PBDB Taxon

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Aeschnidiidae (small darner)

Insecta - Odonata - Aeschnidiidae

Taxonomy

Aeschnidiidae was named by Handlirsch (1906). Its type is Aeschnidium.

It was reranked as the subfamily Aeschnidiinae by Tillyard (1918).

It was assigned to Aeshnidae by Tillyard (1918); to Anisoptera by Carpenter (1992); to Aeschnidioidea by Carle and Wighton (1990), Zhang (1999); to Pananisoptera by Bechly (1998), Bechly (2000); and to Neoanisoptera by Bechly (2007).

Subtaxa

Aegyptidium, Aeschnidiella, Aeschnidiopsis, Aeschnidium (syn. Estemoa, Neimenggolgomphus) (type genus), Angloaeschnidium, Bergeriaeschnidia, Brunetaeschnidium, Brunneaeschnidia, Cooperaeschnidium, Coramaeschnidium, Cratoaeschnidium, Dakotaeschnidium, Delclosaeschidium, Diastatopaeschnidium, Dracontaeschnidium, Gansuaeschnidia, Gigantoaeschnidium, Hebeiaeschnidia, Iberoaeschnidium, Jarzembowskiaeschnidium, Kesseleraeschnidium, Kimmeridgebrachypteraeschnidium, Leptaeschnidium, Libellula abscissa, Linaeschnidium, Lithoaeschnidium, Lleidoaeschnidium, Malmaeschnidium, Misofaeschnidium, Nannoaeschnidium, Neimengogomphus, Neimengogomphus dongwugaiensis, Pseudosamarura, Pseudosamarura largina, Rossaeschnidium, Santanoptera, Sinaeschnidia, Sinostenophlebia, Solnhofenia, Sona, Stylaeschnidium, Tauropteryx, Urogomphus, Wightonia, Yixiangomphus (syn. Archaeogomphus), Yixiangomphus labius

Synonyms

Sonidae was named by Pritykina (1986). It is not extant. Its type is Sona.

It was synonymized subjectively with Aeschnidiidae by Sturm and Mendes (1998).

It was assigned to Aeshnoidea by Pritykina (1986).
Archaeogomphus was named by Lin (1976).

It was replaced with Yixiangomphus by Lin (1986), Nel and Paicheler (1994), Fleck et al. (2002).

It was assigned to Gomphidae by Lin (1976), Lin (1982); and to Gomphinae by Hong (1982).
Pseudosamarura was named by Lin (1976).

It was synonymized subjectively with Aeschnidium by Zhang (1999); it was considered a nomen dubium by Nel and Paicheler (1994), Fleck et al. (2002).

It was assigned to Gomphidae by Lin (1976).
Pseudosamarura largina was named by Lin (1976). It is an impression. Its type locality is AAI511, Dakangpu village, which is in a Barremian lacustrine - large shale in the Yixian Formation of China.

It was synonymized subjectively with Aeschnidium heishankowensis by Zhang (1999); it was considered a nomen dubium by Nel and Paicheler (1994), Fleck et al. (2002).
Hebeiaeschnidia was named by Hong (1982).

It was considered a nomen dubium by Nel and Martínez-Delclòs (1993).

It was assigned to Aeschnidiidae by Hong (1982).
Yixiangomphus was named by Lin (1986).

It was synonymized subjectively with Aeschnidium by Zhang (1999); it was considered a nomen dubium by Fleck et al. (2002).

It was assigned to Gomphidae by Lin (1986).
Libellula abscissa was named by Hagen (1862). It is an impression.

It was recombined as Urogomphus abscissus by Handlirsch (1906); it was recombined as Bergeriaeschnidia abscissa by Bechly (1998); it was considered a nomen dubium by Fleck and Nel (2003).
Neimengogomphus was named by Hong (1982).

It was corrected as Neimenggolgomphus by Nel and Paicheler (1994); it was synonymized subjectively with Aeschnidium by Zhang (1999); it was considered a nomen dubium by Fleck et al. (2002).

