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Systellognatha

Insecta - Plecoptera

Synonyms
Synonymy list
YearName and author
1802Perlomorpha Latreille p. 292
1969Systellognatha Enderlein
1987Perlomorpha Sinitshenkova p. 29
1992Perlomorpha Sinitshenkova p. 48
1998Perlomorpha Sinitshenkova p. 318
2000Systellognatha Zwick
2016Systellognatha Cui et al. p. 3
2018Systellognatha Sroka et al. p. 4

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
RankNameAuthor
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—PolyneopteraMartynov 1938
orderPlecoptera
suborderArctoperlariaZwick 1973
infraorderSystellognathaEnderlein 1969
infraorderSystellognathaEnderlein 1969

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infraor. Systellognatha Enderlein 1969
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Fm. †Crossoperlidae Chen 2024
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G. †Crossoperla Chen 2024
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Superfm. Perloidea Latreille 1802
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Fm. Chloroperlidae Okamoto 1912 [green stonefly]
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G. †Bestioperlisca Sinitshenkova 1990
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†Bestioperlisca inulata Sinitshenkova 1990
G. †Dipsoperla Sinitshenkova 1987
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†Dipsoperla kunikanensis Sinitshenkova 1990
†Dipsoperla serpentis Sinitshenkova 1987
Fm. Perlidae Latreille 1802 [common stonefly]
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Subfm. Acroneuriinae Klapalek 1914
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Tr. Acroneuriini Klapalek 1914
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G. †Euperlida Cifuentes-Ruiz 2007
Tr. Anacroneuriini Stark and Gaufin 1976
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G. †Dominiperla Stark and Lentz 1992
G. †Billoperla Chen and Xu 2021
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†Billoperla starki Chen and Xu 2021
G. †Burmaperla Jouault and Nel 2022
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†Burmaperla pouilloni Jouault and Nel 2022
G. †Burmesoperla Chen 2019
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†Burmesoperla expansa Chen 2019
G. †Cretacroneuria Chen 2020
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G. †Electroneuria Sroka et al. 2018
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†Electroneuria ronwoodi Sroka et al. 2018
Tr. †Largusoperlini Chen 2018
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G. †Largusoperla Chen et al. 2018
G. †Burmacroneuria Chen 2019
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G. †Burperla Chen 2020
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†Burperla decolorata Chen 2020
G. Perla Geoffroy 1762
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Perla burmeisteriana Classen 1936
†Perla prisca Pictet and Hagen 1856
G. †Pinguisoperla Chen 2018
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G. †Starkoperla Chen and Wang 2020
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†Starkoperla longusocollum Chen and Wang 2020
Invalid names: Neoperlini Stark and Caufin 1976 [empty]
Fm. Perlodidae Klapalek 1912 [perlodid stonefly]
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G. †Derancheperla Sinitshenkova 1990
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†Derancheperla collaris Sinitshenkova 1990
Subfm. Isoperlinae Frison 1942
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G. Isoperla Banks 1906
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†Isoperla baltica Jouault et al. 2021
†Isoperla succinica Hagen 1856
G. †Isoperlodes Sinitshenkova 1992
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†Isoperlodes perstrictus Sinitshenkova 1992
G. Perlodes Banks 1903
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†Perlodes resinatus Hagen 1856
Fm. †Petroperlidae Sroka et al. 2018
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G. †Branchioperla Sroka and Staniczek 2019
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†Branchioperla ianstewarti Sroka and Staniczek 2019
G. †Lapisperla Sroka et al. 2018
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†Lapisperla keithrichardsi Sroka et al. 2018
G. †Ovaloperla Chen and Xu 2022
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†Ovaloperla staniczeki Chen and Xu 2022
G. †Petroperla Sroka et al. 2018
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†Petroperla mickjaggeri Sroka et al. 2018
Superfm. Pteronarcyoidea Newman 1853
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Fm. †Cavoperlidae Chen 2023
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G. †Cavoperla Chen 2023
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†Cavoperla excavata Chen 2023
Fm. Peltoperlidae Claassen 1931
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Subfm. †Borisoperlinae Chen and Xu 2020
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G. †Borisoperla Chen and Xu 2020
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†Borisoperla kondratieffi Chen and Xu 2020
G. †Dewaltoperla Chen 2023
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†Dewaltoperla edwardi Chen 2023
G. †Zwickoperla Chen and Wang 2020
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†Zwickoperla brevicauda Chen and Wang 2020
Subfm. Peltoperlinae Claassen 1931
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G. †Siberiopelta Sinitshenkova and Yan 2024
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†Siberiopelta bashkuevi Sinitshenkova and Yan 2024
Fm. Pteronarcyidae Newman 1853 [salmonfly]
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G. †Pteroliriope Cui et al. 2016
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†Pteroliriope sinitshenkovae Cui et al. 2016
Invalid names: Perlomorpha Latreille 1802 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitino
Ontogeny: moltingo
Grouping: solitaryo
Environment: freshwater, terrestrialo
Locomotion: actively mobileo
Life habit: volanto
Diet: detritivoreo
Diet 2: omnivoreo
Vision: well-developedo
Reproduction: oviparouso
Created: 2011-07-02 09:44:25
Modified: 2011-07-01 18:44:25
Source: o = order
Reference: Tillyard 1921

