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Coccoidea (scale insect)

Insecta - Hemiptera

Synonyms
Synonymy list
YearName and author
1814Coccoidea Fallen
1843Orthezioidea Amyot and Audinet-Serville p. 619
1999Orthezioidea Koteja
2005Coccoidea Grimaldi and Engel p. 289
2008Orthezioidea Koteja and Azar p. 135
2008Coccoidea Koteja and Azar p. 158
2015Coccoidea Vea and Grimaldi
2018Coccoidea Johnson et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
RankNameAuthor
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Clareocercaria
—Acercaria()
superorderPancondylognatha
—Condylognatha
orderHemiptera
suborderSternorrhyncha()
superfamilyCoccoideaFallen 1814
superfamilyCoccoideaFallen 1814

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Coccoidea Fallen 1814 [scale insect]
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Fm. †Adocimycolidae Poinar and Vega 2023
+
G. †Adocimycolus Poinar and Vega 2023
+
†Adocimycolus aarondavisii Poinar and Vega 2023
G. †Alacrena Vea and Grimaldi 2015
+
†Alacrena peculiaris Vea and Grimaldi 2015
Fm. †Apticoccidae Vea and Grimaldi 2015
+
G. †Apticoccus Koteja and Azar 2008
+
†Apticoccus fortis Vea and Grimaldi 2015
†Apticoccus longitenuis Vea and Grimaldi 2015
†Apticoccus minutus Koteja and Azar 2008
Fm. †Arnoldidae Koteja 2008
+
G. †Arnoldus Koteja 2008
+
†Arnoldus capitatus Koteja 2008
†Arnoldus clavatus Koteja 2008
Fm. †Burmacoccidae Koteja 2004
+
G. †Burmacoccus Koteja 2004
+
†Burmacoccus danyi Koteja 2004
Fm. Coelostomidiidae Morrison 1928
+
G. †Cancerococcus Koteja 1988
+
†Cancerococcus apterus Koteja 1988
Fm. †Electrococcidae Koteja 2000
+
G. †Electrococcus Beardsley 1969
+
†Electrococcus canadensis Beardsley 1969
G. †Turonicoccus Koteja 2000
+
†Turonicoccus beardsleyi Koteja 2000
†Turonicoccus grimaldii Koteja 2000
Fm. †Grimaldiellidae Koteja 2000
+
G. †Grimaldiella Koteja 2000
+
†Grimaldiella gregaria Koteja 2000
†Grimaldiella resinophila Koteja 2000
Fm. †Grohnidae Koteja 2008
+
G. †Grohnus Koteja 2008
+
†Grohnus eichmanni Koteja 2008
Fm. †Jersicoccidae Koteja 2000
+
G. †Jersicoccus Koteja 2000
+
†Jersicoccus kurthi Koteja 2000
Fm. †Kozariidae Vea and Grimaldi 2015
+
G. †Kozarius Vea and Grimaldi 2015
+
†Kozarius achronus Vea and Grimaldi 2015
†Kozarius perpetuus Vea and Grimaldi 2015
Fm. †Kukaspididae Koteja and Poinar 2001
+
G. †Kukaspis Koteja and Poinar 2001
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†Kukaspis usingeri Koteja and Poinar 2001
Fm. Kuwaniidae MacGillvray 1921
+
G. †Hoffeinsia Koteja 2008
+
†Hoffeinsia foldii Koteja 2008
Fm. †Labiococcidae Koteja 2000
+
G. †Labiococcus Koteja 2000
+
†Labiococcus joosti Koteja 2000
G. †Solicoccus Koteja 2000
+
†Solicoccus nascimbenei Koteja 2000
Fm. †Lebanococcidae Koteja and Azar 2008
+
G. †Lebanococcus Koteja and Azar 2008
+
†Lebanococcus longiventris Koteja and Azar 2008
Fm. †Lithuanicoccidae Koteja 2008
+
G. †Lithuanicoccus Koteja 2008
+
†Lithuanicoccus damzeni Koteja 2008
Fm. †Macrodrilidae Poinar 2020
+
G. †Macrodrilus Poinar 2020
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†Macrodrilus bostrychus Poinar 2020
G. †Magnilens Vea and Grimaldi 2015
+
†Magnilens glaesaria Vea and Grimaldi 2015
Fm. Margarodidae Cockerell 1899
+
G. Heteromargarodes Jakubski 1965
+
