PBDB Collection

Paali Nala level C2 (DBC2) (Oligocene of Pakistan)

Also known as Bugti Hills; Gandoï syncline; DB-C2; Dera Bugti C2

Where: Balochistan, Pakistan (28.9° N, 69.1° E)

• Paleocoordinates: 14.1° N, 70.9° E (Wright 2013)

• coordinate stated in text

• small collection-level geographic resolution

When: Bugti Member (Chitarwata Formation), Early/Lower Oligocene (33.9 - 28.4 Ma)

• "in the lowermost levels of the Lower Chitarwata Formation, which is early Oligocene in age"

• bed-level stratigraphic resolution

Environment/lithology: fluvial-deltaic; sandstone

Size classes: macrofossils, mesofossils

Collected in 2000 - 2004

Collection methods: bulk, sieve,

• material is in the collection of the Institut des Sciences d l'Evolution, Université de Montpellier II; collections were made by "intensive screen-washing"

•

Primary reference: L. Marivaux, P.-O. Antoine, S. R. H. Baqri, M. Benammi, and Y. Chaimanee. 2005. Anthropoid primates from the Oligocene of Pakistan (Bugti Hills): Data on early anthropoid evolution and biogeography. Proceedings of the National Academy of Sciences 102(24):8436-8441 [J. Alroy/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 78520: authorized by John Alroy, entered by John Alroy on 07.02.2008, edited by K. Christopher Beard, Mark Uhen, Pauline Coster and Philip Mannion

