PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Elomeryx

Mammalia - Anthracotheriidae

Taxonomy

Elomeryx was named by Marsh (1894). Its type is Heptacodon armatus.

It was assigned to Anthracotheriinae by Hay (1902); to Anthracotheriidae by Marsh (1894), Carroll (1988), McKenna and Bell (1997), Thewissen et al. (2001) and Sach and Heizmann (2001); to Ruminantiamorpha by Spaulding et al. (2009); to Hippopotamoidea by Orliac et al. (2010); and to Bothriodontinae by Scott (1940), Boisserie et al. (2005), Boisserie et al. (2010), Kostopoulos et al. (2012) and Scherler et al. (2019).

Species

Bakalovia orientalis, Bothriodon chyelingensis, Bothriodon tientongensis, Brachyodus trofimovi, E. armatus (syn. E. mitis, Hyopotamus brachyrhynchus, E. brachystylus), E. asticus, E. borbonicus, E. cluai, E. crispus, E. garbanii, E. japonicus

Species lacking formal opinion data

E. woodi

Synonyms

Bakalovia orientalis was named by Böhme et al. (2013). Its type specimen is SAU 3, a partial skeleton, and it is a 3D body fossil. Its type locality is Na Duong Mine layer 80 , which is in a Bartonian/Priabonian lacustrine - large lignite in the Na Duong Formation of Vietnam.

It was considered a nomen dubium by Ducrocq et al. (2015).
Bothriodon chyelingensis was named by Xu (1977). Its type specimen is IVPP V 4981, a set of teeth, and it is a 3D body fossil.

It was considered a nomen dubium by Ducrocq and Lihoreau (2006).
Bothriodon tientongensis was named by Xu (1977). Its type specimen is IVPP V 4994, a mandible, and it is a 3D body fossil.

It was considered a nomen dubium by Ducrocq and Lihoreau (2006).
Brachyodus trofimovi was named by de Bonis et al. (1997).

It was considered a nomen dubium by Ducrocq and Lihoreau (2006).

Synonymy list

Year	Name and author
1894	Elomeryx Marsh
1902	Elomeryx Hay p. 652
1940	Elomeryx Scott p. 443
1977	Bothriodon chyelingensis Xu
1977	Bothriodon tientongensis Xu
1988	Elomeryx Carroll
1997	Elomeryx McKenna and Bell p. 405
1997	Brachyodus trofimovi de Bonis et al.
2001	Elomeryx Sach and Heizmann p. 30
2001	Elomeryx Thewissen et al. p. suppl.
2005	Elomeryx Boisserie et al.
2009	Elomeryx Spaulding et al.
2010	Elomeryx Boisserie et al. p. 329
2010	Elomeryx Orliac et al. p. 11872 figs. Fig. 2
2012	Elomeryx Kostopoulos et al.
2013	Bakalovia orientalis Böhme et al. p. 132 figs. 10-18
2019	Elomeryx Scherler et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()

Rank	Name	Author
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Euungulata
—	Artiodactylamorpha
—	Artiodactyla	()
—	Whippomorpha
superfamily	Hippopotamoidea	Gray 1821
family	Anthracotheriidae	Leidy 1869
subfamily	Bothriodontinae
genus	Elomeryx

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Elomeryx Marsh 1894

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†Elomeryx armatus Marsh 1894

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Invalid names: Elomeryx brachystylus Troxell 1921 [synonym], Elomeryx mitis Marsh 1894 [synonym], Hyopotamus brachyrhynchus Osborn and Wortman 1894 [synonym]

†Elomeryx asticus Nikolov 1967

†Elomeryx borbonicus Geais 1934

†Elomeryx cluai Deperet 1906

†Elomeryx crispus Gervais 1849

†Elomeryx garbanii Macdonald 1970

†Elomeryx japonicus Matsumoto 1925

Invalid names: Bakalovia orientalis Böhme et al. 2013 [nomen dubium], Bothriodon chyelingensis Xu 1977 [nomen dubium], Bothriodon tientongensis Xu 1977 [nomen dubium], Brachyodus trofimovi de Bonis et al. 1997 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialo

