PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Empididae (dagger fly)

Insecta - Diptera - Empididae

Taxonomy

Empididae was named by Latreille (1804) [includes some dance flies]. It is extant. Its type is Empis.

It was corrected as Empedidae by Cockerell (1917).

It was assigned to Diptera by Cockerell (1917), Cockerell (1917), Cockerell (1921), Cockerell (1921); to Brachycera by Carpenter (1992); and to Empidoidea by Kovalev (1986), Sinclair and Cumming (2006), Wiegmann et al. (2011), Wahlberg and Johanson (2018).

Subtaxa

Acallomyia, Basilorhagio, Brachystomatinae, Burmitempis, Clinocerinae, Empidinae (syn. Hemerodromiinae), Hasmona, Hilarites, Oustaletimyia, Progloma, Protempidinae, Protoedalea, Tachydromia stilpon, Thirza

Synonyms

Brachystomatidae was named by Melander (1908). It is extant. Its type is Brachystoma.

It was synonymized subjectively with Empididae by Wahlberg and Johanson (2018).

It was assigned to Empidoidea by Sinclair and Cumming (2006), Sinclair et al. (2011).
Tachydromia stilpon was named by Meunier (1899). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (1994).

Synonymy list

Year	Name and author
1804	Empididae Latreille p. 310
1899	Tachydromia stilpon Meunier p. 178
1908	Brachystomatidae Melander
1917	Empedidae Cockerell p. 367
1921	Empididae Cockerell
1986	Empididae Kovalev p. 152
1992	Empididae Carpenter
2006	Empididae Sinclair and Cumming
2006	Brachystomatidae Sinclair and Cumming p. 80
2011	Brachystomatidae Sinclair et al.
2011	Empididae Wiegmann et al.
2018	Empididae Wahlberg and Johanson

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta

Rank	Name	Author
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Panorpida
superorder	Antliophora
order	Diptera
—	Neodiptera	Michelsen 1994
suborder	Brachycera	Zetterstedt 1842
—	Heterodactyla
—	Eremoneura
superfamily	Empidoidea	Latreille 1804
family	Empididae	Latreille 1804
family	Empididae	Latreille 1804

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Empididae Latreille 1804 [dagger fly]

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G. Acallomyia Melander 1928

†Acallomyia probolaea Melander 1949

G. †Basilorhagio Ren 1995

†Basilorhagio venustus Ren 1995

Subfm. Brachystomatinae Melander 1908

Tr. Brachystomatini Melander 1908

G. Brachystoma Meigen 1822

Invalid names: Brachystoma spinulosa Loew 1850 [nomen nudum]

Tr. Trichopezini Vaillant 1981

G. Apalocnemis Philippi 1865

†Apalocnemis canadambris Grimaldi and Cumming 1999

†Apalocnemis gracilis Meunier 1908

Invalid names: Timalphes Melander 1928 [synonym]

G. Gloma Meigen 1822

†Gloma hirta Loew 1850

Invalid names: Gloma acuticornis Loew 1850 [nomen nudum], Gloma palpata Loew 1850 [nomen nudum]

G. Heleodromia Haliday 1833

Subg. Heleodromia (Neoilliesiella) Wagner 2006

†Heleodromia (Neoilliesiella) foveata Wagner et al. 2000

Invalid names: Heleodromia (Illiesiella) Wagner 1985 [replaced]

†Heleodromia delicata Meunier 1908

†Heleodromia dolosa Meunier 1908

†Heleodromia starki Hoffeins et al. 1997

G. †Palaeoparamesia Meunier 1902

†Palaeoparamesia proosti Meunier 1902

G. Trichopeza Rondani 1856

†Trichopeza sucina Melander 1928

Invalid names: Trichopeza longicornis Meunier 1908 [replaced]

G. †Burmitempis Cockerell 1917

†Burmitempis halteralis Cockerell 1917

Subfm. Clinocerinae Schiner 1862

G. Proclinopyga Melander 1928

†Proclinopyga ulrichi Sinclair 2010

Invalid names: Fushunempites Hong 2002 [nomen nudum]

Subfm. Empidinae Latreille 1804

Tr. Chelipodini Hendel 1936

G. Chelipoda Macquart 1823

†Chelipoda vaga Meunier 1908

Invalid names: Lepidomya Bigot 1857 [synonym]

G. Phyllodromia Zetterstedt 1837

†Phyllodromia inclusa Shelford 1911

Tr. Empidini Latreille 1804

G. Empis Linnaeus 1758

Subg. Empis (Acallomyia) Melander 1928

†Empis carbonum Germar 1837

†Empis exilis Meunier 1908

†Empis florissantana Cockerell 1915

†Empis infossa Melander 1949

†Empis inscita Meunier 1908

†Empis macilenta Meunier 1908

†Empis macrophthalma Förster 1891

†Empis mala Meunier 1908

†Empis malefica Meunier 1908

†Empis melia von Heyden 1870

†Empis miocenica Cockerell 1915

†Empis mordax Meunier 1908

†Empis morosella Melander 1928

Invalid names: Empis morosa Meunier 1908 [replaced]

