PBDB Collection

Laño L1A (Cretaceous of Spain)

Also known as Laño Quarry

Where: Castilla y Leon, Spain (42.7° N, 2.6° W: paleocoordinates 33.0° N, 0.1° W)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Vitoria Formation, Late/Upper Campanian (83.5 - 70.6 Ma)

• in a sector of the south limb of the Miranda-Trevino syncline, level is also called L1A

•lower pat of subchron C32n

•Also assigned to Sedano Fm. or its equivalent

• bed-level stratigraphic resolution

Environment/lithology: coastal; hardground, conglomeratic sandstone and black siltstone

• upper foreshore and backshore environments to a subcontinental, plant-rich perhaps swampy environment. The vertebrate bearing levels indicate a continental depositional environment affected by high-energy episodes, a fluviatile environment.

• Multidirectional cross-laminated sands with microconglomerates in the bottom-sets, rarely more than 1m thick. Strata become massive toward the top, and are frequently intersected by abrasion surfaces showing an iron-mineralized crust or hard-ground. The most important bone accumulation is between two of these hardgrounds. The unit is topped by two black silt beds.

Size classes: macrofossils, mesofossils, microfossils

Preservation: permineralized, replaced with pyrite

Collection methods: sieve,

• mostly chelonian shell fragments (30)