It was assigned to Gomphinae by Hong (1982); and to Gomphidae by Nel and Paicheler (1994).
Neimengogomphus dongwugaiensis was named by Hong (1982). It is an impression. Its type locality is Dongwugai, northwest of Shilanji, Urad Zhongqi, which is in an Aptian terrestrial horizon in the Huliugou Formation of China.

It was recombined as Neimenggolgomphus dongwugaiensis by Nel and Paicheler (1994); it was synonymized subjectively with Aeschnidium heishankowensis by Zhang (1999); it was considered a nomen dubium by Fleck et al. (2002).

Synonymy list

Year	Name and author
1862	Libellula abscissa Hagen p. 107
1890	Libellula abscissa Kirby p. 167
1906	Urogomphus abscissus Handlirsch p. 595
1906	Aeschnidiidae Handlirsch p. 609
1918	Aeschnidiinae Tillyard
1976	Archaeogomphus Lin p. 99
1976	Pseudosamarura Lin p. 100
1976	Pseudosamarura largina Lin p. 101 figs. pl. 2 figs 2,3
1980	Pseudosamarura largina Wang p. 134 figs. Pl 67, figs 2,3
1982	Archaeogomphus Hong p. 51
1982	Hebeiaeschnidia Hong p. 59
1982	Neimengogomphus Hong p. 86
1982	Neimengogomphus dongwugaiensis Hong pp. 86, 87 figs. Pl. 17 figs 1, 2 pl. 18 fig 1
1982	Archaeogomphus Lin p. 74
1986	Yixiangomphus Lin p. 449
1986	Sonidae Pritykina
1990	Aeschnidiidae Carle and Wighton p. 64
1992	Aeschnidiidae Carpenter p. 82
1994	Neimenggolgomphus Nel and Paicheler p. 69
1994	Neimenggolgomphus dongwugaiensis Nel and Paicheler p. 69
1998	Aeschnidiidae Bechly p. 3
1998	Bergeriaeschnidia abscissa Bechly p. 37 figs. 32-33
1999	Aeschnidiidae Zhang
2000	Aeschnidiidae Bechly p. 2
2007	Aeschnidiidae Bechly

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
infraclass	Hydropalaeoptera	Rohdendorf 1969
—	Euhydropalaeoptera

Rank	Name	Author
—	Odonatoptera	Martynov 1932
—	Palaeodonatoptera
—	Plesiodonatoptera
—	Apodonatoptera
—	Paneodonatoptera
—	Neodonatoptera
—	Euodonatoptera
—	Odonatoclada	Bechly 1999
—	Nodialata
—	Discoidalia
—	Stigmoptera
—	Lobodonata
—	Panodonata
order	Odonata
—	Epiproctophora
—	Euepiproctophora	Bechly 1998
—	Anisopteromorpha
—	Trigonoptera
—	Pananisoptera
—	Neoanisoptera	Bechly 1998
family	Aeschnidiidae
family	Aeschnidiidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Aeschnidiidae Handlirsch 1906 [small darner]

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G. †Aegyptidium Schlüter and Hartung 1982

†Aegyptidium aburasiensis Schlüter and Hartung 1982

G. †Aeschnidiella Zalessky 1953

†Aeschnidiella kabanovi Zalessky 1953

G. †Aeschnidiopsis Tillyard 1918

†Aeschnidiopsis flindersiensis Woodward 1884

G. †Aeschnidium Westwood 1854

†Aeschnidium antiquum Brodie 1845

†Aeschnidium bubas Westwood 1854

Invalid names: Estemoa Giebel 1856 [objective synonym], Neimenggolgomphus Hong 1982 [synonym]