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names

Mammalia
Perissodactyla - Rhinocerotidae	Epiaceratherium cf. magnum² Epiaceratherium cf. magnum² Uhlig 1999 rhinoceros DBC2-RHI-1-3
Artiodactyla - Suidae	Hyotherium sp.¹¹ Hyotherium sp.¹¹ von Meyer 1834 pig UMC-DBC 2242 and 2244
Artiodactyla - Tayassuidae	Pecarichoerus sp.¹¹ Pecarichoerus sp.¹¹ Colbert 1933 peccary UMC-DBC 2240 and 2241
Artiodactyla -	Ruminantia indet.⁹ Ruminantia indet.⁹ ruminant DBC2 5037
Artiodactyla - Lophiomerycidae	Nalameryx savagei¹⁰, Nalameryx sulaimani n. sp.¹⁰, Lophiomeryx kargilensis¹⁰ Nalameryx savagei¹⁰ Nanda and Sahni 1990 ruminant DBC2 specimens Nalameryx sulaimani n. sp.¹⁰ Métais et al. 2009 ruminant DBC2 5108 - type Lophiomeryx kargilensis¹⁰ Nanda and Sahni 1990 ruminant DBC2 5009-5011, 5028, 5029, 5034, 5035, 5037, 5193-5200
Artiodactyla - Palaeomerycidae	aff. Amphitragulus sp.⁹ aff. Amphitragulus sp.⁹ Pomel 1847 ruminant DBC2 5038, 5041, 5157, 5161-5166, 5168 and 5185
Artiodactyla -	Mosaicomeryx aff. quercyi⁹, Paalitherium gurki n. gen. n. sp.⁹, Sanitherium sp.¹¹ Mosaicomeryx aff. quercyi⁹ Jehenne 1987 ruminant DBC2 5040, 5043, 5158, 5160, 5172, 5173 and 5182 Paalitherium gurki n. gen. n. sp.⁹ Métais et al. 2017 ruminant DBC2 5186 - holotype; referred material: DBC2 5036, 5042 and 5181 Sanitherium sp.¹¹ von Meyer 1866 ruminant UMC-DBC 2239
Rodentia - Aplodontidae	Oligopetes sp.⁶ Oligopetes sp.⁶ Heissig 1979 mountain beaver
Rodentia - Cricetidae	Atavocricetodon paaliense n. sp.⁴, Pseudocricetodon nawabi n. sp.⁴ Atavocricetodon paaliense n. sp.⁴ Marivaux et al. 1999 rodent Pseudocricetodon nawabi n. sp.⁴ Marivaux et al. 1999 rodent
Rodentia -	Downsimys margolisi⁶ Downsimys margolisi⁶ rodent
Rodentia - Dipodidae	Allosminthus sp.⁶ Allosminthus sp.⁶ Wang 1985 rodent
Rodentia - Diatomyidae	cf. Fallomus sp.⁶, Fallomus razae⁶, Fallomus quraishyi n. sp.⁶, Fallomus ginsburgi n. sp.⁶ cf. Fallomus sp.⁶ Flynn et al. 1986 rodent DBC 295, 296, 476-494 Fallomus razae⁶ Flynn et al. 1986 rodent DBC 474, 475, 1001-1259, 1277 Fallomus quraishyi n. sp.⁶ Marivaux and Welcomme 2003 rodent DBC 438 - type Fallomus ginsburgi n. sp.⁶ Marivaux and Welcomme 2003 rodent DBC 683 - type
Rodentia -	Hodsahibia azrae⁶, Hodsahibia beamshaiensis n. sp.⁶, Hodsahibia gracilis n. sp.⁶, Bugtimys zafarullahi n. gen. n. sp.⁵ Hodsahibia azrae⁶ Flynn et al. 1986 rodent DBC 308-373 Hodsahibia beamshaiensis n. sp.⁶ Marivaux and Welcomme 2003 rodent DBC 1438 - type Hodsahibia gracilis n. sp.⁶ Marivaux and Welcomme 2003 rodent DBC 302 - type Bugtimys zafarullahi n. gen. n. sp.⁵ Marivaux et al. 2002 rodent DBC 1261 - holotype
Rodentia - Chapattimyidae	Baluchimys chaudryi⁶, Zindapiria quadricollis⁶ Baluchimys chaudryi⁶ Flynn and Cheema 1994 rodent Zindapiria quadricollis⁶ Flynn and Cheema 1994 rodent
Primates - Ekgmowechashalidae	Bugtilemur mathesoni n. gen. n. sp.⁷ Bugtilemur mathesoni n. gen. n. sp.⁷ Marivaux et al. 2001 primate
Primates - Sivaladapidae	Guangxilemur singsilai n. sp.⁸ Guangxilemur singsilai n. sp.⁸ Marivaux et al. 2002 primate
Primates - Eosimiidae	Phileosimias kamali n. gen. n. sp., Phileosimias brahuiorum n. sp. Phileosimias kamali n. gen. n. sp. Marivaux et al. 2005 monkey UMC-DBC 2199 - holotype; referred material: UMC-DBC 2192-2219 Phileosimias brahuiorum n. sp. Marivaux et al. 2005 monkey UMC-DBC 2221 - type; referred material: UMC-DBC 2220 and 2222
Primates - Amphipithecidae	Bugtipithecus inexpectans n. gen. n. sp. Bugtipithecus inexpectans n. gen. n. sp. Marivaux et al. 2005 monkey UMC-DBC 2174 - holotype; referred material: UMC-DBC 2172, 2173, 2175-2178 and 2191
Tribosphenida - Herpetotheriidae	Asiadidelphis akbarbugtii n. sp.³ Asiadidelphis akbarbugtii n. sp.³ Crochet et al. 2007 marsupial PMNH−DBC2−2180 - type; referred material: PMNH−DBC2−2179 and 2223
Chondrichthyes
Carcharhiniformes - Sphyrnidae	Sphyrna sp.¹ Sphyrna sp.¹ Rafinesque 1810 hammerhead shark
Carcharhiniformes - Carcharhinidae	Carcharhinus sp.¹, Carcharhinus perseus n. sp.¹, Rhizoprionodon sp.¹ Carcharhinus sp.¹ de Blainville 1816 requiem shark species 2 Carcharhinus perseus n. sp.¹ Adnet et al. 2007 requiem shark Rhizoprionodon sp.¹ Whitley 1929 sharpnose shark
Myliobatiformes - Gymnuridae	Himantura sp.¹ Himantura sp.¹ Müller and Henle 1837 stingray
Myliobatiformes - Myliobatidae	Aetobatus sp.¹, Myliobatis sp.¹ Aetobatus sp.¹ Blainville 1816 eagle ray Myliobatis sp.¹ Cuvier 1816 eagle ray
Myliobatiformes - Dasyatidae	Dasyatis sp.¹ Dasyatis sp.¹ Rafinesque 1810 whiptail stingray
Rajiformes - Rhinobatidae	Rhinobatos sp.¹ Rhinobatos sp.¹ Link 1790 guitarfish
Rhinopristiformes - Pristidae	? Pristis cf. lathami¹ ? Pristis cf. lathami¹ Galeotti 1837 sawfish
Rhinopristiformes - Rhynchobatidae	Rhynchobatus cf. pristinus¹ Rhynchobatus cf. pristinus¹ Probst 1877 wedgefish

¹Adnet et al. 2007; ²Antoine et al. 2003; ³Crochet et al. 2007; ⁴Marivaux et al. 1999; ⁵Marivaux et al. 2002; ⁶Marivaux and Welcomme 2003; ⁷Marivaux et al. 2001; ⁸Marivaux et al. 2002; ⁹Métais et al. 2017; ¹⁰Métais et al. 2009; ¹¹Orliac et al. 2010