Locomotion:

actively mobilec

Life habit:

ground dwellingf

Diet:

browserf

Reproduction:

viviparoussubc

Created:

2005-08-31 14:33:14

Modified:

2005-08-31 16:33:14

Source: f = family, o = order, subc = subclass, c = class, subp = subphylum

References: Nowak 1999, Hendy et al. 2009, Lillegraven 1979, Kron and Manning 1998, Carroll 1988

Age range: base of the Priabonian to the top of the Arikareean or 38.00000 to 18.50000 Ma

Collections (33 total)

Time interval	Ma	Country or state	Original ID and collection number
Bartonian - Priabonian	41.3 - 33.9	Vietnam (Lang Son)	Bakalovia orientalis (169420)
Priabonian	38.0 - 33.9	Bulgaria	E. asticus (103742)
Priabonian	38.0 - 33.9	France (Provence-Alpes-Cote d'Azur Region)	E. crispus (31747)
Orellan	33.9 - 33.3	USA (Montana)	E. sp. (17090)
Orellan	33.9 - 33.3	Canada (Saskatchewan)	E. armatus (17111) E. sp. (17055)
Early/Lower Oligocene	33.9 - 28.4	Japan	Brachyodus japonicus (180396)
MP 21	33.9 - 28.4	Czech Republic	E. crispus (91466) E. woodi (184467)
MP 25	33.9 - 23.03	France (Alpes-de-Haute Provence)	E. crispus (183364)
Oligocene	33.9 - 23.03	France (Alpes-de-Haute Provence)	E. borbonicus (200439)
Whitneyan	33.3 - 30.8	USA (South Dakota)	E. armatus (185937) Hyopotamus brachyrhynchus (17344)
Whitneyan	33.3 - 30.8	Canada (Saskatchewan)	E. sp. (17372)
Whitneyan - Monroecreekian	33.3 - 24.8	USA (North Dakota)	E. armatus (17330)
Whitneyan - Monroecreekian	33.3 - 24.8	Canada (Saskatchewan)	E. armatus (17306)
Whitneyan - late Early Arikareean	33.3 - 18.5	USA (California)	Bothriodon brachyrhynchus (17527)
Geringian	30.8 - 26.3	USA (Nebraska)	E. sp. (17495)
Geringian	30.8 - 26.3	USA (South Dakota)	E. garbanii (17660)
Arikareean	29.5 - 18.5	USA (South Dakota)	E. garbanii (17520)
Late/Upper Oligocene	28.4 - 23.03	Pakistan (Baluchistan)	E. borbonicus (179812)
Late/Upper Oligocene	28.4 - 23.03	Japan	Brachyodus japonicus (183073) Brachyodus sp. (183066)
Late/Upper Oligocene	28.4 - 23.03	Greece (Thrace)	E. woodi (208602)
Late/Upper Oligocene	28.4 - 23.03	Georgia	E. borbonicus, Bothriodon velaunus, Bothriodon sp., Hyoboops sp. (182744)
Late/Upper Oligocene	28.4 - 23.03	France (Languedoc-Roussillon-Midi-Pyrénées)	E. crispus (182687)
MP 27	28.4 - 23.03	Switzerland	E. borbonicus (184454)
Monroecreekian	26.3 - 24.8	Canada (Saskatchewan)	E. sp. (17526)
MN 1	23.03 - 22.4	Switzerland	E. borbonicus (184456)
Aquitanian	23.03 - 20.44	Austria	E. borbonicus (76129)
Early/Lower Miocene	23.03 - 15.97	Greece (Pieria)	E. sp. (208601)
Burdigalian	20.44 - 15.97	Germany (Baden-Württemberg)	E. borbonicus (49102)
MN 3 - MN 4	20.0 - 15.97	Switzerland (Zürich)	E. borbonicus (46266)