G. †Angloaeschnidium Fleck and Nel 2003

†Angloaeschnidium lacaui Fleck and Nel 2003

†Angloaeschnidium montreuili Fleck and Nel 2003

†Angloaeschnidium toyei Fleck and Nel 2003

G. †Bergeriaeschnidia Nel et al. 1996

†Bergeriaeschnidia inexpectata Nel et al. 1996

G. †Brunetaeschnidium Fleck and Nel 2003

†Brunetaeschnidium nusplingensis Bechly 1998

G. †Brunneaeschnidia Hong 1982

†Brunneaeschnidia jiuquanensis Hong 1982

G. †Cooperaeschnidium Fleck and Nel 2003

†Cooperaeschnidium durandi Fleck and Nel 2003

G. †Coramaeschnidium Fleck and Nel 2003

†Coramaeschnidium minimum Fleck and Nel 2003

G. †Cratoaeschnidium Nel et al. 2022

†Cratoaeschnidium martinsnetoi Nel et al. 2022

G. †Dakotaeschnidium Nel 2021

†Dakotaeschnidium spedeni Nel 2021

G. †Delclosaeschidium Fleck and Nel 2003

†Delclosaeschidium magnum Fleck and Nel 2003

G. †Diastatopaeschnidium Fleck and Nel 2003

†Diastatopaeschnidium reneeheiko Fleck and Nel 2003

G. †Dracontaeschnidium Zhang and Zhang 2001

†Dracontaeschnidium orientale Zhang and Zhang 2001

G. †Gansuaeschnidia Zhang et al. 2022

†Gansuaeschnidia hongi Zhang et al. 2022

G. †Gigantoaeschnidium Nel and Martínez-Delclòs 1993

†Gigantoaeschnidium ibericus Nel and Martínez-Delclòs 1993

G. †Iberoaeschnidium Nel and Martínez-Delclòs 1993

†Iberoaeschnidium conquensis Nel and Martínez-Delclòs 1993

G. †Jarzembowskiaeschnidium Fleck and Nel 2003

†Jarzembowskiaeschnidium polandi Fleck and Nel 2003

G. †Kesseleraeschnidium Fleck and Nel 2003

†Kesseleraeschnidium simonae Fleck and Nel 2003

G. †Kimmeridgebrachypteraeschnidium Fleck and Nel 2003

†Kimmeridgebrachypteraeschnidium etchesi Fleck and Nel 2003

G. †Leptaeschnidium Pritykina 1977

†Leptaeschnidium latum Pritykina 1977

G. †Linaeschnidium Huang et al. 2009

†Linaeschnidium sinensis Huang et al. 2009

G. †Lithoaeschnidium Nel and Martínez-Delclòs 1993

†Lithoaeschnidium viohli Nel and Martínez-Delclòs 1993

G. †Lleidoaeschnidium Nel and Martínez-Delclòs 1993

†Lleidoaeschnidium maculatum Fleck and Nel 2003

†Lleidoaeschnidium valloryi Nel and Martínez-Delclòs 1993

G. †Malmaeschnidium Nel and Martínez-Delclòs 1993

†Malmaeschnidium mayeri Nel and Martínez-Delclòs 1993

G. †Misofaeschnidium Fleck and Nel 2003

†Misofaeschnidium densum Hagen 1862

G. †Nannoaeschnidium Nel and Martínez-Delclòs 1993

†Nannoaeschnidium pumilio Nel and Martínez-Delclòs 1993

G. †Rossaeschnidium Fleck and Nel 2003

†Rossaeschnidium patriciae Fleck and Nel 2003

G. †Santanoptera Martill and Nel 1996

†Santanoptera gabbotti Martill and Nel 1996

G. †Sinaeschnidia Hong 1965

†Sinaeschnidia heishankowense Hong 1965

†Sinaeschnidia huzhouensis Zhou and Wei 1980

†Sinaeschnidia martinezdelclosi Fleck and Nel 2003

G. †Sinostenophlebia Hong 1984

†Sinostenophlebia zhanjiakouensis Hong 1984

G. †Solnhofenia Bechly 2000

†Solnhofenia stoebeneri Bechly 2000

G. †Stylaeschnidium Zhang and Zhang 2001

†Stylaeschnidium rarum Zhang and Zhang 2001

G. †Tauropteryx Pritykina 1993

†Tauropteryx krassilovi Pritykina 1993

G. †Urogomphus Handlirsch 1906

†Urogomphus giganteus Germar 1839

Invalid names: Petalura eximia Hagen 1862 [synonym], Petalura latialata Hagen 1862 [objective synonym]

