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Ellipsocephalidae

Trilobita - Redlichiida - Ellipsocephalidae

Synonymy list
YearName and author
1887Ellipsocephalidae Matthew
1997Ellipsocephalidae Whittington et al. p. 302
2002Ellipsocephalidae Jell and Adrain p. 470
2008Ellipsocephalidae Landing et al. p. 893
2011Ellipsocephalidae Adrain p. 104
2022Ellipsocephalidae Cederström et al. p. 31
2024Ellipsocephalidae El Albani et al. p. S7

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
RankNameAuthor
subphylumArtiopoda(Hou and Bergstrom)
classTrilobitaWalch 1771
orderRedlichiidaRichter 1932
suborderRedlichiinaRichter 1932
superfamilyEllipsocephaloideaLin 1990
familyEllipsocephalidaeMatthew 1887
familyEllipsocephalidaeMatthew 1887

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Ellipsocephalidae Matthew 1887
show all | hide all
G. †Acadolenus Sdzuy 1967
G. †Alueva Lotze 1958
Subfm. †Antatlasiinae Hupe 1953
+
G. †Antatlasia Hupe 1952
G. †Berabichia Geyer 1990
+
†Berabichia subditus Palmer and Gatehouse 1972
G. †Argunaspis Repina 1966
G. †Asiatella Repina 1964
+
†Asiatella elegans Repina 1964
G. †Bergeroniaspis Lermontova 1951
G. †Bergeroniellus Lermontova 1940
G. †Bhargavia Peng et al. 2009
+
†Bhargavia prakritika Peng et al. 2009
G. †Blayacina Cobbold 1932
+
Invalid names: Sardaspis Brasier 1976 [synonym]
G. †Brevitermierella Hupé 1953
+
Invalid names: Paratermierella Hupe 1952 [synonym]
G. †Cambrunicornia Geyer 1990
G. †Catadoxides Matthew 1899
G. †Culmenaspis Repina 1972
Subfm. †Ellipsocephalinae Mathew 1887
+
G. †Cambrosaurura Cederström et al. 2022
G. †Dellingia Cederström et al. 2022
G. †Ellipsocephalus Zenker 1833
+
†Ellipsocephalus gripi Kautsky 1945
Invalid names: Germaropyge Snajdr 1957 [synonym]
G. †Ellipsostrenua Kautsky 1945
+
†Ellipsostrenua sularpensis Ahlberg and Bergström 1978
†Ellipsostrenua troedssoni Cederström et al. 2022
G. †Kingaspidoides Hupe 1953
+
†Kingaspidoides alberti Geyer 2017
†Kingaspidoides meieri Geyer 2017
†Kingaspidoides usitata Geyer 1990
Invalid names: Elatius Orlowski 1985 [synonym]
G. †Ornamentaspis Geyer 1990
+
†Ornamentaspis hupei Orłowski 1964
G. †Geyerorodes Özdikmen 2009
+
†Geyerorodes howleyi Walcott 1890
†Geyerorodes schmitti Geyer 1990
Invalid names: Orodes Geyer 1990 [replaced]
G. †Glabrella Lermontova 1940
+
†Glabrella pitans Palmer and Gatehouse 1972
†Glabrella ventrosa Lermontova 1940
G. †Hamatolenus Hupe 1952
+
Subg. †Hamatolenus (Hamatolenus) Hupé 1952
+
†Hamatolenus (Hamatolenus) vincenti Geyer and Landing 2004
Subg. †Hamatolenus (Myopsolenus) Hupe 1952
+
Invalid names: Collyrolenus Hupe 1952 [synonym]
G. †Issafeniella Geyer 1990
G. †Kadyella Pokrovskaya 1959
G. †Kameschkoviella Repina 1959
G. †Kijanella Repina 1958
G. †Kingaspis Kobayashi 1935
+
Subg. †Kingaspis (Kingaspidoides) Hupe 1953
+
†Kingaspis (Kingaspidoides) armatus Liñán et al. 2003
+
Invalid names: Kingaspis (Kingaspis) brevifrons Hupé 1953 [synonym]
Subg. †Kingaspis (Kingaspis) Hupé 1954
+
†Kingaspis avalonensis Geyer and Landing 2001
†Kingaspis convexus Palmer and Rowell 1995
Invalid names: Mesetaia Hupe 1953 [synonym]
G. †Krolina Repina 1960
G. †Kymataspis Geyer 1990
G. †Latikingaspis Geyer 1990
+
†Latikingaspis alatus Hupé 1954
G. †Latoucheia Hupe 1953
G. †Latuzella Repina et al. 1999
G. †Lermontovia Suvorova 1956
G. †Limataceps Repina 1969
