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Haplohamulida

Insecta - Odonata

Taxonomy
Haplohamulida was named by Bechly (1996). It is extant.

It was assigned to Anisoptera by Bechly (2005); and to Liberaponsida by Bechly (2007).

Subtaxa
Italoansida

Synonymy list
YearName and author
1996Haplohamulida Bechly
2005Haplohamulida Bechly p. 2
2007Haplohamulida Bechly

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
infraclassHydropalaeopteraRohdendorf 1969
—Euhydropalaeoptera
—OdonatopteraMartynov 1932
—Palaeodonatoptera
—Plesiodonatoptera
—Apodonatoptera
—Paneodonatoptera
—Neodonatoptera
—Euodonatoptera
—OdonatocladaBechly 1999
—Nodialata
—Discoidalia
—Stigmoptera
RankNameAuthor
—Lobodonata
—Panodonata
orderOdonata
—Epiproctophora
—EuepiproctophoraBechly 1998
—Anisopteromorpha
—Trigonoptera
—Pananisoptera
—NeoanisopteraBechly 1998
suborderAnisopteraSelys-Longchamps and Hagen 1854
—Euanisoptera
—Exophytica
—Cavilabiata
—Cristotibiata
—Brachystigmata
—Paucipostnodalia
—Eubrachystigmata
—NeobrachystigmataBechly 2003
—PaneurypalpidomorphaBechly 2003
—EurypalpidomorphaBechly 2003
—EurypalpidiformiaBechly 2003
—Paneurypalpida
—Eurypalpida
—Neolamellida
—Valvulida
—Trichodopalpida
—Laterocarinida
—Mediocostida
—Longiansida
—Liberaponsida
—Haplohamulida
—Haplohamulida

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Haplohamulida Bechly 1996
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Unr. Italoansida Bechly 1996
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Unr. Anauriculida Bechly 1996
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Fm. †Bolcacorduliidae Gentilini 2002
+
G. †Bolcacordulia Gentilini 2002
+
†Bolcacordulia paradoxa Gentilini 2002
Unr. Libellulida Bechly 1996
+
Fm. Libellulidae Rambur 1842 [common skimmer]
+
G. †Caussanelia Nel et al. 1997
G. †Jeanlegrandia Nel et al. 2005
Subfm. Leucorrhininae Tillyard and Fraser 1940
Subfm. Libellulinae Rambur 1842
G. †Miorhodopygia Riou and Nel 1995
G. †Molertrum Zessin 2019
G. Onychothemis Brauer 1868
G. †Oryctodiplax Cavallo and Galletti 1987
G. †Palaeolibellula Fleck et al. 1999
Subfm. Pantaliinae Fraser 1957
Subfm. Pantalinae Jacobson and Bianchi 1905
+
Invalid names: Trameinae Tillyard 1917 [synonym]
G. †Prorhyothemis Prokop et al. 2003
G. †Sloveniatrum Zessin et al. 2008
Subfm. Sympetrinae Tillyard 1917
Subfm. Tetrathemistinae Tillyard 1917
Subfm. Trithemistinae Tillyard 1917
Invalid names: Libellula eusebioi Piton 1935 [nomen dubium], Libellula minuscula Oustalet 1870 [nomen dubium]
Fm. †Palaeomacromiidae Petrulevičius et al. 1999
+
G. †Bolcathemis Gentilini 2002
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†Bolcathemis nervosa Gentilini 2002
G. †Curviarculia Petrulevičius and Nel 2002
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†Curviarculia delicata Petrulevičius and Nel 2002
†Curviarculia lamasi Petrulevičius and Nel 2002
G. †Palaeomacromia Petrulevičius et al. 1999
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†Palaeomacromia multicellulata Petrulevičius et al. 1999
Invalid names: Bolcathemidae Gentilini 2002 [synonym]
G. †Austrolibellula Petrulevičius and Nel 2003
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†Austrolibellula noroestenia Petrulevičius and Nel 2003
Fm. Corduliidae Selys-Longchamps 1850 [green-eyed skimmer]
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G. †Cellulocordulia Riou and Nel 1995
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†Cellulocordulia fasciata Riou and Nel 1995
Subfm. Corduliinae Selys-Longchamps 1850
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G. Cordulia Leach 1815 [emerald]
G. †Croatocordulia Kiauta 1969
+
†Croatocordulia platyptera Charpentier 1843
G. Epitheca Burmeister 1839
+
†Epitheca annae Gentilini 1992
†Epitheca miocenica Gentilini 1992
G. †Miocordulia Kennedy 1931
+
†Miocordulia latipennis Kennedy 1931
G. Somatochlora Selys-Longchamps 1871
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†Somatochlora brisaci Nel et al. 1996
†Somatochlora oregonica Cockerell 1927
G. †Molercordulia Bechly 2005
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†Molercordulia karinae Bechly 2005
G. †Stenogomphus Scudder 1892
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†Stenogomphus carletoni Scudder 1892
Invalid names: Gomphomacromiinae Tillyard and Fraser 1940 [empty]
G. †Jujusia Petrulevičius and Nel 2002
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†Jujusia maizgorda Petrulevičius and Nel 2002
Fm. Urothemistidae Lieftinck 1954
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G. †Eomacrodiplax Nel and Fleck 2014
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†Eomacrodiplax incompleta Nel and Fleck 2014
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitino
Form: roller-shapedo
Environment: freshwater, terrestrialo
Locomotion: fast-movingo
Life habit: volanto
Diet: carnivoreo
Diet 2: insectivoreo
Vision: well-developedo
Dispersal: watero
Created: 2011-03-07 03:41:10
Modified: 2011-03-06 10:41:10
Source: o = order
Reference: Ren and Guo 1996