Primary reference: B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 28406: authorized by John Alroy, entered by Erin Leckey on 12.02.2003, edited by Matthew Carrano and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Actinopteri
Amiiformes - Palaeolabridae	? Palaeolabrus sp.¹³ ? Palaeolabrus sp.¹³ Casier 1967 bowfin
Lepisosteiformes - Lepisosteidae	Atractosteus sp.¹³ Atractosteus sp.¹³ Rafinesque 1820 gar
Elopiformes - Phyllodontidae	Phyllodontidae indet.¹³ Phyllodontidae indet.¹³ Sauvage 1875
Reptilia
Pterosauria - Azhdarchidae	cf. Azhdarcho sp.¹³ cf. Azhdarcho sp.¹³ Nessov 1981 pterosaur
Ornithischia - Rhabdodontidae	Rhabdodon sp.¹⁴ Rhabdodon sp.¹⁴ Matheron 1869 rhabdodontid
Ornithischia - Hadrosauridae	Hadrosauridae indet.¹³ Hadrosauridae indet.¹³ Cope 1869 hadrosaurid MCNA 10510, tooth
Ornithischia - Nodosauridae	Nodosauridae indet.¹, cf. Struthiosaurus austriacus¹² Nodosauridae indet.¹ Marsh 1890 ankylosaur cf. Struthiosaurus austriacus¹² Bunzel 1871 ankylosaur a partial skeleton, including the synsacrum, portions of the pelvic girdle and hindlimbs, a dorsal vertebra, dorsal rib fragments and a dermal scute
Saurischia -	Lirainosaurus astibiae n. gen. n. sp.¹³ Lirainosaurus astibiae n. gen. n. sp.¹³ Sanz et al. 1999 sauropod orig. 1 tooth
Theropoda -	Coelurosauria indet.², ? Ornithomimosauria indet.², cf. Paraves "indet. A"⁷, cf. Paraves "indet. B"⁷, "cf. Paronychodon sp." = Paronychodon⁷, "cf. Euronychodon sp." = Paronychodon¹³ Coelurosauria indet.² coelurosaur ? Ornithomimosauria indet.² Barsbold 1976 ornithomimosaur cf. Paraves "indet. A"⁷ Sereno 1997 maniraptoran teeth, MCNA 14523–14529 cf. Paraves "indet. B"⁷ Sereno 1997 maniraptoran teeth, MCNA 14530–14546, 14548–14561 "cf. Paronychodon sp." = Paronychodon⁷ Cope 1876 maniraptoran teeth, MCNA 14547, 14562–14565, 16755 "cf. Euronychodon sp." = Paronychodon¹³ Cope 1876 maniraptoran
Theropoda - Dromaeosauridae	Dromaeosauridae indet.⁷, Dromaeosauridae indet.¹³, cf. Richardoestesia "sp. A"⁷, cf. Richardoestesia "sp. B"⁷ Dromaeosauridae indet.⁷ Colbert and Russell 1969 maniraptoran teeth, MCNA 14623–14626 Dromaeosauridae indet.¹³ Colbert and Russell 1969 maniraptoran cf. Richardoestesia "sp. A"⁷ Currie et al. 1990 maniraptoran teeth, MCNA 14566–14610, 14612–14622, 16768, 16779, 16780, 16786–16788, 16790, 16803, 16804, 16816–16823, 16825–16830, 16832, 16833, 16837, 16841, 16846–16849, 16851, 16852, 16857–16861 cf. Richardoestesia "sp. B"⁷ Currie et al. 1990 maniraptoran MCNA 16765–16767, 16769–16778, 16781–16785, 16789, 16791–16802, 16805–16807, 16814, 16815, 16824, 16831, 16834–16836, 16838–16840, 16842–16845, 16850, 16853–16856
Theropoda - Abelisauridae	? Abelisauridae indet.¹, Abelisauridae indet.⁸, Arcovenator sp.⁷ ? Abelisauridae indet.¹ Bonaparte and Novas 1985 ceratosaur Abelisauridae indet.⁸ Bonaparte and Novas 1985 ceratosaur femora, MHNV Arcovenator sp.⁷ Tortosa et al. 2014 ceratosaur teeth, MCNA 1852, 1853, 1855, 2205, 2206, 8589, 8600, 10082, 14067, 14073, 14074, 14520–14522, 16756–16764, 16808–16813, 16862–16864
Loricata - Hylaeochampsidae	Acynodon iberoccitanus¹³ Acynodon iberoccitanus¹³ Buscalioni et al. 1997 crocodilian
Loricata - Allodaposuchidae	Musturzabalsuchus buffetauti¹³, Allodaposuchus sp.¹⁵ Musturzabalsuchus buffetauti¹³ Buscalioni et al. 1997 crocodilian Allodaposuchus sp.¹⁵ Nopcsa 1928 crocodilian may be Allodaposuchus
Loricata -	Ischyrochampsa sp.¹³ Ischyrochampsa sp.¹³ Vasse 1995 crocodilian