G. †Lusatiops Richter 1941
+
†Lusatiops harveyi Walcott 1890
G. †Nelegeria Korobov 1966
+
†Nelegeria lata Korobov 1966
G. †Olekmaspis Suvorova 1956
G. †Ourikaia Geyer 1990
G. †Paramicmacca Lermontova 1951
G. †Paraprotolenella Zhu and Jiang 1981
+
†Paraprotolenella subcylindrica Zhu and Jiang 1981
G. †Planolimbus Geyer 1990
G. †Protagraulos Matthew 1895
G. †Protolenella Qian and Yao 1974
+
†Protolenella conica Qian and Yao 1974
†Protolenella lata Zhou 1974
Subfm. †Protoleninae Richter and Richter
+
G. †Anaxarete Geyer 2016
+
†Anaxarete senior Cobbold 1910
G. †Chorbusulina Lazarenko 1962
+
†Chorbusulina bella Lazarenko 1964
†Chorbusulina wilkesi Palmer and Gatehouse 1972
G. †Cuneoaxiella Cederström et al. 2022
+
†Cuneoaxiella grandis Ahlberg and Bergström 1978
G. †Protolenus Matthew 1892
+
†Protolenus elegans Matthew 1892
+
Invalid names: Protolenus paradoxoides Matthew 1892 [synonym]
†Protolenus pustulatus Cobbold 1910
Invalid names: Bergeronia Matthew 1895 [objective synonym]
G. †Pruvostinoides Hupe 1952
G. †Pseudokadyella Repina 1966
G. †Pseudolenus Hupe 1953
G. †Pseudoprotolenella Zhu and Lin 1983
+
G. †Ptychoparopsis Hupe 1953
G. †Rinconia Hupe 1953
G. †Sailycaspis Repina 1965
G. †Sectigena Geyer 1990
Subfm. †Strenuaevinae Cederström et al. 2022
+
G. †Epichalnipsus Geyer et al. 2004
G. †Strenuaeva Richter and Richter 1940
+
†Strenuaeva inflata Ahlberg and Bergström 1978
†Strenuaeva primaeva Brøgger 1878
†Strenuaeva spinosa Ahlberg and Bergström 1978
†Strenueva annio Cobbold 1910
Invalid names: Hindermeyeria Hupe 1953 [synonym]
G. †Strenuella Matthew 1887
+
Subg. †Strenuella (Comluella) Hupe 1953
†Strenuella calceola Cobbold 1931
†Strenuella cobboldi Richter 1919
†Strenuella platycephala Cobbold 1910
†Strenuella polonica Czarnocki 1926
†Strenuella strenua Billings 1874
G. †Tadakoustia Geyer 1990
G. †Termieraspis Repina 1960
G. †Termierella Hupe 1952
+
†Termierella totanesensis Sepúlveda et al. 2024
G. †Triangulaspis Lermontova 1940
+
Invalid names: Acutaspis Repina 1976 [synonym], Angustaeva Hupé 1953 [synonym], Triangulina Repina 1965 [synonym]
G. †Yeshanaspis Zhu and Jiang 1981
+
†Yeshanaspis subconicus Zhu and Jiang 1981
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingo
Life habit: low-level epifaunalo
Diet: deposit feedero
Vision: well-developedf
Created: 2009-12-19 04:00:08
Modified: 2009-12-19 06:00:08
Source: f = family, o = order, c = class
References: Aberhan et al. 2004, Whittington et al. 1997, Hendy et al. 2009

Age range: base of the Branchian to the top of the Undillan or 516.20000 to 500.50000 Ma

Collections (176 total)


Time interval Ma Country or state Original ID and collection number
Caerfai538.8 - 506.5Canada (New Brunswick) Ellipsocephalus galeatus (34572 34597) Ellipsocephalus galeatus, Protolenus elegans (34571) Kingaspidoides obliquoculatus (34573) Protolenus articephalus, Protolenus elegans (34567 34751) Protolenus elegans (34569 34570 34595 34596 34752)
Early/Lower Cambrian538.8 - 506.5Antarctica Glabrella pitans, Chorbusulina wilkesi, Chorbusulina subdita (154030)
Early/Lower Cambrian538.8 - 506.5Norway (Troms) Ellipsocephalus sp. (190927) Strenuaeva inflata (190925 190926)
Early/Lower Cambrian538.8 - 506.5Spain (Aragon) Alueva undulata, Hamatolenus ibericus, Kingaspis campbelli, Protolenus pisidianus, Protolenus jilocanus (120550 120552)