Age range

Maximum range based only on fossils: base of the Kungurian to the top of the Piacenzian or 283.30000 to 2.58000 Ma
Minimum age of oldest fossil (stem group age): 274.4 Ma

Collections (92 total)


Time interval Ma Country or state Original ID and collection number
Kungurian283.3 - 274.4Russian Federation (Suksun) Tshekardoperla expulsa, Tshekardoperla depicta, Tshekardoperla squarrosa, Sylvoperlodes zhiltzovae (108206)
Roadian274.4 - 266.9Russian Federation (Kemerovo) Palaeoperla exacta, Palaeoperla prisca (104704 122568 122572) Palaeoperla prisca (122567 122569)
Wordian266.9 - 264.28Russian Federation (Udmurtia) Properla incrassata, Properla umbrosa, Kargaloperla grata, Kargaloperla fibrosa, Kargaloperla queata, Kargaloperla postica (104750)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Kargaloperla exuperata (104057) Palaeoperla perfracta (104717) Permoleuctropsis gracilis (104994)
Wuchiapingian259.51 - 254.14Russian Federation (Vologda) Issadoperla permiana, Properla issadensis, Kargaloperla decipiens, Kargaloperla furcata, Mirumoperla multinerva (104754)
Changhsingian254.14 - 251.902Kazakhstan (Zaisan) Kargaloperla avulsa (105062)
Changhsingian254.14 - 251.902Russian Federation Properla tungussica (151553) Tungussonympha meyeni (151565)
Ladinian241.464 - 237.0Kyrgyzstan (Osh) Mesoleuctra brachypoda (151569 151570)
Carnian237.0 - 227.3Argentina Platyperla marquati (151962)
Norian227.3 - 205.7Ukraine (Kharkiv) Trianguliperla innoxia, Berekia neglecta (105057)
Norian227.3 - 205.7Kazakhstan Trianguliperla aequalis (105055 114782 151554 151566)
Norian227.3 - 205.7China (Yunnan) Triassoperla yongrenensis (165249)
Norian227.3 - 205.7South Korea Platyperla sp. (191410)
Early/Lower Jurassic - Middle Jurassic201.4 - 161.5Kazakhstan Platyperla rigida (151561 151562 151563)
Pliensbachian192.9 - 184.2Russian Federation (Kemerovo) Mesoleuctra gigantea (117990 125341) Mesoleuctra gigantea, Mesoleuctra tibialis (125496)
Pliensbachian - Toarcian192.9 - 174.7Kazakhstan (Karagandy) Mesoleuctra gigantea (125495)
Toarcian184.2 - 174.7Russian Federation (Kemerovo) Platyperla caudiculata (125493)
Toarcian184.2 - 174.7Kyrgyzstan (Batken) Mesoleuctra quadrata, Mesoleuctra exserta (113381)
Toarcian184.2 - 174.7Russian Federation (Irkutsk) Mesoleuctra gracilis, Platyperla platypoda (114696) Perlisca aufuga (122950 125228 125494)
Aalenian174.7 - 170.9Mongolia Platyperla admissa (123958)
Bajocian - Bathonian170.9 - 165.3Mongolia Platyperla propera (151564)
Bajocian - Bathonian170.9 - 165.3Russian Federation (Krasnoyarsk) Platyperla conferta (118010)
Bathonian168.2 - 165.3Mongolia Mesoleuctroides latus, Trianguliperla orbiculata (119621)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Pteroliriope sinitshenkovae (113378 155751 172659)
Callovian - Oxfordian165.3 - 154.8China (Liaoning) Mesoleuctra peipiaoensis (156597) Platyperla platypoda (154823 173107)
Callovian - Oxfordian165.3 - 154.8China (Jilin) Platyperla platypoda (154827)
Oxfordian161.5 - 154.8Russian Federation Trianguliperla attenuata (124813)
