PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Dorylinae (army ant)

Insecta - Hymenoptera - Formicidae

Taxonomy

Dorylinae was named by Leach (1815). It is extant.

It was assigned to Formicidae by Werner (1969), Hong (2002), Bolton (2003) and Borowiec (2016).

Subtaxa

Acanthostichini, Acanthostichus, Aenictus, Cerapachyini, Curticorna, Cylindromyrmecini, Cylindromyrmex, Dorylus (syn. Anomma, Dissimulodorylus), Eoaenictites, Lioponera (syn. Phyracaces), Neivamyrmex, Parasyscia, Procerapachys, Yunodorylus

Synonyms

Curticorna was named by Hong (2002).

It was considered a nomen nudum by Hong (2002) and Hong (2002).

It was assigned to Formicidae by Bolton (2012).
Curticorna leptogastrosa was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002) and Hong (2002).
Aenictinae was named by Emery (1901). It is extant.

It was synonymized subjectively with Dorylinae by Borowiec (2016).

It was assigned to Formicidae by Hong (2002) and Bolton (2003).
Eoaenictites was named by Hong (2002).

It was considered a nomen nudum by Hong (2002) and Hong (2002).

It was assigned to Formicidae by Bolton (2012).
Eoaenictites castanifurvus was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002) and Hong (2002).
Cerapachyini was named by Forel (1893). It is extant. Its type is Cerapachys.

It was considered an invalid subgroup of Dorylinae by Borowiec (2016).

It was assigned to Ponerinae by Wheeler (1915); and to Cerapachyinae by Bolton (2003) and Bolton (2012).
Cerapachyinae was named by Forel (1893). It is extant. Its type is Cerapachys.

It was synonymized subjectively with Dorylinae by Borowiec (2016).

It was assigned to Formicidae by Bolton (2003), Bolton (2012) and Dlussky et al. (2015).
Acanthostichini was named by Emery (1901). It is extant.

It was considered an invalid subgroup of Dorylinae by Borowiec (2016).

It was assigned to Cerapachyinae by Bolton (2003) and Bolton (2012).
Cylindromyrmecini was named by Emery (1901). It is extant.

It was considered an invalid subgroup of Dorylinae by Borowiec (2016).

It was assigned to Cerapachyinae by Bolton (2003) and Bolton (2012).

Synonymy list

Year	Name and author
1815	Dorylinae Leach
1893	Cerapachyinae Forel
1893	Cerapachyini Forel
1901	Acanthostichini Emery
1901	Aenictinae Emery
1901	Cylindromyrmecini Emery
1915	Cerapachyini Wheeler
1969	Dorylinae Werner p. 128
2002	Curticorna Hong p. 537
2002	Dorylinae Hong p. 537
2002	Curticorna leptogastrosa Hong p. 538 figs. 2.8.331-340
2002	Aenictinae Hong p. 541
2002	Eoaenictites Hong p. 541
2002	Eoaenictites castanifurvus Hong p. 542 figs. Pl 42, fig 2; text-fig 2.8.341-349
2003	Cerapachyinae Bolton p. 32
2003	Acanthostichini Bolton p. 33
2003	Cerapachyini Bolton p. 33
2003	Cylindromyrmecini Bolton p. 33
2003	Aenictinae Bolton p. 36
2003	Dorylinae Bolton p. 36
2012	Acanthostichini Bolton
2012	Cerapachyinae Bolton
2012	Cerapachyini Bolton
2012	Curticorna Bolton
2012	Curticorna leptogastrosa Bolton
2012	Cylindromyrmecini Bolton
2012	Eoaenictites Bolton
2012	Eoaenictites castanifurvus Bolton
2015	Cerapachyinae Dlussky et al. p. 140
2016	Dorylinae Borowiec p. 14

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()

Rank	Name	Author
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Hymenopterida
order	Hymenoptera
—	Unicalcarida
suborder	Apocrita	Gerstaecker 1867
—	Aculeata	()
superfamily	Formicoidea
family	Formicidae	()
subfamily	Dorylinae	Leach 1815
subfamily	Dorylinae	Leach 1815

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Dorylinae Leach 1815 [army ant]

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G. Acanthostichus Mayr 1887

†Acanthostichus hispaniolicus de Andrade 1998

G. Aenictus Shuckard 1840

G. Cylindromyrmex Mayr 1870

†Cylindromyrmex antillanus de Andrade 1998

†Cylindromyrmex electrinus de Andrade 1998

†Cylindromyrmex inopinatus de Andrade 2001

G. Dorylus Fabricius 1793 [driver ant]

Dorylus molestus Gerstäcker 1859

Invalid names: Anomma Shuckard 1840 [synonym], Dissimulodorylus Sosiak et al. 2022 [synonym], Dissimulodorylus perseus Sosiak et al. 2022 [invalid subgroup]

G. Lioponera Mayr 1879

Lioponera suscitata Viehmeyer 1913

Invalid names: Phyracaces Emery 1902 [synonym]

G. Neivamyrmex Borgmeier 1940

†Neivamyrmex ectopus Wilson 1985

Neivamyrmex iridescens Borgmeier 1950

G. Parasyscia Emery 1882

†Parasyscia conservata Viehmeyer 1913

G. †Procerapachys Wheeler 1915

†Procerapachys annosus Wheeler 1915

†Procerapachys favosus Wheeler 1915

†Procerapachys sulcatus Dlussky 2009

G. Yunodorylus Xu 2000

Invalid names: Acanthostichini Emery 1901 [invalid subgroup], Aenictinae Emery 1901 [synonym], Cerapachyinae Forel 1893 [synonym], Cerapachyini Forel 1893 [invalid subgroup], Curticorna Hong 2002 [nomen nudum], Cylindromyrmecini Emery 1901 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Ypresian to the top of the Holocene or 56.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 47.8 Ma

Collections (22 total)

Time interval	Ma	Country or state	Original ID and collection number
Ypresian	56.0 - 47.8	China (Liaoning)	Curticorna leptogastrosa, Eoaenictites castanifurvus (124820)
Priabonian	38.0 - 33.9	Poland	Procerapachys sulcatus (120813)
Priabonian	38.0 - 33.9	Russian Federation (Primorskiy)	Cerapachyinae indet. (172678)
Priabonian	38.0 - 33.9	Germany (Halle)	Procerapachys annosus (129227 129228)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Procerapachys annosus (110735 124073 124951) Procerapachys annosus, Procerapachys favosus (123927)
Rupelian	33.9 - 28.1	Germany	Lioponera sp. (122407)
Early/Lower Miocene	23.03 - 15.97	Mexico (Chiapas)	Neivamyrmex sp. (123456)
Burdigalian - Langhian	20.44 - 13.82	Dominican Republic	Acanthostichus hispaniolicus, Cylindromyrmex antillanus (122474) Cylindromyrmex electrinus, Neivamyrmex ectopus (21978) Cylindromyrmex inopinatus (122472) Neivamyrmex ectopus (222615)
Langhian	15.97 - 13.82	China (Fujian)	Cerapachys sp., Aenictus sp., Yunodorylus sp. (220273)
Pliocene - Holocene	5.333 - 0.0	Tanzania	Dorylus molestus (125729)
Blancan	4.9 - 1.8	USA (Texas)	Dorylinae indet. (154909)
Holocene	0.0117 - 0.0	Colombia (Santander)	Neivamyrmex iridescens (213295)
Holocene	0.0117 - 0.0	Tanzania	Dissimulodorylus perseus (229685) Dorylus molestus (234540)
Holocene	0.0117 - 0.0	Indonesia	Cerapachys conservatus, Phyracaces suscitatus (182105)