PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Forcipomyia

Insecta - Diptera - Ceratopogonidae

Taxonomy

Forcipomyia was named by Meigen (1818). It is extant. It is the type genus of Forcipomyiinae, Forcipomyiini.

It was reranked as Ceratopogon (Forcipomyia) by Kieffer (1906).

It was assigned to Ceratopogon by Kieffer (1906); to Ceratopogonidae by Solórzano Kraemer (2007); to Forcipomyiinae by Szadziewski (1988), Szadziewski (1990), Borkent and Wirth (1997), Sontag and Szadziewski (2011), Borkent (2013), Pielowska et al. (2018); and to Forcipomyiini by Szadziewski and Grogan (1998), Borkent and Dominiak (2020), Szadziewski et al. (2021).

Species

F. (Euprojoannisia), F. (Forcipomyia), F. (Lasiohelea), F. (Lepidohelea), F. (Phytohelea), F. (Synthyridomyia), F. (Thyridomyia), F. (Trichohelea), F. eobreviflagellata, F. krzeminskii, F. kulickae, F. nadicola, F. orientalus

Synonyms

Aspinus was named by Hong (1981).

It was replaced with Succinaspinus by Hong (2002); it was replaced with Amberaspinus by Evenhuis (1994), Borkent et al. (2022).

It was assigned to Tendipedinae by Hong (1981).
Succinaspinus was named by Hong (2002).

It was synonymized objectively with Amberaspinus by Borkent et al. (2022).

It was assigned to Hamicaudini by Hong (2002), Hong (2002).
Amberaspinus was named by Evenhuis (1994).

It was synonymized subjectively with Forcipomyia by Szadziewski (2019), Borkent and Dominiak (2020), Borkent et al. (2022).

It was assigned to Chironomidae by Evenhuis (1994).

Synonymy list

Year	Name and author
1818	Forcipomyia Meigen p. 73
1906	Ceratopogon (Forcipomyia) Kieffer p. 52
1981	Aspinus Hong p. 52
1988	Forcipomyia Szadziewski p. 190
1990	Forcipomyia Szadziewski p. 78
1994	Amberaspinus Evenhuis
1997	Forcipomyia Borkent and Wirth p. 29
1998	Forcipomyia Szadziewski and Grogan p. 266
2002	Succinaspinus Hong p. 140
2007	Forcipomyia Solórzano Kraemer p. 54
2011	Forcipomyia Sontag and Szadziewski p. 793
2013	Forcipomyia Borkent p. 28
2018	Forcipomyia Pielowska et al. p. 241
2020	Forcipomyia Borkent and Dominiak p. 68
2021	Forcipomyia Szadziewski et al. p. 441

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta

Rank	Name	Author
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Panorpida
superorder	Antliophora
order	Diptera
infraorder	Culicomorpha
superfamily	Chironomoidea	Macquart 1838
family	Ceratopogonidae
subfamily	Forcipomyiinae	Lenz 1934
tribe	Forcipomyiini	Lenz 1934
genus	Forcipomyia

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Forcipomyia Meigen 1818

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Subg. Forcipomyia (Euprojoannisia) Brèthes 1914