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Gelasian or 93.90000 to 1.80000 Ma
Minimum age of oldest fossil (stem group age): 89.8 Ma

Collections (74 total)


Time interval Ma Country or state Original ID and collection number
Turonian93.9 - 89.8Kazakhstan Palaeolibellula zherikhini (119502)
Late/Upper Paleocene59.24 - 56.0Argentina (Jujuy) Austrolibellula noroestenia (106995) Curviarculia lamasi (114155) Jujusia maizgorda, Curviarculia delicata, Palaeomacromia multicellulata (113004)
Ypresian56.0 - 48.07Denmark Molercordulia karinae (105202) Molertrum eburneum (157040)
Ypresian56.0 - 48.07Italy Bolcathemis nervosa, Bolcacordulia paradoxa (122230)
Bridgerian50.5 - 46.2USA (Colorado) Stenogomphus carletoni (114681)
Bridgerian50.5 - 46.2USA (Wyoming) Libellulidae indet. (121322)
MP 1148.07 - 41.03Germany (Hessen) Corduliidae indet. (120798)
Uintan46.2 - 39.7USA (Colorado) Libellulidae indet. (156492)
Priabonian37.71 - 33.9United Kingdom (England) Corduliidae indet. A, Eomacrodiplax incompleta (123960) Corduliidae indet. B, Corduliidae indet. C, Eomacrodiplax incompleta (21976) Pantalinae indet. (207259)
Priabonian37.71 - 33.9United Kingdom (Isle of Wight) Corduliidae indet. A (157667)
Early/Lower Oligocene33.9 - 27.3USA (Oregon) Somatochlora oregonica (117157)
MP 2333.9 - 27.3France Tauriphila cerestensis (123913)
Rupelian33.9 - 27.3France (Auvergne) Libellula minuscula (127630)
Rupelian33.9 - 27.3France (Alpes-de-Haute) Paleotauriphila marquesi (155208) Paleotauriphila sp. (114016)
Chattian27.3 - 23.04France (Provence-Alpes-Côte d'Azur) Libellulidae larva, Libellula martini (122487)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Caussanelia papaziani, Palaeotramea aquisextana, Sympetrum bigoti (127415) Libellula requieniana, Libellula pourqueryi (153412) Libellulidae indet. (157194) Libellulidae larva (113712 122501) Paleotramea aquisextana (177171)
Chattian27.3 - 23.04Germany (North Rhine-Westphalia) Libellulidae sp. A, Palaeotramea cellulosa, Paleotramea cellulosa (122010)
Late/Upper Oligocene27.3 - 23.04France (Aix-en-Provence) Jeanlegrandia oligocenica (114117)
MP 2827.3 - 23.04Germany (Rheinland-Pfalz) Libellulidae indet. (71942)
MP 3027.3 - 23.04Germany Libellula cellulosa (123945)
MP 3027.3 - 23.04Germany (Nordrhein-Westfalen) Libellula cellulosa (138696)
Aquitanian23.04 - 20.45Germany Libellula sp., Trameobasileus moguntiacus (123872) Libellulidae indet. (237236)
Aquitanian23.04 - 20.45Türkiye Libellulidae indet., Trameini indet., Paleotramea aquisextana, Paleotramea fasciata, Paleotramea acutibucula (123307)
