Åsen (Cretaceous of Sweden)

Also known as Asen

Where: Skåne, Sweden (56.1° N, 14.5° E: paleocoordinates 46.6° N, 14.5° E)

• coordinate stated in text

• outcrop-level geographic resolution

When: Belemnellocamax mammillatus ammonoid zone, Early/Lower Campanian (83.5 - 70.6 Ma)

• from two beds in lower part (1 m) of section; the "marine sand unit"; latest Early Campanian

• group of beds-level stratigraphic resolution

Environment/lithology: coastal; unlithified, shelly/skeletal, calcareous sandstone and unlithified, coarse-grained, shelly/skeletal, green sandstone

• shallow water marine storm deposit
• small tooth from "highly calcareous quartz sand with extremely abundant and well-preserved belemnite guards and articulated oysters" from uppermost 0.5 m of uppermost part of sequence; remainder from "a mottled, 10-cm thick storm deposit comprising greenish, coarse quartz sand mixed with fragmented oysters, belemnites and common vertebrate remains."

Size classes: macrofossils, mesofossils

Collected in 1990s, 2001, 2010

Collection methods: bulk, quarrying, chemical,

Primary reference: M. Siverson. 1992. Late Cretaceous Paraorthacodus (Palaeospinacidae, Neoselachii) from Sweden. Journal of Paleontology 66:994-1001 [M. Clapham/M. Clapham]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 74334: authorized by Matthew Carrano, entered by Matthew Carrano on 02.08.2007, edited by Philip Mannion, Jonathan Tennant and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Bivalvia
 Ostreida - Ostreidae
Ostreidae indet.4
Ostreidae indet.4 Rafinesque 1815 oyster
Cephalopoda
 Belemnitida - Belemnitellidae
Belemnitella mucronata5
Belemnitella mucronata5 Schlotheim 1813 belemnite
 Belemnitida -
Belemnellocamax mammillatus2, Gonioteuthis quadrata5
Chondrichthyes
 Synechodontiformes - Paraorthacodontidae
Paraorthacodus conicus, Paraorthacodus andersoni
Paraorthacodus conicus Davis 1890 elasmobranch
Paraorthacodus andersoni Case 1978 elasmobranch
 Synechodontiformes - Palaeospinacidae
Synechodus "sp. 1"13, Synechodus "sp. 2"13, Synechodus filipi n. sp.11
Synechodus "sp. 1"13 Woodward 1888 elasmobranch
Synechodus "sp. 2"13 Woodward 1888 elasmobranch
Synechodus filipi n. sp.11 Siversson et al. 2015 elasmobranch
 Orectolobiformes - Parascylliidae
Pararhincodon spp.13
Pararhincodon spp.13 Herman 1975 collared carpetshark
 Orectolobiformes - Orectolobidae
Cretorectolobus sp.13, Cederstroemia nilsi n. sp.10
Cretorectolobus sp.13 Case 1978 wobbegong
Cederstroemia nilsi n. sp.10 Siverson 1995 wobbegong
 Orectolobiformes - Hemiscylliidae
Chiloscyllium sp.13, Hemiscyllium hermani13
Chiloscyllium sp.13 Müller and Henle 1837 carpet shark
Hemiscyllium hermani13 Müller 1989 carpet shark
 Lamniformes - Mitsukurinidae
Anomotodon hermani9, Scapanorhynchus perssoni9
Anomotodon hermani9 Siverson 1992 goblin shark
Scapanorhynchus perssoni9 Siverson 1992 goblin shark
 Lamniformes - Otodontidae
"Cretalamna borealis" = Cretolamna borealis12, Cretalamna sarcoportheta n. sp.12
"Cretalamna borealis" = Cretolamna borealis12 Priem 1897 mackerel shark
Part of C. appendiculata in Siverson, 1992
Cretalamna sarcoportheta n. sp.12 Siversson et al. 2015 mackerel shark
 Lamniformes - Serratolamnidae
Serratolamna sp.13
Serratolamna sp.13 Landemaine 1991 mackerel shark
 Lamniformes - Carchariidae
Carcharias latus9, Carcharias aasenensis9, Carcharias tenuis9
Carcharias latus9 Davis 1890 sand shark
Carcharias aasenensis9 Siverson 1992 sand shark
Carcharias tenuis9 Davis 1890 sand shark
 Lamniformes - Alopiidae
Paranomotodon sp.9
Paranomotodon sp.9 Herman 1975 thresher shark
 Lamniformes - Anacoracidae
Squalicorax kaupi9
Squalicorax kaupi9 Agassiz 1843 crow shark
 Lamniformes - Pseudoscapanorhynchidae
"Cretodus borodini" = Protolamna borodini9
"Cretodus borodini" = Protolamna borodini9 Cappetta and Case 1975 mackerel shark
 Lamniformes - Archaeolamnidae
Archaeolamna kopingensis9
Archaeolamna kopingensis9 Davis 1890 mackerel shark
 Lamniformes - Pseudocoracidae
Pseudocorax laevis13
Pseudocorax laevis13 Leriche 1906 mackerel shark
 Carcharhiniformes - Scyliorhinidae
Scyliorhinidae "sp. 1"13, "Scyliorhinus germanicus" = Crassescyliorhinus germanicus13