†Heteromargarodes hukamsinghi Vea and Grimaldi 2015
Fm. Matsucoccidae Morrison 1927 [pine bark scales]
+
G. †Eomatsucoccus Koteja 1988
+
†Eomatsucoccus andrewi Koteja 1999
†Eomatsucoccus casei Koteja 2000
†Eomatsucoccus popovi Koteja 1988
†Eomatsucoccus sukachevae Koteja 1988
G. Matsucoccus Cockerell 1909
+
†Matsucoccus apterus Koteja 1984
†Matsucoccus crenata Koch and Berendt 1854
†Matsucoccus electrinus Koteja 1984
†Matsucoccus larssoni Koteja 1984
†Matsucoccus pinnatus Germar and Berendt 1856
†Matsucoccus saxonicus Koteja 1986
Invalid names: Acreagris Koch and Berendt 1854 [synonym]
Fm. Monophlebidae Signoret 1875 [giant scale]
+
G. Monophlebus Guerin-Meneville 1827
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†Monophlebus irregularis Germar and Berendt 1856
†Monophlebus simplex Scudder 1890
G. †Palaeophlebus Simon 2015
+
†Palaeophlebus kotejai Simon 2015
Fm. Ortheziidae Amyot and Audinet-Serville 1843 [ensign scale insect]
+
G. †Burmorthezia Vea and Grimaldi 2012
+
†Burmorthezia insolita Vea and Grimaldi 2012
†Burmorthezia kotejai Vea and Grimaldi 2012
G. †Cretorthezia Koteja and Azar 2008
+
†Cretorthezia hammanaica Koteja and Azar 2008
Subfm. Newsteadiinae Koteja 1974
+
Tr. Newsteadiini Koteja 1974
+
G. Mixorthezia Morrison 1925
+
†Mixorthezia dominicana Vea and Grimaldi 2012
†Mixorthezia kozari Vea and Grimaldi 2012
G. Newsteadia Green 1902
+
†Newsteadia succini Koteja and Zac-Ogaza 1988
G. Nipponorthezinella Kuwana 1916
Invalid names: Mixortheziini Kozár 2004 [synonym], Nipponortheziini Kozár 2004 [synonym]
Tr. Ortheziolini Kozár 2004
+
Invalid names: Matileortheziolini Kozár 2004 [synonym], Ortheziolacoccini Kozár 2004 [synonym], Ortheziolamametini Kozár 2004 [synonym]
Invalid names: Nipponortheziinae Kozár 2004 [synonym]
G. †Ochyrocoris Menge 1856
+
†Ochyrocoris electrina Menge 1856
Subfm. Ortheziinae Amyot and Audinet-Serville 1843
+
Tr. Ortheziini Amyot and Audinet-Serville 1843
+
G. Arctorthezia Cockerell 1912
+
†Arctorthezia antiqua Koteja and Zac-Ogaza 1988
†Arctorthezia baltica Vea and Grimaldi 2012
G. Orthezia Bosc d'Antic 1784
Invalid names: Arctortheziini Kozár 2004 [synonym]
G. †Palaeonewsteadia Koteja 1987
+
†Palaeonewsteadia huaniae Koteja 1987
G. †Protorthezia Koteja 1987
+
†Protorthezia aurea Koteja 1987
G. †Wathondara Simon et al. 2015
+
†Wathondara kotejai Simon et al. 2015
G. †Pedicellicoccus Vea and Grimaldi 2015
+
†Pedicellicoccus marginatus Vea and Grimaldi 2015
Fm. †Pennygullaniidae Koteja and Azar 2008
+
G. †Pennygullania Koteja and Azar 2008
+
†Pennygullania electrina Koteja and Azar 2008
Fm. Pityococcidae McKenzie 1942
+
G. Pityococcus McKenzie 1942
+
†Pityococcus moniliformalis Vea and Grimaldi 2015
Fm. Putoidae Beardsley 1969
+
G. †Palaeotupo Koteja and Azar 2008
+
†Palaeotupo danieleae Koteja and Azar 2008
G. Puto Signoret 1875
+
†Puto trivenosus Germar and Berendt 1856
Invalid names: Ceroputo Sulc 1898 [synonym]
G. †Xiphos Vea and Grimaldi 2015
+
†Xiphos vani Vea and Grimaldi 2015
Invalid names: Orthezioidea Amyot and Audinet-Serville 1843 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Hauterivian to the top of the Serravallian or 136.40000 to 11.62000 Ma
Minimum age of oldest fossil (stem group age): 130.0 Ma

