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Cheilotrichia (Empeda)
Insecta - Diptera - Limoniidae
Taxonomy
Empeda was named by Osten Sacken (1869). It is extant. Its type is Empeda stigmatica.
It was reranked as Erioptera (Empeda) by Alexander (1931); it was reranked as Cheilotrichia (Empeda) by Evenhuis (1994), Gelhaus and Johnson (1996), Podenas (1999), Kopeć and Kania (2013), Krzemiński (2019), Kopeć et al. (2025).
It was assigned to Erioptera by Alexander (1931); to Tipulidae by Aldrich (1905), Carpenter (1992); and to Cheilotrichia by Evenhuis (1994), Gelhaus and Johnson (1996), Podenas (1999), Kopeć and Kania (2013), Krzemiński (2019), Kopeć et al. (2025).
It was reranked as Erioptera (Empeda) by Alexander (1931); it was reranked as Cheilotrichia (Empeda) by Evenhuis (1994), Gelhaus and Johnson (1996), Podenas (1999), Kopeć and Kania (2013), Krzemiński (2019), Kopeć et al. (2025).
It was assigned to Erioptera by Alexander (1931); to Tipulidae by Aldrich (1905), Carpenter (1992); and to Cheilotrichia by Evenhuis (1994), Gelhaus and Johnson (1996), Podenas (1999), Kopeć and Kania (2013), Krzemiński (2019), Kopeć et al. (2025).
Subtaxa
Cheilotrichia (Empeda) axillaris, Cheilotrichia (Empeda) bella, Cheilotrichia (Empeda) budrysi, Cheilotrichia (Empeda) cretacea, Cheilotrichia (Empeda) diacantha, Cheilotrichia (Empeda) duplicata, Cheilotrichia (Empeda) ferruginea, Cheilotrichia (Empeda) gallica, Cheilotrichia (Empeda) hyalina (syn. Gonomyia grisea), Cheilotrichia (Empeda) minuta (syn. Empeda elongata, Empeda prolifica), Cheilotrichia (Empeda) oisensis, Cheilotrichia (Empeda) palaeocenica, Cheilotrichia (Empeda) pawlowskii, Cheilotrichia (Empeda) platyphylla, Cheilotrichia (Empeda) rectistyla, Cheilotrichia (Empeda) schummeli, Cheilotrichia (Empeda) subabortiva, Cheilotrichia (Empeda) weitschati, Empeda stigmatica (type species)
Synonymy list
| Year | Name and author |
|---|---|
| 1869 | Empeda Osten Sacken p. 183 |
| 1905 | Empeda Aldrich p. 86 |
| 1931 | Erioptera (Empeda) Alexander p. 105 |
| 1992 | Empeda Carpenter |
| 1994 | Cheilotrichia (Empeda) Evenhuis |
| 1996 | Cheilotrichia (Empeda) Gelhaus and Johnson p. 59 |
| 1999 | Cheilotrichia (Empeda) Podenas p. 243 |
| 2013 | Cheilotrichia (Empeda) Kopeć and Kania p. 538 |
| 2019 | Cheilotrichia (Empeda) Krzemiński p. 508 |
| 2025 | Cheilotrichia (Empeda) Kopeć et al. p. 2 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subg. Cheilotrichia (Empeda) Osten Sacken 1869
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†Cheilotrichia (Empeda) axillaris Alexander 1931
†Cheilotrichia (Empeda) bella Podenas 1999
†Cheilotrichia (Empeda) budrysi Podenas 1999
†Cheilotrichia (Empeda) cretacea Gelhaus and Johnson 1996
†Cheilotrichia (Empeda) diacantha Alexander 1931
†Cheilotrichia (Empeda) duplicata Alexander 1931
†Cheilotrichia (Empeda) ferruginea Cockerell 1921
†Cheilotrichia (Empeda) gallica Kopeć et al. 2025
†Cheilotrichia (Empeda) hyalina Cockerell 1921
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Invalid names: Gonomyia grisea Cockerell 1921 [synonym]
†Cheilotrichia (Empeda) minuta Meunier 1899
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Invalid names: Empeda elongata Meunier 1906 [synonym], Empeda prolifica Meunier 1906 [synonym]
†Cheilotrichia (Empeda) oisensis Kopeć et al. 2025
†Cheilotrichia (Empeda) palaeocenica Krzemiński and Krzemińska 1994
†Cheilotrichia (Empeda) pawlowskii Kopeć 2019
†Cheilotrichia (Empeda) platyphylla Alexander 1931
†Cheilotrichia (Empeda) rectistyla Alexander 1931
†Cheilotrichia (Empeda) schummeli Meunier 1917
†Cheilotrichia (Empeda) subabortiva Alexander 1931
†Cheilotrichia (Empeda) weitschati Kopeć and Kania 2013
Empeda stigmatica Osten-Sacken 1869
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Turonian to the top of the Priabonian or 93.90000 to 33.90000 Ma
Minimum age of oldest fossil (stem group age): 89.8 Ma
Minimum age of oldest fossil (stem group age): 89.8 Ma
Collections (17 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Turonian | USA (New Jersey) | Cheilotrichia cretacea (105001) | |
| Ypresian | France (Oise) | Cheilotrichia oisensis, Cheilotrichia gallica (123850) | |
| Lutetian | Russian Federation | Cheilotrichia palaeocenica (107340) | |
| Priabonian | Ukraine | Cheilotrichia pawlowskii (153882) | |
| Priabonian | Russian Federation | Cheilotrichia pawlowskii (123419) | |
| Priabonian | Poland | Cheilotrichia pawlowskii (110737 120813 123215) | |
| Priabonian | Russian Federation (Kaliningrad) | Cheilotrichia bella (140808) Cheilotrichia pawlowskii (123962) Erioptera minuta (127170) Erioptera minuta, Empeda schummeli, Erioptera duplicata, Erioptera diacantha, Erioptera subabortiva, Erioptera rectistyla, Empeda prolifica, Empeda elongata (123927) Erioptera minuta, Erioptera platyphylla, Erioptera duplicata, Erioptera axillaris, Empeda prolifica (124073) | |
| Priabonian | Lithuania | Cheilotrichia budrysi (123977) | |
| Priabonian | United Kingdom (England) | Empeda ferruginea (123959) Empeda hyalina, Gonomyia grisea (123960) | |
| Priabonian | Germany (Halle) | Cheilotrichia weitschati (129228) |