Early/Lower Miocene23.04 - 15.98Spain (Castellon) Libellulidae indet. (140796)
Burdigalian20.45 - 15.98Austria Libellulidae indet. (139942)
Burdigalian20.45 - 15.98Czechia Onychothemis rihai (181765)
Orleanian20.0 - 15.98Germany (Baden-Wurttemberg) Parabrachydiplax miocenica (106002)
Hemingfordian18.5 - 16.3USA (California) Orthemis ferruginea (106431) Orthemis sp. (123022 123023 123025 123042)
MN 416.9 - 15.98Czechia Libellulidae indet., Libellulidae indet. larvae, Libellula knetti (157202) Onychothemis rihai (105501) Prorhyothemis bubiki (105500)
Langhian15.98 - 13.82USA (Washington) Miocordulia latipennis (140652)
MN 515.98 - 13.82Germany (Baden-Württemberg) Randecktrum ebi (215189)
Middle Miocene15.98 - 11.63Australia (New South Wales) Corduliidae indet. (224044)
Middle Miocene15.98 - 11.63Russian Federation Pantalinae indet., Sympetrum sp. (105571)
Sarmatian12.8 - 11.63Slovenia Sloveniatrum robici (103103)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Cordulia sp. (115305) Libellula oeningensis (151877)
Sarmatian12.8 - 11.63Croatia Libellula platyptera (117246) Lithemis lejeunecarpentieri, Oligocaemia imperfecta (145135) Protopaltothemis hageni (113698)
Vallesian11.63 - 8.725Spain Trithemis pseudodistanti (114010)
Late/Upper Miocene11.63 - 5.333Ukraine (Kerch) Libellula ukrainensis (127948)
Late/Upper Miocene11.63 - 5.333Poland Libellula sieboldiana, Libellula kieseli, Libellula pannewitziana (138936)
Vallesian - Turolian11.63 - 5.09Spain (Lleida) Libellulidae indet. (154246)
Turolian8.725 - 5.09France (Ardèche) Corduliinae indet., Somatochlora brisaci (113893) Libellulidae indet., Miorhodopygia andancensis, Cellulocordulia fasciata (156164)
Messinian7.246 - 5.333Italy (Tuscany) Libellulidae indet., Diplax sp. (113751)
Messinian7.246 - 5.333Italy Corduliinae indet., Epitheca miocenica, Epitheca annae (113886) Libellula sp., Sympetrum italicum, Sympetrum krzeminskii, Sympetrum elongatum, Libellula mediterranea, Tramea miocenica, Pisaurum coloratum, Deielia sarae, Trapezostigma sp., Trapezostigma barbaresii, Libellula adriatica, Celithemis zavattinii, Sympetrum marinum (157306) Libellulidae indet. (157203)
Messinian7.246 - 5.333Italy (Piedmont) Oryctodiplax gypsorum (124409) Oryctodiplax sp., Oryctodiplax gypsorum (113998)
Messinian7.246 - 5.333France (Cantal) Celithemis cantalensis, Sympetrum cantalensis (122195)
Pliocene5.333 - 2.58India (Jharkhand) Libellulidae indet. (214996)
Blancan4.7 - 1.4USA (Texas) Libellulidae indet. (154909)
Piacenzian3.6 - 2.58USA (California) Libellulidae indet. (162798)
Gelasian2.58 - 1.8France Libellula eusebioi (113996)