PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Heterogaster

Insecta - Hemiptera - Heterogastridae

Taxonomy

Heterogaster was named by Schilling (1829). It is extant. It is the type genus of Heterogasterinae.

It was assigned to Lygaeidae by Förster (1891); and to Heterogastridae by Gao et al. (2017).

Species

H. antiquus, H. breviscutatus, H. famosus, H. pumilio, H. radobojanus, H. redivivus, H. sundgoviensis, H. tristis, H. troglodytes

Synonyms

Heterogaster tristis was named by Heer (1864). It is a compression fossil. Its type locality is Oeningen (Zurich collection), which is in a Sarmatian crater lake limestone in the Upper Freshwater-Molasse Formation of Germany.

It was considered a nomen nudum by Heer (1864).

Synonymy list

Year	Name and author
1829	Heterogaster Schilling
1864	Heterogaster tristis Heer p. 392
1891	Heterogaster Förster p. 545
2017	Heterogaster Gao et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata

Rank	Name	Author
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Clareocercaria
—	Acercaria	()
superorder	Pancondylognatha
—	Condylognatha
order	Hemiptera
suborder	Heteroptera
infraorder	Pentatomomorpha	Leston et al. 1954
superfamily	Lygaeoidea	Schilling 1829
family	Heterogastridae	(Stal 1872)
genus	Heterogaster	Schilling 1829

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Heterogaster Schilling 1829

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†Heterogaster antiquus Heer 1853

†Heterogaster breviscutatus Theobald 1937

†Heterogaster famosus Förster 1891

†Heterogaster pumilio Heer 1853

†Heterogaster radobojanus Heer 1853

†Heterogaster redivivus Heer 1853

†Heterogaster sundgoviensis Theobald 1937

†Heterogaster troglodytes Heer 1853

Invalid names: Heterogaster tristis Heer 1864 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the MP 23 to the top of the Sarmatian or 33.90000 to 11.63000 Ma

Collections (14 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Oligocene	33.9 - 27.3	France	H. famosus, Lygaeus gracilentus, H. troglodytes (121473) H. sundgoviensis (122408)
MP 23	33.9 - 27.3	France	H. pumilio (122536) H. pumilio, H. breviscutatus (122535)
Rupelian	33.9 - 27.3	Germany	H. radobojanus, H. sundgoviensis (122407)
Chattian	27.3 - 23.04	France (Provence-Alpes-Côte d'Azur)	H. pumilio (122488)
Chattian	27.3 - 23.04	France (Bouches-Du-Rhone)	H. antiquus, H. pumilio (122501 122877) H. pumilio (113712) H. sp., H. antiquus (122511)
Sarmatian	12.8 - 11.63	Croatia	H. radobojanus (117246) H. redivivus (122874) H. troglodytes (117761)
Sarmatian	12.8 - 11.63	Germany (Baden-Württemberg)	H. tristis (117252)