G. †Wightonia Carle 1990

†Wightonia araripina Carle and Wighton 1990

Invalid names: Hebeiaeschnidia Hong 1982 [nomen dubium], Libellula abscissa Hagen 1862 [nomen dubium], Neimengogomphus Hong 1982 [nomen dubium], Neimengogomphus dongwugaiensis Hong 1982 [nomen dubium], Pseudosamarura Lin 1976 [nomen dubium], Pseudosamarura largina Lin 1976 [nomen dubium], Sonidae Pritykina 1986 [synonym], Yixiangomphus Lin 1986 [nomen dubium], Yixiangomphus labius Lin 1976 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitino

Form:

roller-shapedo

Environment:

freshwater, terrestrialo

Locomotion:

fast-movingo

Life habit:

volanto

Diet:

carnivoreo

Diet 2:

insectivoreo

Vision:

well-developedo

Dispersal:

watero

Created:

2011-03-07 03:41:10

Modified:

2011-03-06 10:41:10

Source: o = order

Reference: Ren and Guo 1996

Age range: base of the Late/Upper Kimmeridgian to the top of the Chattian or 152.21000 to 23.04000 Ma

Collections (78 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Kimmeridgian	152.21 - 149.2	United Kingdom (England)	Kimmeridgebrachypteraeschnidium etchesi (106418)
Late/Upper Kimmeridgian	152.21 - 149.2	Germany (Swabian Alb)	Misofaeschnidium densum, Urogomphus giganteus (106402)
Early/Lower Tithonian	149.2 - 145.06	Germany (Bayern)	Aeschnidium densum (123144) Bergeriaeschnidia inexpectata (139550) Bergeriaeschnidia inexpectata, Urogomphus giganteus (122730) Estemoa densa (133578) Libellula densa, Aeschna gigantea, Petalura eximia, Lithoaeschnidium viohli, Brunetaeschnidium sp. (123985) Solnhofenia stoebeneri (152902) Urogomphus eximius (117531) Urogomphus giganteus (105798 131959 139551)
Early/Lower Tithonian	149.2 - 145.06	Germany	Urogomphus eximius, Urogomphus giganteus (122880) Urogomphus giganteus (151861 151862)
Early/Lower Tithonian	149.2 - 145.06	Germany (Bavaria)	Malmaeschnidium mayeri (152603) Urogomphus giganteus (122586 123994 139876)
Middle Berriasian	141.0 - 139.1	United Kingdom (England)	Aeschnidium bubas, Delclosaeschidium magnum (106401) Libellula antiqua (104008)
Late/Upper Berriasian	139.1 - 137.05	United Kingdom (England)	Aeschnidiidae indet., Delclosaeschidium magnum (104003) Aeschnidiidae indet., Diastatopaeschnidium reneeheiko, Jarzembowskiaeschnidium polandi (103833) Aeschnidiidae indet., Urogomphus sp., Jarzembowskiaeschnidium polandi (160681) Aeschnidium bubas (117342 153649 153650)
Late/Upper Hauterivian	132.6 - 125.77	United Kingdom (England)	Aeschnidiidae indet. (119697) Kesseleraeschnidium simonae, Coramaeschnidium minimum (106417)
Barremian	125.77 - 121.4	China (Hebei)	Sinostenophlebia zhanjiakouensis (155155)
Early/Lower Barremian	125.77 - 121.4	Spain	Aeschnidiidae indet. (123437) Lleidoaeschnidium valloryi (104052)
Early/Lower Barremian	125.77 - 121.4	United Kingdom (England)	Aeschnidiidae indet., Lleidoaeschnidium maculatum (45193) Angloaeschnidium lacaui (118921) Angloaeschnidium toyei, Cooperaeschnidium durandi, Rossaeschnidium patriciae (104005)