†Empis orapaensis Waters 1989

†Empis perdita Cockerell 1916

†Empis personata Meunier 1908

†Empis poeppigi Giebel 1856

†Empis pulvillata Loew 1850

†Empis spinifera Statz 1940

†Empis tritava Melander 1928

Invalid names: Empis tristis Meunier 1908 [replaced]

†Empis umbonata Statz 1940

Invalid names: Empis bulbirostris Meunier 1899 [nomen nudum], Empis stilicornis Meunier 1899 [nomen nudum], Empis tibialis Meunier 1899 [nomen nudum]

G. Rhamphomyia Meigen 1822

†Rhamphomyia ablata Meunier 1908

†Rhamphomyia aeterna Melander 1949

†Rhamphomyia angusta Meunier 1908

†Rhamphomyia corrupta Meunier 1908

†Rhamphomyia craterae Melander 1949

†Rhamphomyia crinitarsis Loew 1850

†Rhamphomyia distans Loew 1850

†Rhamphomyia enena Cockerell 1921

†Rhamphomyia errabunda Meunier 1908

†Rhamphomyia fossa Melander 1949

†Rhamphomyia fulvescens Statz 1940

†Rhamphomyia hypolitha Cockerell 1917

†Rhamphomyia inanimata Melander 1949

†Rhamphomyia infernalis Melander 1949

†Rhamphomyia insolita Meunier 1908

†Rhamphomyia interita Melander 1949

†Rhamphomyia involuta Meunier 1908

†Rhamphomyia latipennata Statz 1940

†Rhamphomyia media Meunier 1908

†Rhamphomyia morticina Melander 1949

†Rhamphomyia obtusa Meunier 1908

†Rhamphomyia oedaloides Meunier 1908

†Rhamphomyia polymorpha Meunier 1899

†Rhamphomyia porrecta Meunier 1908

†Rhamphomyia rottensis Statz 1940

†Rhamphomyia senecta Melander 1949

†Rhamphomyia sepulta Cockerell 1916

†Rhamphomyia spodites Melander 1949

†Rhamphomyia tumulata Melander 1949

†Rhamphomyia ungulina Loew 1850

Invalid names: Rhamphomyia antipedalis Loew 1850 [nomen nudum], Rhamphomyia formosa Loew 1850 [nomen nudum], Rhamphomyia pteropa Loew 1850 [nomen nudum], Rhamphomyia ptilopa Loew 1850 [nomen nudum], Rhamphomyia remitarsis Loew 1850 [nomen nudum]

G. †Emplita Grimaldi and Cumming 1999

†Emplita casei Grimaldi and Cumming 1999

Tr. Hemerodromiini Schiner 1862

G. Chelifera Macquart 1823

†Chelifera detestata Meunier 1908

G. Hemerodromia Meigen 1822

†Hemerodromia grootaerti Plant and Shamshev 2011

Tr. Hilarini Collin 1961

G. Hilara Meigen 1822

†Hilara heerii Meunier 1915

†Hilara litigiosa Meunier 1908

†Hilara royoi Gil Collado 1926

†Hilara tarda Meunier 1908

G. Hilarempis Bezzi 1905

†Hilarempis otagoensis Kaulfuss and Sinclair 2024

G. †Turonempis Grimaldi and Cumming 1999

†Turonempis styx Grimaldi and Cumming 1999

Invalid names: Hemerodromiinae Schiner 1862 [synonym]

G. †Hasmona Giebel 1856

†Hasmona leo Giebel 1856

G. †Hilarites Heer 1856

†Hilarites bellus Heer 1856

G. †Oustaletimyia Meunier 1893

†Oustaletimyia succinorum Meunier 1893

G. †Progloma James 1937

†Progloma rohweri James 1937

Subfm. †Protempidinae Ussatchov 1968

G. †Homalocnemimus Mostovski 1999

†Homalocnemimus abjugatus Mostovski 1999

G. †Protempis Ussatchov 1968

†Protempis antennata Ussatchov 1968

†Protempis minuta Ren 1998

G. †Protoreogeton Mostovski 1999

†Protoreogeton admirabilis Mostovski 1999

G. †Protoedalea Cockerell 1921

†Protoedalea brachystoma Cockerell 1921

G. †Thirza Giebel 1856

†Thirza naumanni Giebel 1856

Invalid names: Brachystomatidae Melander 1908 [synonym], Tachydromia stilpon Meunier 1899 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Bajocian to the top of the Holocene or 170.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 165.3 Ma

Collections (46 total)