Cambrian538.8 - 486.85USA (Wisconsin) Ellipsocephalus sp. (250)
Atdabanian521.0 - 516.0Sweden Ellipsocephalus nordenskioeldi, Proampyx grandis, Proampyx sularpensis (87212) Proampyx grandis, Proampyx sularpensis (87121)
Atdabanian521.0 - 516.0Morocco (Ant-Atlas) Antatlasia hollardi (232863)
Atdabanian521.0 - 516.0Morocco Antatlasia hollardi (225084) Antatlasia sp. indet. (225083) Pruvostinoides sp. (80900)
Atdabanian521.0 - 516.0United Kingdom (Wales) Hamatolenus douglasi (41750) Myopsolenus sp. (41747)
Atdabanian521.0 - 516.0Antarctica Berabichia subditus (56123)
Atdabanian521.0 - 516.0Poland Ellipsocephalus sanctacrucensis (7905) Ellipsocephalus sanctacrucensis, Protolenus bodzanti, Protolenus medius, Strenuaeva orlowinensis, Strenuella lakei, Termierella sandomirensis, Protolenus czarnockii, Strenuaeva primaeva, Strenuella polonica (72261) Protolenus sp. (72268)
Atdabanian521.0 - 516.0Russian Federation (Khakasiya) Kijanella sp. (81312) Kijanella sp., Kijanella diffusica, Kijanella batenica, Asiatella elegans (81327) Limataceps bajangolica (81314) Strenuella paraspinosa (81328)
Atdabanian521.0 - 516.0Spain Lusatiops sp., Protolenus schneideri, Strenueva vigilans, Strenueva annio (225087)
Atdabanian521.0 - 516.0Russian Federation (Tuva) Bergeroniaspis vladimirskii (81465)
Atdabanian521.0 - 516.0Russian Federation (Siberia) Triangulaspis lermontovae, Triangulaspis annio (96839) Triangulaspis sp. (56855) Triangulaspis sp., Protolenidae indet. (232892)
Atdabanian521.0 - 516.0Russian Federation (Kemorovo) Strenuella sp. (57019)
Nangaoian521.0 - 514.5China (Guizhou) Protolenella conica, Protolenella lata (107130)
Nangaoian521.0 - 514.5China (Xinjiang) Pseudoprotolenella subcylindrica, Pseudoprotolenella xinjiangensis (127612)
Nangaoian521.0 - 514.5China (Jiangsu) Yeshanaspis subconicus, Paraprotolenella subcylindrica (127483)
Atdabanian - Lenian521.0 - 508.2Germany (Saxony) Comluella sp., Lusatiops sp. (6899)
Stage 3 - Stage 4521.0 - 506.5Sweden (Lappland) Ellipsocephalus gripi (190924)
Branchian516.2 - 506.5Canada (New Brunswick) Berabichia milleri (34579 34580) Berabichia milleri, Kingaspidoides obliquoculatus (34575 34576 34577 34578)
Branchian516.2 - 506.5USA (Massachusetts) Strenuella strenua (56838 56854)
Branchian516.2 - 506.5United Kingdom (England) Protolenus indet., Strenuella coboldi, Strenuella calceola, Pagetia annio, Strenuella platycephala, Protolenus pustulatus (79928)
Branchian516.2 - 506.5Canada (Newfoundland and Labrador) Solenopleura harveyi, Solenopleura howleyi, Agraulos nasutus (99495)
Botomian516.0 - 512.1Russian Federation (Yakutia) Bergeroniaspis sp., Bergeroniellus sp. (90438) Triangulaspis sp. (174246 232979)
Botomian516.0 - 512.1Russian Federation (Tuva) Bergeroniaspis divergens, Bergeroniaspis dualis, Nelegeria lata (9978) Ellipsocephalus sp. (81480) Kadyella ubsanurica (81484) Strenuaeva repentina (81472)
Botomian516.0 - 512.1Russian Federation (Kemorovo) Latuzella poletaevae (57020)
Botomian516.0 - 512.1Morocco Berabichia vertumnia (80850 80851)
Botomian516.0 - 512.1Norway (Mjosen) Ellipsocephalus nordenskjoldi, Strenuella primaeva, Strenuella linnarssoni (42220)
Botomian516.0 - 512.1Russian Federation Bergeroniaspis dualis (233054) Bergeroniellus expansus (225138) Lermontovia dzevanovskii (225139) Triangulaspis lermontovae, Triangulaspis annio (233047)
