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Olenellinae
Taxonomy
Olenellinae was named by Walcott (1890).
It was assigned to Olenellidae by Harrington (1956), Palmer and Repina (1993), Lieberman (1998), Palmer (1998), Lieberman (1999), Skovsted (2006).
It was assigned to Olenellidae by Harrington (1956), Palmer and Repina (1993), Lieberman (1998), Palmer (1998), Lieberman (1999), Skovsted (2006).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 1890 | Olenellinae Walcott |
| 1956 | Olenellinae Harrington p. 57 |
| 1993 | Olenellinae Palmer and Repina p. 21 |
| 1998 | Olenellinae Lieberman p. 68 |
| 1998 | Olenellinae Palmer p. 659 |
| 1999 | Olenellinae Lieberman p. 8 |
| 2006 | Olenellinae Skovsted p. 1093 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. †Olenellinae Walcott 1890
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G. †Olenellus Hall 1861
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Subg. †Olenellus (Angustolenellus) Palmer and Repina 1993
Subg. †Olenellus (Georgiellus) Moberg 1899
Subg. †Olenellus (Mesolenellus) Palmer and Repina 1993
Subg. †Olenellus (Olenelloides) Peach 1894
Subg. †Olenellus (Paedeumias) Walcott 1910
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†Olenellus (Paedeumias) chiefensis Palmer 1998
†Olenellus (Paedeumias) puertoblancoensis Lochman 1952
†Olenellus (Paedeumias) terminatus Palmer 1998
†Paedeumias granulatus Palmer 1964
†Paedeumias transitans Walcott 1910
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Invalid names: Olenellus perkinsi Resser and Howell 1938 [synonym]
†Olenellus agellus Resser and Howell 1938
†Olenellus alius Resser and Howell 1938
†Olenellus brachyomma Palmer and Halley 1979
†Olenellus bufrontis Fritz 1991
†Olenellus buttsi Resser 1938
†Olenellus clarki Resser 1928
†Olenellus claytoni Walcott 1910
†Olenellus crassimarginatus Walcott 1910
†Olenellus georgiensis Resser and Howell 1938
†Olenellus getzi Dunbar 1925
†Olenellus gigas Peach 1894
†Olenellus gilberti White 1874
†Olenellus howelli White 1874
†Olenellus latilimbatus Resser and Howell 1938
†Olenellus megafrontalis Vaccari 1988
†Olenellus nevadensis Walcott 1910
†Olenellus parvofrontatus Fritz 1991
†Olenellus peculiaris Resser and Howell 1938
†Olenellus puertoblancoensis Lochman 1952
†Olenellus robsonensis Burling 1916
†Olenellus roddyi Resser and Howell 1938
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Invalid names: Olenellus jonasae Resser and Howell 1938 [synonym], Olenellus nitidus Resser and Howell 1938 [synonym], Olenellus nodosus Resser and Howell 1938 [synonym], Olenellus similaris Resser and Howell 1938 [synonym], Paedeumias eboracense Resser and Howell 1938 [synonym], Paedeumias glabrum Resser and Howell 1938 [synonym], Paedeumias yorkense Resser and Howell 1938 [synonym]
†Olenellus romensis Resser and Howell 1938
†Olenellus santuccii Webster and Caron 2024
†Olenellus schucherti Resser and Howell 1938
†Olenellus terminatus Palmer 1998
†Olenellus thompsoni Hall 1859
†Olenellus walcotti Shaler and Foerste 1888
†Olenellus wanneri Resser and Howell 1938
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| A. R. Palmer and L. N. Repina 1993 | Olenellidae with glabellar furrows weakly to moderately defined. Transverse width of anterior part of LI equal to or only slightly less than transverse width of occipital ring. L4 slightly to moderately expanded anteriorly. Posterior tips of ocular lobes ranging from opposite L2 to opposite posterior part of occipital ring. Posterior margin of cephalon nearly straight or slightly deflected forward distal to intergenal spine or swelling. Intergenal spines or distinct intergenal angle usually present. Prothorax, where known, with axis narrower than inner part of pleurae. Third thoracic segment weakly to moderately macropleural, with pleural spine not greatly elongated. Fifteenth thoracic segment with long axial spine. Opisthothorax with variable number of uniformly small segments, each bearing narrow pleurae. Pygidium small, subquadrate, with 1 or 2 pairs of short marginal spines. | |
| B. S. Lieberman 1999 | The Olenellinae is characterized by the following shared characters when present in combination: 1. posterolateral margins of L4 divergent anteriorly; 2. transverse profile of ocular lobes flattened dorsally; 3. posterior tips of ocular lobes developed opposite medial part of distal margin of L0 or distal tip of S0; 4. S3 not conjoined medially; 5. line between ends of S2 directed inward and posteriorly at 45 degree angle relative to transverse line; 6. S0 not conjoined medially; 7. lateral lobes on L0 absent; 8. glabellar furrows moderately to strongly incised; 9. genal ridge prominently developed; 10. extraocular region opposite L1 broad, width (tr.) greater than or equal to 75% of width of glabella at L1; 11. genal spine near where it hits cephalic border directed posterolaterally at roughly 10 to 20 degree angle relative to sagittal line; 12. length (exsag.) of genal spine roughly equal to length (sag.) of first four to six thoracic segments; 13. genal spine angle developed opposite medial part of distal margin of L0; 14. intergenal angle developed adjacent to or directly behind genal spine angle; 15. intergenal angle with small, pointed spine; 16. intergenal angle deflected at roughly 10 degree angle relative to transverse line; 17. medial part of cephalic posterior border between intergenal angle and L0 weakly flexed anteriorly; 18. anterior margin of third thoracic pleural segment, before flexing strongly posteriorly, parallel to transverse line or weakly deflected posteriorly; 19. posterior margin of thoracic pleural furrow on third thoracic segment directed evenly posterolaterally; 20. thoracic pleural spines on seg- ments T5 to T8 developed as broad sweeping projections extending roughly four segments back; 21. length (exsag.) of thoracic pleural furrows (excluding those of T3) at medial part of segment equal to roughly 1.3 times length (exsag.) of posterior band of pleural segment; 22. width of thoracic pleural spines T5 to T8 at spine midlength more than two-thirds length (exsag.) of medial part of inner pleural region; 23. length (sag.) of spine on 15th thoracic segment roughly equal to length of whole prothorax; 24. base of spine on 15th thoracic segment nearly as wide (tr.) as axis of segment; 25. segments of opisthothorax with reduced pleurae; 26. pygidium length (sag.) 1.5 times width (tr.); 27. posterior mar- gin of pygidium weakly convex. Of these characters, only character 25 is unique to the Olenellidae. |