Scyliorhinidae "sp. 1"13 Gill 1862 cat shark
"Scyliorhinus germanicus" = Crassescyliorhinus germanicus13 Herman 1982 cat shark
Listed as ‘Scyliorhinus’ germanicus
 Carcharhiniformes - Triakidae
Palaeogaleus sp.13, ? Paratriakis sp.13, Galeorhinus sp.13
Palaeogaleus sp.13 Gurr 1963 hound shark
? Paratriakis sp.13 hound shark
Galeorhinus sp.13 de Blainville and Prévost 1816 school shark
 Rajiformes - Rhinobatidae
Rhinobatos "sp. 1"13, Rhinobatos "sp. 2"13, Rhinobatos "sp. 3"13, Rhinobatos casieri13
Rhinobatos "sp. 1"13 Link 1790 guitarfish
Rhinobatos "sp. 2"13 Link 1790 guitarfish
Rhinobatos "sp. 3"13 Link 1790 guitarfish
Rhinobatos casieri13 Herman 1977 guitarfish
 Rajiformes -
Squatirhina sp.13
Squatirhina sp.13 Casier 1947 ray
 Selachii -
Selachii indet.2
Selachii indet.2 shark
coprolite morphotypes 3, 4
 Heterodontiformes - Heterodontidae
Heterodontus "sp. 1"13, Heterodontus "sp. 2"13
Heterodontus "sp. 1"13 de Blainville 1816 bullhead shark
Heterodontus "sp. 2"13 de Blainville 1816 bullhead shark
 Squatiniformes - Squatinidae
Squatina spp.13, Squatina (Squatina) fortemordeo n. sp.11, Squatina (Squatina) lundegreni n. sp.11
Squatina spp.13 Dumeril 1806 angel shark
Squatina (Squatina) fortemordeo n. sp.11 Siversson et al. 2015 angel shark
Squatina (Squatina) lundegreni n. sp.11 Siversson et al. 2015 angel shark
 Squaliformes - Squalidae
Squalidae spp.13
Squalidae spp.13 Bonaparte 1834 dogfish shark
 Batoidea -
Rhinobatoidea "sp. 2"13
 Hybodontiformes - Hybodontidae
Hybodus sp.13
Hybodus sp.13 Agassiz 1834 elasmobranch
 Hybodontiformes - Polyacrodontidae
Polyacrodus siversoni13, Polyacrodus sp.13
Polyacrodus siversoni13 elasmobranch
Polyacrodus sp.13 Jaekel 1889 elasmobranch
Gnathostomata
  -
Osteichthyes indet.2
Osteichthyes indet.2 bony fish
coprolite morphotype 5
Reptilia
 Testudines -
"Chelonia indet." = Testudines13
"Chelonia indet." = Testudines13 Batsch 1788 turtle
 Squamata - Mosasauridae
Tylosaurus ivoensis5, Hainosaurus sp.3, ? Platecarpus sp.13, Clidastes propython6, "Dollosaurus sp." = Prognathodon13, Eonatator sternbergi13, "Halisaurus sternbergi" = Eonatator sternbergii7
Tylosaurus ivoensis5 Russell 1967 mosasaur
Hainosaurus sp.3 Dollo 1885 mosasaur
? Platecarpus sp.13 Cope 1869 mosasaur
Clidastes propython6 Cope 1869 mosasaur
100 marginal teeth (LO 8231t, 8232t and 86 unnumbered, Lund University; RM PZ R1776, 1777, 1794–1803), 13 pterygoid teeth (RM PZ R1804, 1805; 11 unnumbered, Lund University), four cervical vertebrae (RM PZ R1778, 1779, 1782, 1789), six cervical-dorsal vertebrae (RM PZ R1783–1788), two dorsal vertebrae (RM PZ R1780, 1781) and 11 caudal vertebrae (RM PZ R1806–1816)
"Dollosaurus sp." = Prognathodon13 Dollo 1889 mosasaur
7 unnumbered teeth, Lund University
"Halisaurus sternbergi" = Eonatator sternbergii7 Wiman 1920 mosasaur
58 teeth (RM PZ R1811-1819; 49 unnumbered) and one caudal vertebra (RM PZ R1810)
 Ornithischia - Leptoceratopsidae
Leptoceratopsidae indet.4
Leptoceratopsidae indet.4 Nopcsa 1923 leptoceratopsid
RM PZ R1827-R1834; LO 12044t, right manual phalanx
 Ornithischia -
Ornithopoda indet.8
Ornithopoda indet.8 Marsh 1881 ornithopod
LO 12045t, left pedal phalanx; LO 12046t, left pedal phalanx
 Plesiosauria - Elasmosauridae
Scanisaurus sp.13
Scanisaurus sp.13 Persson 1960 elasmosaur
Actinopteri
 Lepisosteiformes - Lepisosteidae
Lepisosteidae indet.1
Lepisosteidae indet.1 Cuvier 1825 gar
 Pycnodontiformes - Pycnodontidae
Pycnodontidae indet.1, Anomoeodus subclavatus1
Pycnodontidae indet.1 Agassiz 1833
Anomoeodus subclavatus1 Agassiz 1833
 Pachycormiformes - Pachycormidae
Protosphyraena sp.1
Protosphyraena sp.1 Leidy 1857
 Teleostei -
Teleostei indet.1
Teleostei indet.1 Müller 1846
 Aulopiformes - Enchodontidae
Enchodus sp.1
Enchodus sp.1 Agassiz 1835
 Crossognathiformes - Pachyrhizodontidae
cf. Pachyrhizodus sp.1
cf. Pachyrhizodus sp.1 Agassiz 1850
 Ichthyodectiformes - Ichthyodectidae
Ichthyodectidae indet.1
1Bazzi et al. 2015; 2Erikkson et al. 2011; 3Lindgren 2005; 4Lindgren et al. 2007; 5Lindgren and Siverson 2002; 6Lindgren and Siverson 2004; 7Lindgren and Siverson 2005; 8Poropat et al. 2016; 9Siverson 1992; 10Siverson 1995; 11Siversson et al. 2015; 12Siversson et al. 2015; 13Sørensen et al. 2013