Collections (42 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Hauterivian136.4 - 130.0United Kingdom (England) Eomatsucoccus andrewi (119697)
Early/Lower Barremian130.0 - 125.45Lebanon Apticoccus longitenuis (138801) Cretorthezia hammanaica, Hammanococcus sp., Hammanococcus setosus, Lebanococcus longiventris, Apticoccus minutus, Palaeotupo danieleae, Pennygullania sp., Pennygullania electrina, Apticoccus fortis, Xiphos vani (122471) Cretorthezia sp. (123000)
Aptian125.0 - 113.0Russian Federation (Buryatia) Eomatsucoccus popovi (113353 122549) Eomatsucoccus sukachevae (125413)
Albian113.0 - 100.5Jordan Margarodidae indet. (124847)
Cenomanian100.5 - 93.9USA (Alaska) Kukaspis usingeri (123051)
Early/Lower Cenomanian99.6 - 93.5Myanmar Burmacoccus danyi (123020) Burmorthezia sp., Burmorthezia kotejai, Burmorthezia insolita, Kozarius perpetuus, Kozarius achronus, Alacrena peculiaris, Magnilens glaesaria, Pedicellicoccus marginatus (111688) Wathondara kotejai (167760)
Early/Lower Cenomanian99.6 - 93.5Myanmar (Kachin) Macrodrilus bostrychus, Adocimycolus aarondavisii (104053)
Turonian93.9 - 89.8USA (New Jersey) Matsucoccidae indet., Jersicoccus kurthi, Grimaldiella sp., Grimaldiella resinophila, Grimaldiella gregaria, Eomatsucoccus casei, Turonicoccus beardsleyi, Turonicoccus grimaldii, Labiococcus joosti, Solicoccus nascimbenei (110895)
Late/Upper Campanian83.5 - 70.6Canada (Manitoba) Electrococcus canadensis (122354)
Ypresian56.0 - 47.8India (Gujarat) Heteromargarodes hukamsinghi (124463)
Priabonian38.0 - 33.9Poland Acreagris crenata, Monophlebus pinnatus, Monophlebus trivenosus, Monophlebus irregularis (109625) Arnoldus sp., Arnoldus capitatus, Arnoldus clavatus, Hoffeinsia foldii, Lithuanicoccus sp., Lithuanicoccus damzeni, Lithuanicoccus kosmowskae, Grohnus eichmanni, Palaeophlebus hoffeinorum, Palaeophlebus kotejai (165567) Hoffeinsia foldii (110737 120813) Ochyrocoris electrina, Monophlebus pinnatus (120811)
Priabonian38.0 - 33.9Russian Federation Hoffeinsia foldii, Lithuanicoccus sp., Lithuanicoccus damzeni, Palaeophlebus hoffeinorum (123419)
Priabonian38.0 - 33.9United Kingdom (England) Coccoidea indet. (207291)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Arctorthezia antiqua (123766) Arctorthezia baltica, Pityococcus moniliformalis (123493) Arctorthezia sp. (123773) Arnoldus capitatus (165568) Coccoidea indet. (201682) Lithuanicoccus damzeni, Grohnus eichmanni (110735) Matsucoccus pinnatus (123962) Matsucoccus pinnatus, Matsucoccus larssoni, Matsucoccus electrinus (123927) Newsteadia succini (124849) Protorthezia aurea (123420)
Priabonian38.0 - 33.9Denmark Matsucoccus sp., Matsucoccus pinnatus, Cancerococcus apterus, Matsucoccus larssoni, Matsucoccus electrinus, Matsucoccus apterus, Palaeonewsteadia huaniae (123762)
Priabonian38.0 - 33.9Germany (Halle) Hoffeinsia foldii (149491 165569) Hoffeinsia foldii, Lithuanicoccus damzeni (124273) Matsucoccus pinnatus, Matsucoccus saxonicus (129227)
Priabonian38.0 - 33.9Germany Hoffeinsia foldii (123769)
Chadronian37.2 - 33.9USA (Colorado) Monophlebus simplex (106384)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Mixorthezia kozari, Mixorthezia dominicana, Nipponorthezinella sp., Newsteadia sp. (122982)
Serravallian13.82 - 11.62India Coccoidea indet. (152647)