostris n. sp." = Apateopholis laniatus1 Davis 1887
 Aulopiformes - Ichthyotringidae
Rhinellus curtirostris n. sp.1 Davis 1887
 Aulopiformes - Halecidae
Phylactocephalus microlepis n. gen. n. sp.1 Davis 1887
 Ctenothrissiformes - Ctenothrissidae
"Beryx ovalis n. sp." = Ctenothrissa vexillifer1
"Beryx ovalis n. sp." = Ctenothrissa vexillifer1 Pictet 1850
 Gonorhynchiformes - Gonorhynchidae
"Spaniodon hakelensis n. sp." = Charitosomus hakelensis1
"Spaniodon hakelensis n. sp." = Charitosomus hakelensis1 Davis 1887
Osteichthyes
  -
Exocoetoides minor n. gen. n. sp.1, "? Engraulis tenuis n. sp." = Exocoetoides tenuis1
Exocoetoides minor n. gen. n. sp.1 Davis 1887
"? Engraulis tenuis n. sp." = Exocoetoides tenuis1 Davis 1887
 Halecomorphi - Ionoscopidae
"Spathiurus dorsalis n. gen. n. sp." = Spathiurus dorsalis1, "Amphilaphurus major n. gen. n. sp." = Spathiurus dorsalis1
"Spathiurus dorsalis n. gen. n. sp." = Spathiurus dorsalis1 Davis 1887 ray-finned fish
"Amphilaphurus major n. gen. n. sp." = Spathiurus dorsalis1 Davis 1887 ray-finned fish
 Actinopterygii -
"Petalopteryx dorsalis n. sp." = Aphanepygus dorsalis1
"Petalopteryx dorsalis n. sp." = Aphanepygus dorsalis1 Davis 1887 ray-finned fish
Chondrichthyes
 Rajiformes - Rajidae
"Rhinobatus expansus n. sp." = Pararaja expansa1
"Rhinobatus expansus n. sp." = Pararaja expansa1 Davis 1887 skate
 Rajiformes - Rhinobatidae
"Rhinobatus grandis n. sp." = Rhinobatos maronita1
"Rhinobatus grandis n. sp." = Rhinobatos maronita1 Pictet and Humbert 1866 guitarfish
Malacostraca
 Stomatopoda - Squillidae
Squilla lewesii n. sp. Woodward 1879 mantis shrimp
Xiphosura
 Xiphosurida - Limulidae
"Limulus syriacus n. sp." = Tachypleus syriacus
"Limulus syriacus n. sp." = Tachypleus syriacus Woodward 1879 horseshoe crab