Oxfordian161.5 - 154.8Russian Federation (Buryatia) Mesoleuctroides saturatus (123194 123195 125403 125497) Perlomimus multus (125229)
Oxfordian161.5 - 154.8Russian Federation (Chita) Mesoleuctroides saturatus (125499 125500 125501 125502)
Oxfordian161.5 - 154.8Russian Federation (Buryat) Mesoleuctroides saturatus (105017)
Oxfordian - Kimmeridgian161.5 - 149.2Russian Federation (Buryat) Chloroperloides fusiformis (104950)
Late/Upper Jurassic161.5 - 143.1China (Xinjiang) Sinoephemera kingi (124528)
Tithonian149.2 - 143.1Mongolia Bestioperlisca inulata, Trianguliperla optanda (121394) Platyperla mendosa, Trianguliperla optanda, Mesoleuctroides derelictus (104748)
Tithonian149.2 - 143.1Russian Federation (Chita) Dipsoperla serpentis (114307 123197) Pectinoperla notabilis (125407) Perlitodes aenigmaticus (119397) Savina laeta (140476)
Tithonian149.2 - 143.1Russian Federation Dipsoperla kunikanensis (124812)
Berriasian - Barremian143.1 - 121.4Russian Federation (Yakutiya) Isoperlodes perstrictus, Perlitodes borealis (110818)
Late/Upper Berriasian139.1 - 137.05United Kingdom (England) Ecdyoperla fairlightensis (123449)
Valanginian - Barremian137.05 - 121.4Mongolia (Dornogov) Trianguliperla quassa (113407)
Hauterivian132.6 - 125.77Russian Federation (Chita) Platyperla parricidalis (152177)
Barremian125.77 - 121.4Russian Federation (Chita) Trianguliperla limosa, Mesoleuctrisca pronotalis (125782) Trianguliperla limosa, Trianguliperla volucris, Mesoleuctrisca pronotalis (113599) Trianguliperla volucris (125784)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Archaeoperla rarissimus (140490) Sinosharaperla zhaoi (118075)
Early/Lower Aptian121.4 - 119.57Mongolia (Ara-Hangayn aymag) Derancheperla collaris (123464)
Aptian121.4 - 113.2Russian Federation (Buryatia) Perlomorpha indet. (113349) Siberiopelta bashkuevi (113350)
Early/Lower Cenomanian100.5 - 93.9Myanmar Acroneuriinae indet., Largusoperla sp., Petroperla mickjaggeri, Lapisperla keithrichardsi, Electroneuria ronwoodi, Largusoperla charliewattsi, Largusoperla billwymani, Largusoperla micktaylori, Largusoperla brianjonesi, Branchioperla ianstewarti (123053) Burmaperla pouilloni (212258) Largusoperla acus, Largusoperla flata, Largusoperla arcus, Largusoperla reni, Starkoperla longusocollum, Zwickoperla brevicauda, Borisoperla kondratieffi, Billoperla starki, Ovaloperla staniczeki (161726) Perlidae indet., Pinguisoperla yangzhouensis, Largusoperla crassus, Largusoperla difformitatem, Burmesoperla expansa, Largusoperla dewalti, Largusoperla borisi, Burmacroneuria projecta, Cretacroneuria angularis, Burperla decolorata, Largusoperla zhouchangfai, Dewaltoperla edwardi, Cavoperla excavata, Crossoperla teslenkoae (194479)
Priabonian37.71 - 33.9Poland Perla succinica, Perla resinata (120811) Perla succinica, Perla resinata, Perla prisca (109625)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Isoperla baltica (123936)
Oligocene33.9 - 23.04Mexico (Puebla) Euperlida parvicercifera (31519)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Dominiperla antigua (122473)
MN 320.0 - 16.9Czechia Perla burmeisteriana (105508)
Piacenzian3.6 - 2.58Japan Perlodidae indet. (188973)