PBDB Taxon

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Pimplinae

Insecta - Hymenoptera - Ichneumonidae

Taxonomy

Pimplinae was named by Wesmael (1845). It is extant. Its type is Pimpla.

It was assigned to Ichneumonidae by Handlirsch (1910), Meunier (1923), Statz (1936), Kasparyan and Rasnitsyn (1992), Khalaim (2014), Kopylov et al. (2018), Spasojevic et al. (2018) and Spasojevic et al. (2018); and to Pimpliformes by Klopfstein et al. (2019) and Klopfstein and Spasojevic (2019).

Subtaxa

Crusopimpla, Ephialtini, Firkantus, Ichninsum, Itoplectis saxosa, Lithoserix, Pimplini, Pimplites, Polysphinctini, Theroniini

Synonyms

Itoplectis saxosus was named by Cockerell (1921). Its type specimen is BMNH H. 183, a forewing, and it is a compression fossil. Its type locality is Bembridge Marls (Hooley coll), which is in a Priabonian lagoonal/restricted shallow subtidal lime mudstone in the Bouldnor Formation of the United Kingdom.

It was considered a nomen dubium by Khalaim (2014).
Pimploidae was named by Förster (1869). It is extant.

It was synonymized objectively with Pimplinae by Wahl (2014).

It was assigned to Ichneumonoidea by Förster (1869).

Synonymy list

Year	Name and author
1845	Pimplinae Wesmael
1869	Pimploidae Förster p. 162
1910	Pimplinae Handlirsch p. 99
1921	Itoplectis saxosus Cockerell pp. 7-8 fig. 5
1923	Pimplinae Meunier p. 84
1936	Pimplinae Statz p. 270
1992	Pimplinae Kasparyan and Rasnitsyn
2014	Pimplinae Khalaim p. 344
2018	Pimplinae Kopylov et al. p. 323
2018	Pimplinae Spasojevic et al. p. 45
2018	Pimplinae Spasojevic et al. p. 4
2019	Pimplinae Klopfstein et al.
2019	Pimplinae Klopfstein and Spasojevic

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()

Rank	Name	Author
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Hymenopterida
order	Hymenoptera
—	Unicalcarida
suborder	Apocrita	Gerstaecker 1867
superfamily	Ichneumonoidea
family	Ichneumonidae	()
—	Pimpliformes	Wahl 1990
subfamily	Pimplinae	Wesmael 1845
subfamily	Pimplinae	Wesmael 1845

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Pimplinae Wesmael 1845

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G. †Crusopimpla Kopylov et al. 2018

†Crusopimpla collina Klopfstein 2022

†Crusopimpla elongata Klopfstein 2022

†Crusopimpla klopfsteinae Manukyan 2023

†Crusopimpla minuta Klopfstein 2022

†Crusopimpla rediviva Brues 1910

†Crusopimpla rettigi Klopfstein 2022

†Crusopimpla tadushiensis Kopylov et al. 2018

†Crusopimpla violina Klopfstein 2022

†Crusopimpla weltii Viertler et al. 2022

Tr. Ephialtini Hellén 1915

G. Dolichomitus Smith 1877

†Dolichomitus saxeus Kopylov et al. 2018

G. Exeristes Förster 1869

†Exeristes gurnetor Khalaim 2014

G. †Polyhelictes Spasojevic et al. 2018

†Polyhelictes bipolarus Spasojevic et al. 2018

G. Scambus Hartig 1838

†Scambus fossilis Khalaim 2014

†Scambus fossilobus Spasojevic et al. 2018

†Scambus mandibularis Spasojevic et al. 2018

†Scambus parachuti Spasojevic et al. 2018

G. Zabrachypus Cushman 1920

†Zabrachypus tumidus Khalaim 2008

G. †Firkantus Viertler et al. 2023

†Firkantus freddykruegeri Viertler et al. 2023

G. †Ichninsum Spasojevic et al. 2018

†Ichninsum appendicrassum Spasojevic et al. 2018

G. †Lithoserix Brown 1986

†Lithoserix antiquus Saussure 1852

†Lithoserix oublierus Viertler 2024

†Lithoserix williamsi Brown 1986

Tr. Pimplini Wesmael 1845

G. Itoplectis Förster 1869

G. Pimpla Fabricius 1804

†Pimpla amplifemura Lin 1988

†Pimpla anomalensis Theobald 1937

†Pimpla antiquus Saussure 1852

Invalid names: Ichneumon sublongaevus Meunier 1914 [synonym]