Botomian516.0 - 512.1Spain (Andalucia) Strenueva sampelayoi, Strenueva melendezi (8093)
Botomian516.0 - 512.1Poland Ellipsocephalus sanctacrucensis, Strenuaeva orlowinensis (153007)
Lenian516.0 - 508.2Sweden Comluella scanica, Ellipsocephalus lunatus (87214) Comluella scanica, Ellipsocephalus lunatus, Strenuaeva sp. (87123) Comluella scanica, Strenuaeva sp. (87187) Proampyx linnarssoni (87122 87213)
Lenian516.0 - 508.2Morocco Hamatolenus sp. (161461) Kingaspidoides neglectus, Latoucheia epichara, Latoucheia ourikaensis, Ourikaia calva, Protolenus dimarginatus, Protolenus hupei, Protolenus termierelloides (9801) Kingaspis sp., Hamatolenus vincenti, Protolenus densigranulatus (161460) Latikingaspis sulcatus (9793)
Stage 4514.5 - 506.5Sweden (Laisvall) Strenuella primaeva (239931)
Stage 4514.5 - 506.5Morocco Protolenus sp. (235700)
Stage 4514.5 - 506.5Sweden Cuneoaxiella grandis (240025) Cuneoaxiella grandis, Ellipsostrenua linnarssoni (236574) Cuneoaxiella grandis, Ellipsostrenua linnarssoni, Ellipsostrenua sularpensis (240026) Cuneoaxiella grandis, Ellipsostrenua linnarssoni, Ellipsostrenua sularpensis, Ellipsostrenua troedssoni (240027 240028) Cuneoaxiella grandis, Ellipsostrenua linnarssoni, Ellipsostrenua sularpensis, Ellipsostrenua troedssoni, Ellipsostrenua spinosa (236573 240029) Cuneoaxiella grandis, Ellipsostrenua linnarssoni, Ellipsostrenua troedssoni, Ellipsostrenua spinosa (236571 236572) Strenuaeva inflata (237137)
Toyonian512.1 - 508.2Russian Federation (Khakasiya) Glabrella mrassina (81331)
Toyonian512.1 - 508.2Morocco Protolenus densigranulatus (9123)
Toyonian512.1 - 508.2Antarctica Kingaspis convexus (56127)
Toyonian512.1 - 508.2Norway (Mjøsa) Ellipsocephalus gripi, Ellipsocephalus linnarssoni, Ellipsostrenua sularpensis, Ellipsostrenua spinosa (174259)
Toyonian512.1 - 508.2Spain (Zaragoza) Kingaspis campbelli, Protolenus dimarginatus, Protolenus termierelloides, Protolenus jilocanus (233072)
Delamaran509.7 - 505.3Morocco Hamatolenus maroccanus, Hamatolenus sp., Kingaspidoides laetus, Kingaspidoides neglectus, Hamatolenus vincenti (174203) Hamatolenus maroccanus, Hamatolenus sp., Kingaspidoides neglectus, Kingaspidoides laetus, Latoucheia epichara, Protolenus densigranulatus (174223) Hamatolenus maroccanus, Kingaspidoides laetus, Hamatolenus meridionalis, Hamatolenus vincenti, Protolenus densigranulatus, Ornamentaspis angustigena (174208) Hamatolenus maroccanus, Kingaspidoides laetus, Latoucheia epichara, Hamatolenus meridionalis, Hamatolenus vincenti, Protolenus densigranulatus (174210) Hamatolenus maroccanus, Latoucheia epichara, Hamatolenus lotzei, Ornamentaspis frankenwaldensis (174294) Hamatolenus sp., Latoucheia epichara (174204) Kingaspidoides laetus (174222) Kingaspidoides laetus, Hamatolenus meridionalis, Hamatolenus lotzei, Hamatolenus vincenti, Ornamentaspis frankenwaldensis (174293) Kingaspidoides laetus, Hamatolenus vincenti, Ornamentaspis angustigena (174220) Kingaspidoides laetus, Hamatolenus vincenti, Ornamentaspis frankenwaldensis (174290) Kingaspidoides laetus, Latoucheia epichara, Hamatolenus meridionalis, Hamatolenus vincenti (174292) Kingaspidoides laetus, Latoucheia epichara, Protolenus densigranulatus (174217) Kingaspidoides laetus, Protolenus densigranulatus (174291) Kingaspidoides neglectus (174205 174206 174207) Kingaspidoides neglectus, Kingaspidoides laetus, Latoucheia epichara, Hamatolenus draensis, Protolenus densigranulatus, Ornamentaspis angustigena (174219)
