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Holcoptera schlotheimi

Insecta - Coleoptera - Coptoclavidae

Taxonomy
Harpalus schlotheimi was named by Giebel (1856). Its type specimen is NHMUK I.10783, an exoskeleton, and it is a compression fossil. Its type locality is Apperley, which is in a Rhaetian terrestrial horizon in the Lilstock Formation of the United Kingdom. It is the type species of Holcoptera.

It was recombined as Holcoelytrum schlotheimi by Zeuner (1962); it was recombined as Holcoptera schlotheimi by Handlirsch (1906), Cockerell (1915), Whalley (1985), Whalley (1986), Kelly et al. (2017).

Synonyms
Synonymy list
YearName and author
1856Harpalus schlotheimi Giebel p. 63
1906Holcoptera schlotheimi Handlirsch p. 453 figs. Pl XLI fig 63
1915Holcoptera confluens Cockerell p. 480 figs. Pl 61, fig 8
1915Holcoptera schlotheimi Cockerell p. 480 figs. Pl 61, fig 7
1962Holcoelytrum schlotheimi Zeuner
1985Holcoptera schlotheimi Whalley
1986Holcoptera schlotheimi Whalley p. 52
2017Holcoptera schlotheimi Kelly et al. p. 664 figs. 4-6

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
RankNameAuthor
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCoptoclavidaePonomarenko 1961
subfamilyCoptoclaviscinae
genusHolcoptera
speciesschlotheimi()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Holcoptera schlotheimi Giebel 1856
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Invalid names: Holcoptera confluens Cockerell 1915 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the Sevatian to the top of the Sinemurian or 211.18000 to 192.90000 Ma

Collections (15 total)


Time interval Ma Country or state Original ID and collection number
Sevatian211.18 - 205.7Italy (Lombardia) Holcoptera schlotheimi (94207)
Rhaetian205.7 - 201.4United Kingdom (England) Harpalinae indet. 1 (114643) Holcoptera schlotheimi (113961 114644 115223 115246 187761)
Rhaetian - Early/Lower Hettangian205.7 - 199.5United Kingdom (England) Holcoptera confluens (124688)
Early/Lower Hettangian201.4 - 199.5United Kingdom (England) Holcoptera schlotheimi (113956 113958 123986)
Planorbis201.4 - 199.5United Kingdom (England) Holcoptera schlotheimi (115460 123982)
Hettangian - Sinemurian201.4 - 192.9USA (Connecticut) Holcoptera schlotheimi (187762)
Turneri199.5 - 192.9United Kingdom (Dorset) Holcoptera schlotheimi (187763)