PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Carupinae

Malacostraca - Decapoda - Portunidae

Taxonomy

Carupinae was named by Paulson (1875). It is extant. Its type is Carupa.

It was assigned to Portunidae by Karasawa et al. (2008), Schweitzer et al. (2010), Schweitzer et al. (2021).

Subtaxa

Atoportunus, Carupa (type genus), Catoptrus, Euronectes, Laleonectes, Libystes (syn. Carcinoplacoides), Neptocarcinus, Rakosia, Richerellus

Synonyms

Catoptrinae was named by Borradaile (1902). It is extant.

It was reranked as the family Catoptridae by Schweitzer et al. (2010); it was synonymized subjectively with Carupinae by Schweitzer et al. (2021).

It was assigned to Portunoidea by Schweitzer et al. (2010).
Atoportunini was named by Stevcic (2005). It is extant.

It was reranked as the subfamily Atoportuninae by Karasawa et al. (2008), Schweitzer et al. (2010); it was synonymized subjectively with Carupinae by Schweitzer et al. (2021).

It was assigned to Portunidae by Karasawa et al. (2008), Schweitzer et al. (2010).

Synonymy list

Year	Name and author
1875	Carupinae Paulson
1902	Catoptrinae Borradaile
2008	Atoportuninae Karasawa et al. p. 103
2008	Carupinae Karasawa et al. p. 103
2010	Catoptridae Schweitzer et al. p. 106
2010	Atoportuninae Schweitzer et al. p. 109
2010	Carupinae Schweitzer et al. p. 109
2021	Carupinae Schweitzer et al. p. 20

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Hymenostraca	(Rolfe 1969)
superclass	Multicrustacea	Regier et al. 2010
class	Malacostraca	()

Rank	Name	Author
subclass	Eumalacostraca	Grobben 1892
—	Caridoida	Hessler 1982
order	Decapoda
suborder	Pleocyemata	Burkenroad 1963
—	Reptantia	(Boas)
—	Meiura	Saint Laurent 1980
infraorder	Brachyura	()
—	Eubrachyura	de Saint Laurent 1980
—	Heterotremata	Guinot 1977
superfamily	Portunoidea	Rafinesque 1815
family	Portunidae	Rafinesque 1815
subfamily	Carupinae	Paulson 1875
subfamily	Carupinae	Paulson 1875

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Carupinae Paulson 1875

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G. Carupa Dana 1851

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Carupa tenuipes Dana 1852

G. Catoptrus Milne-Edwards 1870

G. †Euronectes Karasawa et al. 2008

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†Euronectes grumiensis Beschin et al. 2001

Euronectes vocans Müller 1993

G. Laleonectes Manning and Chace 1990

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Laleonectes vocans Milne-Edwards 1878

G. Libystes Milne Edwards 1867

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Libystes nitidus Milne-Edwards 1867

Invalid names: Carcinoplacoides Kesling 1958 [synonym]

G. †Neptocarcinus Lorenthey 1898

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†Neptocarcinus dezanchei Beschin et al. 2015

†Neptocarcinus millenaris Lörenthey 1898

†Neptocarcinus millenarusdup Lorenthey 1898

†Neptocarcinus spinosus Lorenthey 1929

G. †Rakosia Muller 1984

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†Rakosia carupoides Müller 1984

†Rakosia grumiensis Beschin et al. 2001

†Rakosia rectifrons Muller 1996

Rakosia vocans Müller 1993

G. Richerellus Crosnier 2003

Invalid names: Atoportuninae Stevcic 2005 [synonym], Catoptrinae Borradaile 1902 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Locomotion:

actively mobilec

Life habit:

semi-infaunalsuperf

Diet:

carnivoreo

Vision:

well-developedc

Created:

2022-11-16 17:54:45

Modified:

2022-11-16 17:54:45

Source: superf = superfamily, o = order, c = class

References: Fearon and Clapham 2023, Turnsek 1997, Aberhan 1992

Age range

Maximum range based only on fossils: base of the Lutetian to the top of the Late/Upper Pleistocene or 48.07000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 41.03 Ma

Collections (17 total)

Time interval	Ma	Country or state	Original ID and collection number
Paleogene	66.0 - 23.04	Hungary (Budapest)	Neptocarcinus millenaris (138811)
Lutetian	48.07 - 41.03	Italy (Vicenza)	Neptocarcinus millenaris (145128)
Lutetian	48.07 - 41.03	Italy	Neptocarcinus dezanchei (207556) Neptocarcinus millenaris (207553 226615)
Lutetian	48.07 - 41.03	Hungary	Neptocarcinus millenaris (207773)
Priabonian	37.71 - 33.9	Italy	Neptocarcinus millenaris (207550)
Early/Lower Oligocene	33.9 - 27.3	Italy (Vicenza)	Euronectes grumiensis (69701) Rakosia grumiensis (147005)
Rupelian	33.9 - 27.3	Italy (Veneto)	Euronectes grumiensis (91651)
Neogene	23.04 - 2.58	Poland	Rakosia rectifrons (139754)
Badenian	13.82 - 12.8	Hungary	Carupa tenuipes (207754) Rakosia carupoides (149366)
Serravallian	13.82 - 11.63	Hungary	Rakosia carupoides (34739)
Pleistocene	2.58 - 0.0117	Guam	Carcinoplacoides flottei (207017)
Middle Pleistocene	0.774 - 0.129	Barbados	Portunus vocans (185882)
Late/Upper Pleistocene	0.129 - 0.0117	Jamaica	Portunus vocans (185031)