Amgan508.2 - 504.5Poland Ellipsocephalus polytomus, Ellipsocephalus sandomiri, Ellipsocephalus guerichi, Ellipsocephalus jugoszovi, Kingaspis henningsmoeni, Strenuella samsonowiczi, Protolenus polonicus (72270)
Amgan508.2 - 504.5Sweden (Oland) Ellipsocephalus polytomus (225039 225040)
Amgan508.2 - 504.5Germany Cambrunicornia saxonica (162995) Cambrunicornia saxonica, Ornamentaspis todraensis, Protolenus bergstroemi (162993) Cambrunicornia saxonica, Protolenus bergstroemi (162994 162996) Protolenus bergstroemi (162997)
Amgan508.2 - 504.5Kyrgyzstan Glabrella ventrosa (156378)
Amgan508.2 - 504.5Czechia (Bohemia) Ellipsocephalus vetustus, Acadolenus snajdri (67709) Ellipsocephalus vetustus, Ellipsocephalus hoffi, Acadolenus snajdri (67708) Ellipsocephalus vetustus, Germaropyge sp. nov. (67738)
Solvan508.2 - 504.5Sweden (Norrbotten) Ellipsocephalus polytomus (9782)
Solvan - Menevian508.2 - 499.5Czechia Ellipsocephalus hoffi (73537 73539) Ellipsocephalus vetustus (73545 73561) Ellipsocephalus vetustus, Germaropyge germari (73529 73557) Germaropyge germari (203343)
Middle Cambrian506.5 - 497.0Italy (Iglesiente) Protolenus pisidianus (139878)
Middle Cambrian506.5 - 497.0India Bhargavia prakritika (116546 116547 116548 116549)
Middle Cambrian506.5 - 497.0Czechia (Bohemia) Germaropyge germari (67775)
Middle Cambrian506.5 - 497.0Germany (Bayern) Latikingaspis sp., Ornamentaspis frankenwaldensis (97598)
Middle Cambrian506.5 - 497.0Spain (Aragon) Hamatolenus lotzei, Alueva hastata, Alueva moratrix (120553) Hamatolenus sp., Alueva hastata, Alueva moratrix (120551)
Middle Cambrian506.5 - 497.0Poland Ellipsocephalus polytomus, Ellipsocephalus guerichi, Ellipsocephalus puschi, Ellipsocephalus jugoszovi, Kingaspis henningsmoeni, Strenuella samsonowiczi, Strenuella hupei, Protolenus polonicus (72269)
Middle Cambrian506.5 - 497.0Czechia (Jince) Ellipsocephalus vetustus (76483)
Middle Cambrian506.5 - 497.0Spain Kymataspis bernardi (55265)
Middle Cambrian506.5 - 497.0Czechia (Litavka) Ellipsocephalus vetustus (76453) Ellipsocephalus vetustus, Acadolenus snajdri (76454)
St Davids506.5 - 497.0Spain (Leon) Acadolenus decorus (55503)
St Davids506.5 - 497.0USA (Massachusetts) Kingaspis avalonensis (49340)
Drumian504.5 - 500.5Poland Ellipsocephalus polytomus, Ellipsocephalus sandomiri, Strenuella samsonowiczi (72271)
Drumian504.5 - 500.5Czechia (Central Boemian) Ellipsocephalus vetustus (76456)
Undillan503.0 - 500.5Germany Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186152) Kingaspidoides frankenwaldensis (186149 186150 186153 186154 186159 186165 186166 186170) Kingaspidoides meieri, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186167) Kingaspidoides meieri, Kingaspidoides frankenwaldensis (186151 186157 186158) Kingaspidoides sp. (186162 197474) Latikingaspis alatus, Kingaspidoides meieri, Kingaspidoides frankenwaldensis (186169) Ornamentaspis crassilimbata, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186155 186156 186163) Ornamentaspis crassilimbata, Kingaspidoides frankenwaldensis (186148 186160) Ornamentaspis crassilimbata, Kingaspidoides meieri, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186168) Ornamentaspis sp., Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186164)