†Pimpla appendigera Brues 1906

†Pimpla aquensis Theobald 1937

†Pimpla bibosa Khalaim 2008

†Pimpla cyclostigmata Statz 1936

†Pimpla decessa Scudder 1877

†Pimpla eocenica Cockerell 1919

†Pimpla impuncta Lin 1988

†Pimpla morleyi Meunier 1923

†Pimpla renevieri Meunier 1903

†Pimpla saussurei Heer 1856

†Pimpla saxea Scudder 1877

†Pimpla senecta Scudder 1877

†Pimpla seyrigi Theobald 1937

†Pimpla succini Giebel 1856

G. Xanthopimpla Saussure 1892

†Xanthopimpla biamosa Khalaim 2008

†Xanthopimpla ciboisae Klopfstein 2022

†Xanthopimpla crescendae Klopfstein 2022

†Xanthopimpla messelensis Spasojevic et al. 2018

†Xanthopimpla praeclara Spasojevic et al. 2018

G. †Pimplites Steinbach 1967

†Pimplites praeparatus Steinbach 1967

Tr. Polysphinctini Hellén 1915

G. Polysphincta Gravenhorst 1829

†Polysphincta inundata Brues 1910

†Polysphincta statzi Meunier 1923

Tr. Theroniini Cushman and Rohwer 1921

G. Epitheronia Gupta 1962

†Epitheronia stigmatica Henriksen 1922

G. Theronia Holmgren 1859

†Theronia furensis Klopfstein 2022

†Theronia nigriscutum Klopfstein 2022

Invalid names: Itoplectis saxosa Cockerell 1921 [nomen dubium], Pimploidae Förster 1869 [objective synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the MP 11 to the top of the Chibanian or 48.60000 to 0.12900 Ma
Minimum age of oldest fossil (stem group age): 47.8 Ma

Collections (55 total)

Time interval	Ma	Country or state	Original ID and collection number
Ypresian	56.0 - 47.8	Russian Federation (Primorsky Krai)	Crusopimpla tadushiensis, Dolichomitus saxeus (123616)
Ypresian	56.0 - 47.8	Denmark	Crusopimpla collina (226442) Crusopimpla elongata, Crusopimpla rettigi, Crusopimpla violina, Theronia furensis, Xanthopimpla crescendae (152156) Crusopimpla minuta, Crusopimpla rettigi, Crusopimpla violina, Xanthopimpla crescendae (113147) Crusopimpla minuta, Crusopimpla violina, Epitheronia stigmatica, Xanthopimpla ciboisae, Xanthopimpla crescendae (114154) Crusopimpla weltii (236577) Epitheronia stigmatica (67711 152357 224404 226443 226444) Pimpla stigmatica (156743) Theronia nigriscutum (113029) Xanthopimpla ciboisae (226445) Xanthopimpla crescendae (226446)
Bridgerian	50.3 - 46.2	USA (Colorado)	Ichninsum appendicrassum (192071) Pimpla eocenica (115103) Scambus mandibularis (138671) Scambus parachuti (138673)
MP 11	48.6 - 40.4	Germany (Hessen)	Polyhelictes bipolarus (166830) Scambus fossilobus (125466 153473) Xanthopimpla messelensis, Xanthopimpla praeclara (193978)
Priabonian	38.0 - 33.9	Canada (British Columbia)	Pimpla saxea, Pimpla senecta, Pimpla decessa (121480)
Priabonian	38.0 - 33.9	United Kingdom (England)	Exeristes gurnetor (127743) Pimplinae indet. 1, Itoplectis saxosus (123960) Pimplinae indet. 2 (123959) Scambus fossilis (123961)
Priabonian	38.0 - 33.9	Russian Federation (Primorskiy)	Pimpla bibosa (123794) Xanthopimpla biamosa (172679) Xanthopimpla biamosa, Zabrachypus tumidus (172678)
Priabonian	38.0 - 33.9	France	Pimpla sp. (113709)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Crusopimpla klopfsteinae (192258) Pimpla succini (147344)
Priabonian	38.0 - 33.9	China (Shandong)	Pimpla amplifemura, Pimpla impuncta (156317)
Priabonian	38.0 - 33.9	Denmark	Firkantus freddykruegeri (123762)
Chadronian	37.2 - 33.9	USA (Colorado)	Lithoserix williamsi (144630) Pimpla appendigera (122976) Polysphincta inundata, Pimpla rediviva (106384)
MP 23	33.9 - 28.4	France	Lithoserix oublierus (157327) Pimpla anomalensis, Pimpla aquensis (122535)
Rupelian	33.9 - 28.1	Germany	Pimpla seyrigi (122407)
MP 30	28.4 - 23.03	Germany (Nordrhein-Westfalen)	Pimpla morleyi (138696)
Chattian	28.1 - 23.03	Germany (North Rhine-Westphalia)	Pimpla cyclostigmata, Polysphincta statzi (122010)
Chattian	28.1 - 23.03	France (Provence-Alpes-Côte d'Azur)	Ichneumon sublongaevus (122487) Pimpla anomalensis (122488)
Chattian	28.1 - 23.03	France (Bouches-Du-Rhone)	Pimpla aquensis (113712) Pimpla renevieri (153144) Pimpla saussurei, Pimpla antiqua (122501) Pimpla saussurii (122877)
Langhian	15.97 - 13.82	USA (Idaho)	Pimplinae indet. (97616)
Piacenzian	3.6 - 2.588	Germany (Niedersachsen)	Pimplites praeparatus (109196)
Gelasian	2.588 - 1.806	Greenland	Pimplinae indet. (160143 160146)
Chibanian	0.774 - 0.129	Japan	Pimpla sp. A, Pimpla sp. B, Pimplinae indet. (211426)