Time interval	Ma	Country or state	Original ID and collection number
Bajocian - Bathonian	170.9 - 165.3	Mongolia (Bayan-Khongor)	Protoreogeton admirabilis (114615)
Callovian - Oxfordian	165.3 - 154.8	Kazakhstan (Chimkent)	Homalocnemimus abjugatus (105970) Homalocnemimus abjugatus, Protempis antennata (106101) Homalocnemimus sp. (105971) Homalocnemimus sp., Homalocnemimus abjugatus (105426)
Middle Berriasian	141.0 - 139.1	United Kingdom (England)	Empididae indet. (104008)
Late/Upper Barremian	125.77 - 121.4	China (Hebei)	Basilorhagio venustus (107001)
Late/Upper Barremian	125.77 - 121.4	China (Liaoning)	Protempis minuta (138436)
Aptian	121.4 - 113.2	Russian Federation (Buryatia)	Empididae indet. (113349)
Aptian	121.4 - 113.2	Mongolia (Govi-Altai)	Empididae indet. (122332)
Early/Lower Cenomanian	100.5 - 93.9	Myanmar	Burmitempis halteralis (123020)
Turonian	93.9 - 89.8	USA (New Jersey)	Turonempis styx, Emplita casei (110895)
Turonian	93.9 - 89.8	Botswana	Empis orapaensis (123430)
Late/Upper Campanian	83.6 - 72.2	Canada (Manitoba)	Apalocnemis canadambris (122354)
Early/Lower Eocene	56.0 - 48.07	USA (Colorado)	Protoedalea brachystoma (121194)
Ypresian	56.0 - 48.07	China (Liaoning)	Fushunempites furvis (124820)
Ypresian	56.0 - 48.07	USA (Colorado)	Rhamphomyia enena (105004)
Priabonian	37.71 - 33.9	Ukraine	Hemerodromia grootaerti (122997)
Priabonian	37.71 - 33.9	Poland	Rhamphomyia ungulina, Empis pulvillata, Gloma hirta, Rhamphomyia distans (109625)
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	Empis poeppigi, Thirza naumanni (147344) Heleodromia starki (154618) Oustaletimyia succinorum, Hemerodromia detestata, Phyllodromia dolosa, Phyllodromia delicata, Lepidomyia vaga, Parathalassiella problematica, Palaeoparamesia proosti, Trichopeza longicornis, Brachystoma gracilis, Rhamphomyia corrupta, Rhamphomyia ablata, Rhamphomyia involuta, Rhamphomyia ungulina, Rhamphomyia crinitarsis, Rhamphomyia errabunda, Rhamphomyia obtusa, Rhamphomyia insolita, Rhamphomyia angusta, Rhamphomyia media, Hilara litigiosa, Hilara tarda, Hilara macilenta, Empis exilis, Empis inscita, Empis personata, Empis mordax, Empis tristis, Empis pulvillata, Empis morosa, Brachystoma spinulosa, Rhamphomyia polymorpha, Tachydromia stilpon (123927) Phyllodromia sp., Rhamphomyia ablata, Rhamphomyia ungulina, Rhamphomyia crinitarsis, Rhamphomyia errabunda, Rhamphomyia porrecta, Rhamphomyia oedaloides, Rhamphomyia insolita, Rhamphomyia media, Empis malefica, Empis mala, Empis morosa (124073) Proclinopyga ulrichi (125512)
Priabonian	37.71 - 33.9	Germany (Halle)	Heleodromia starki (124273)
Priabonian	37.71 - 33.9	Russian Federation	Heleodromia starki, Heleodromia foveata (123419)
Chadronian	37.0 - 33.9	USA (Colorado)	Acallomyia probolaea, Rhamphomyia aeterna, Rhamphomyia interita, Rhamphomyia spodites, Rhamphomyia tumulata (106381) Empis miocenica, Empis florissantana (113749) Empis perdita (124692) Progloma rohweri, Rhamphomyia sepulta (106379) Rhamphomyia hypolitha, Empis infossa, Rhamphomyia craterae, Rhamphomyia fossa, Rhamphomyia inanimata, Rhamphomyia infernalis (124624) Rhamphomyia morticina (124680) Rhamphomyia senecta (123119)
Early/Lower Oligocene	33.9 - 27.3	France	Empis macrophthalma (121473)
MP 23	33.9 - 27.3	France	Hilara heerii (122535)
Chattian	27.3 - 23.04	France (Bouches-Du-Rhone)	Hilara heerii (113712) Hilarites bellus (122876)
Chattian	27.3 - 23.04	Germany (North Rhine-Westphalia)	Empis spinifera, Empis umbonata, Rhamphomyia fulvescens, Rhamphomyia latipennata, Rhamphomyia rottensis (122010)
MP 30	27.3 - 23.04	Germany	Empis melia (123945)
MP 30	27.3 - 23.04	Germany (Nordrhein-Westfalen)	Empis melia (138696)
Aquitanian	23.04 - 20.45	New Zealand	Hilarempis otagoensis (50035)
Early/Lower Miocene	23.04 - 15.98	Spain (Castellon)	Hilara royoi (122583 152716)
Early/Lower Miocene	23.04 - 15.98	Japan	Empididae indet. (114003)
Late/Upper Miocene	11.63 - 5.333	Germany	Empis carbonum (117341)
Holocene	0.0117 - 0.0	Tanzania	Phyllodromia inclusa (123991)