Late/Upper Barremian	125.77 - 121.4	China (Nei Mongol)	Sinaeschnidia heishankowensis (173105)
Late/Upper Barremian	125.77 - 121.4	Spain (Castilla-La Mancha)	Sinaeschnidia martinezdelclosi, Gigantoaeschnidium ibericus, Iberoaeschnidium conquensis (28006)
Late/Upper Barremian	125.77 - 121.4	China (Henan)	Sinaeschnidia heishankowense (229524)
Late/Upper Barremian	125.77 - 121.4	Spain (Cuenca)	Aeschnidiidae indet., Sinaeschnidia martinezdelclosi, Gigantoaeschnidium ibericus, Nannoaeschnidium pumilio, Iberoaeschnidium conquensis (152992)
Late/Upper Barremian	125.77 - 121.4	China (Liaoning)	Aeschnidiidae indet. (105809) Aeschnidium heishankowense (106315) Aeschnidium heishankowense, Stylaeschnidium rarum, Dracontaeschnidium orientale (111725) Archaeogomphus labius (146277) Linaeschnidium sinensis (152005) Pseudosamarura largina (151535) Sinaeschnidia sp. (105795)
Early/Lower Aptian	121.4 - 119.57	Russian Federation	Aeschnidiella kabanovi (160748)
Aptian	121.4 - 113.2	China (Liaoning)	Sinaeschnidia heishankowensis (151521 152382)
Aptian	121.4 - 113.2	Mongolia (Gobi-Altai aimak)	Sona nectes (104055)
Aptian	121.4 - 113.2	Russian Federation (Buryatia)	Leptaeschnidium latum (105536)
Aptian	121.4 - 113.2	China (Nei Mongol)	Neimengogomphus dongwugaiensis (165570)
Aptian	121.4 - 113.2	Australia (Queensland)	Aeschna flindersiensis (106404) Aeschnidiopsis flindersiensis (152158)
Aptian	121.4 - 113.2	China (Shandong)	Aeschnidium heishankowense (106313) Sinaeschnidia heishankowensis (154886)
Aptian	121.4 - 113.2	China (Gansu)	Aeschnidium heishankowense (106314) Archaeogomphus labius (154779) Archaeogomphus sp. (155009) Archaeogomphus sp., Brunneaeschnidia sp., Brunneaeschnidia jiuquanensis (155034) Gansuaeschnidia hongi (172664)
Aptian	121.4 - 113.2	Russian Federation	Aeschnidiella kabanovi (160749)
Aptian	121.4 - 113.2	Mongolia (Kobdo aimak)	Sona nectes (104054)
Aptian	121.4 - 113.2	China (Zhejiang)	Sinaeschnidia huzhouensis (156529)
Late/Upper Aptian	119.57 - 113.2	Brazil (Ceará)	Cratoaeschnidium martinsnetoi (217013) Santanoptera gabbotti (123012) Wightonia araripina (122352 122922 122926) Wightonia araripina, Santanoptera gabbotti (122557 122919)
Late/Upper Aptian	119.57 - 113.2	Egypt	Aegyptidium aburasiensis (106415)
Late/Upper Aptian	119.57 - 113.2	Colombia	Aeschnidiidae indet. (224197)
Albian	113.2 - 100.5	Australia (Queensland)	Aeschnidiidae indet. (106419)
Early/Lower Cenomanian	100.5 - 93.9	Ukraine (Bakhchisarayskiy)	Tauropteryx krassilovi (106283)
Late/Upper Maastrichtian	72.2 - 66.0	USA (North Dakota)	Aeschnidiidae indet. (229450)
Late/Upper Maastrichtian	72.2 - 66.0	USA (South Dakota)	Dakotaeschnidium spedeni (229449)
Chattian	27.3 - 23.04	Kazakhstan (Saisan)	Aeschnidiidae indet. (114018)