†Forcipomyia (Euprojoannisia) berendti Szadziewski 1988

†Forcipomyia (Euprojoannisia) henningseni Szadziewski 1988

†Forcipomyia (Euprojoannisia) miocaenica Szadziewski 1993

†Forcipomyia (Euprojoannisia) piriformis Meunier 1904

†Forcipomyia (Euprojoannisia) tuberculosa Szadziewski 1993

Subg. Forcipomyia (Forcipomyia) Meigen 1818

†Forcipomyia (Forcipomyia) eobreviflagellata Szadziewski 1988

†Forcipomyia (Forcipomyia) eocostata Szadziewski 1988

†Forcipomyia (Forcipomyia) fusiparamera Szadziewski and Grogan 1998

†Forcipomyia (Forcipomyia) gedanicola Szadziewski 1988

†Forcipomyia (Forcipomyia) grimaldii Szadziewski and Grogan 1998

†Forcipomyia (Forcipomyia) krzeminskii Szadziewski 1988

†Forcipomyia (Forcipomyia) kulickae Szadziewski 1988

†Forcipomyia (Forcipomyia) lyneborgi Szadziewski 1988

†Forcipomyia (Forcipomyia) pseudomicrohelea Szadziewski 1988

†Forcipomyia (Forcipomyia) rasnitsyni Szadziewski et al. 2021

†Forcipomyia (Forcipomyia) subgedanicola Szadziewski 1993

†Forcipomyia (Forcipomyia) turbinata Meunier 1904

†Forcipomyia (Forcipomyia) uncula Meunier 1904

†Forcipomyia (Forcipomyia) unculiformis Szadziewski 1993

Subg. Forcipomyia (Lasiohelea) Kieffer 1921

†Forcipomyia (Lasiohelea) americana Szadziewski and Grogan 1998

†Forcipomyia (Lasiohelea) succinea Szadziewski 1988

†Forcipomyia (Lasiohelea) woodruffi Szadziewski and Grogan 1998

Subg. Forcipomyia (Lepidohelea) Kieffer 1917

†Forcipomyia (Lepidohelea) antilleana Szadziewski and Grogan 1998

†Forcipomyia (Lepidohelea) chrysosuccinea Szadziewski and Grogan 1998

†Forcipomyia (Lepidohelea) domibicolor Szadziewski and Grogan 1998

†Forcipomyia (Lepidohelea) lepidosuccinea Szadziewski and Grogan 1998

Subg. Forcipomyia (Phytohelea) Remm 1971

†Forcipomyia (Phytohelea) eophytoheleana Szadziewski 1988

Subg. Forcipomyia (Synthyridomyia) Saunders 1957

†Forcipomyia (Synthyridomyia) tertiaricola Szadziewski and Grogan 1998

Subg. Forcipomyia (Thyridomyia) Saunders 1925

†Forcipomyia (Thyridomyia) frutetosuccinea Szadziewski and Grogan 1998

Subg. Forcipomyia (Trichohelea) Goetghebuer 1920

†Forcipomyia (Trichohelea) bifidicola Szadziewski 1993

†Forcipomyia (Trichohelea) eotrichoheleana Szadziewski 1988

†Forcipomyia (Trichohelea) succinicola Szadziewski 1993

†Forcipomyia eobreviflagellata Szadziewski 1988

†Forcipomyia krzeminskii Szadziewski 1988

†Forcipomyia kulickae Szadziewski 1988

†Forcipomyia nadicola Szadziewski 2013

†Forcipomyia orientalus Hong 1981

Invalid names: Amberaspinus Evenhuis 1994 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Ypresian to the top of the Burdigalian or 56.00000 to 15.98000 Ma
Minimum age of oldest fossil (stem group age): 48.07 Ma

Collections (34 total)

Time interval	Ma	Country or state	Original ID and collection number
Ypresian	56.0 - 48.07	China (Liaoning)	Aspinus orientalus (124530)
Ypresian	56.0 - 48.07	India (Gujarat)	F. sp. (124463 183742 183743)
Lutetian	48.07 - 41.03	Russian Federation	F. nadicola (151918) F. sp. indet., F. sp., F. sp. (107340)
Priabonian	37.71 - 33.9	Germany (Halle)	F. sp., F. sp., F. miocaenica, F. tuberculosa, F. gedanicola, F. subgedanicola, F. unculiformis, F. succinicola (129227) F. sp., F. sp., F. sp., F. tuberculosa, F. bifidicola (129228)
Priabonian	37.71 - 33.9	Poland	F. sp., F. sp. (154301) F. spp., F. sp., F. sp. A, F. sp., F. turbinata, F. uncula, F. sp., F. piriformis, F. eocostata, F. gedanicola, F. krzeminskii, F. kulickae, F. eobreviflagellata (120813) F. spp., F. sp., F. uncula, F. berendti, F. gedanicola, F. eophytoheleana, F. eotrichoheleana (109625)
Priabonian	37.71 - 33.9	Denmark	F. spp., F. sp., F. sp., F. sp. B, F. uncula, F. piriformis, F. henningseni, F. gedanicola, F. lyneborgi, F. succinea, F. eophytoheleana, F. krzeminskii (123762)
Priabonian	37.71 - 33.9	Ukraine	F. sp. (113621 122996 122997)
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	Ceratopogon turbinatus, Ceratopogon unculus, Ceratopogon piriformis (123927) F. sp., F. uncula, F. pseudomicrohelea (123221) F. spp., F. sp., F. piriformis (123962) F. spp., F. sp., F. turbinata, F. uncula, F. krzeminskii (123993)
Early/Lower Miocene	23.04 - 15.98	Ethiopia	F. rasnitsyni (222682)
Early/Lower Miocene	23.04 - 15.98	Mexico (Chiapas)	F. sp. (106432 124467 124474 140215 140217 140833)
Early/Lower Miocene	23.04 - 15.98	Mexico	F. sp., F. sp. 1, F. sp. 2 (21977)
Burdigalian	20.45 - 15.98	New Zealand (Otago)	F. sp. (191650)
Burdigalian - Langhian	20.45 - 13.82	Dominican Republic	F. sp., F. antilleana, F. chrysosuccinea, F. domibicolor (125970) F. sp., F. sp. (123393) F. sp., F. sp., F. antilleana (140835) F. sp., F. sp., F. fusiparamera, F. grimaldii, F. woodruffi, F. sp., F. antilleana, F. domibicolor (123857) F. sp., F. sp., F. grimaldii, F. americana, F. woodruffi, F. sp., F. antilleana, F. domibicolor, F. lepidosuccinea, F. sp., F. frutetosuccinea, F. tertiaricola, F. sp. (122473) F. sp., F. sp., F. sp., F. grimaldii, F. woodruffi, F. sp., F. antilleana, F. domibicolor, F. sp. (122982)