Testudinata - Helochelydridae	Helochelydra nopcsai¹¹, Solemys vermiculata n. sp.¹⁰, Solemys vermiculata⁹ Helochelydra nopcsai¹¹ Lapparent de Broin and Murelaga 1999 turtle partial shell, lots of fragments Solemys vermiculata n. sp.¹⁰ Lapparent de Broin and Murelaga 1996 turtle hypodigme around 260 shell fragments, most of them are peripherals and pleural fragments, around 25 fragments of plastron including the hologype Solemys vermiculata⁹ Lapparent de Broin and Murelaga 1996 turtle
Testudines - Dortokidae	Dortoka vasconica n. gen. n. sp.¹⁰ Dortoka vasconica n. gen. n. sp.¹⁰ Lapparent de Broin and Murelaga 1996 turtle
Testudines - Bothremydidae	? Elochelys sp.¹¹, "Polysternon atlanticum n. sp." = Iberoccitanemys atlanticum¹⁰ ? Elochelys sp.¹¹ Nopsca 1931 sideneck turtle right partial epiplastron "Polysternon atlanticum n. sp." = Iberoccitanemys atlanticum¹⁰ Lapparent de Broin and Murelaga 1996 sideneck turtle
Squamata -	Lacertilia "indet. 1"¹⁶, Lacertilia "indet. 2"¹⁶, Lacertilia "indet. 3"¹⁶, Lacertilia indet.¹⁶ Lacertilia "indet. 1"¹⁶ lizard numerous remains (teeth) Lacertilia "indet. 2"¹⁶ lizard teeth Lacertilia "indet. 3"¹⁶ lizard teeth Lacertilia indet.¹⁶ lizard vertebrae
Squamata - Iguanidae	Iguanidae indet.¹⁶ Iguanidae indet.¹⁶ Oppel 1811 squamates
Squamata -	Amphisbaenia indet.¹, ? Amphisbaenia indet.¹⁶, Scincomorpha indet.¹⁶ Amphisbaenia indet.¹ Gray 1844 worm lizard ? Amphisbaenia indet.¹⁶ Gray 1844 worm lizard 10 vertebrae Scincomorpha indet.¹⁶ squamates
Squamata - Madtsoiidae	Herensugea caristiorum¹⁶, "Madtsoia laurasiae" = Menarana laurasiae¹⁶ Herensugea caristiorum¹⁶ Rage 1996 snake "Madtsoia laurasiae" = Menarana laurasiae¹⁶ Rage 1996 snake
Mammalia
Tribosphenida -	Theria "indet. B"¹, Eutheria indet.⁵ Theria "indet. B"¹ Parker and Haswell 1897 mammal Eutheria indet.⁵ eutherian
Tribosphenida - Zhelestidae	Lainodon " new species "⁶, Lainodon orueetxebarriai n. gen. n. sp.⁴, Lainodon cf. orueetxebarriai⁶, Lainodon orueetxebarriai⁶, Lainodon ragei n. sp.⁶ Lainodon " new species "⁶ Gheerbrant and Astibia 1994 eutherian L1AT 18 (talonid of left M/3) and L1AT 13 and (talonid of right M/1 or M/2) Lainodon orueetxebarriai n. gen. n. sp.⁴ Gheerbrant and Astibia 1994 eutherian Lainodon cf. orueetxebarriai⁶ Gheerbrant and Astibia 1994 eutherian L1AT 20 (distal part of right lower premolar) Lainodon orueetxebarriai⁶ Gheerbrant and Astibia 1994 eutherian new species Lainodon ragei n. sp.⁶ Gheerbrant and Astibia 2012 eutherian may be Marsupialia
Amphibia
Temnospondyli - Albanerpetontidae	Albanerpetontidae indet.³ Albanerpetontidae indet.³ Fox and Naylor 1982 tetrapod
Salientia -	Anura indet.³ Anura indet.³ frog distal part of 2 humeri, 8 ilia, fragmentary maxillaries, 2 broken sphenethmoids
Salientia - Discoglossidae	Discoglossidae indet.¹ Discoglossidae indet.¹ Günther 1859 painted frog 6 prearticulars, 5 presacral verts, 7 sacral verts, 1 urostyle, 7 ilia, 2 humeri
Salientia - Palaeobatrachidae	Palaeobatrachidae indet.³ Palaeobatrachidae indet.³ Cope 1865 frog 43 prearticulars, 1 presacral vert, 2 synsacrums, 6 distal parts of humeri
Caudata -	Caudata indet.³ Caudata indet.³ salamander may be Salamandra

¹Astibia et al. 1990; ²Canudo and Ruiz-Omeñaca 2003; ³Duffaud and Rage 1999; ⁴Gheerbrant and Astibia 1994; ⁵Gheerbrant and Astibia 1999; ⁶Gheerbrant and Astibia 2012; ⁷Isasmendi et al. 2022; ⁸Isasmendi et al. 2021; ⁹Joyce 2017; ¹⁰Lapparent de Broin and Murelaga 1996; ¹¹Lapparent de Broin and Murelaga 1999; ¹²Le Loeuff and Buffetaut 1995; ¹³Pereda-Suberbiola et al. 2000; ¹⁴Pereda-Suberbiola and Sanz 1999; ¹⁵Puértolas-Pascual et al. 2016; ¹⁶Rage 1999