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Icoisoptera

Insecta - Blattodea

Taxonomy
Icoisoptera was named by Engel et al. (2013). It is extant.

It was assigned to Artisoptera by Engel (2021); and to Euisoptera by Engel and Kaulfuss (2016), Wang et al. (2022).

Synonymy list
YearName and author
2013Icoisoptera Engel et al. p. 331
2016Icoisoptera Engel and Kaulfuss p. 95
2021Icoisoptera Engel
2022Icoisoptera Wang et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
RankNameAuthor
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—PolyneopteraMartynov 1938
—Holopandictyoptera
superorderDictyopteraLatreille 1829
orderBlattodea(Latreille 1810)
—Xylophagodea
infraorderIsoptera()
—Euisoptera
—Artisoptera
—Icoisoptera
—Icoisoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Icoisoptera Engel et al. 2013
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Fm. Kalotermitidae Froggatt 1897
+
G. Calcaritermes Snyder 1925
+
†Calcaritermes vetus Emerson 1969
G. †Cratokalotermes Bechly 2007
+
G. Cryptotermes Banks 1906 [drywood termite]
+
†Cryptotermes glaesarius Engel and Krishna 2007
†Cryptotermes pouilloni Jouault and Nel 2022
†Cryptotermes ryshkoffi Pierce 1958
†Cryptotermes yamini Krishna and Bacchus 1987
Invalid names: Cryptotermes batheri von Rosen 1913 [nomen nudum]
G. †Electrotermes von Rosen 1913
+
†Electrotermes affinis Hagen 1856
†Electrotermes flecki Nel and Bourguet 2006
†Electrotermes girardi Giebel 1856
G. †Eotermes Statz 1939
+
†Eotermes grandaeva Statz 1939
G. Glyptotermes Froggatt 1897
+
†Glyptotermes abyssinicus Bouju et al. 2021
†Glyptotermes grimaldii Engel and Krishna 2007
†Glyptotermes paleoliberatus Engel and Krishna 2007
†Glyptotermes pusillus Heer 1849
+
Invalid names: Termes (Eutermes) diaphanus Giebel 1856 [synonym], Termes (Eutermes) punctatus Giebel 1856 [synonym]
G. †Glyptotermites Armbruster 1941
+
†Glyptotermites assmuthi Armbruster 1941
G. †Huguenotermes Engel and Nel 2015
+
†Huguenotermes septimaniensis Engel and Nel 2015
G. Incisitermes Krishna 1961
+
†Incisitermes krishnai Emerson 1969
†Incisitermes peritus Engel and Krishna 2007
G. †Kachinitermes Engel et al. 2007
+
†Kachinitermes tristis Cockerell 1917
G. †Kachinitermopsis Engel and Delclòs 2010
+
G. Kalotermes Hagen 1853
+
Subg. Calotermes (Calotermes) Hagen 1853
†Kalotermes disruptus Cockerell 1917
†Kalotermes fossus Zhang et al. 1994
†Kalotermes nigellus Zhang et al. 1994
Kalotermes nigritus Snyder 1946
†Kalotermes nisus Zhang et al. 1994
†Kalotermes oeningensis von Rosen 1913
†Kalotermes piacentinii Piton and Théobald 1937
†Kalotermes rhenanus Hagen 1863
Invalid names: Calotermes bosniaskii Handlirsch 1907 [nomen nudum]
G. Neotermes Holmgren 1911
+
†Neotermes grassei Piton 1940
G. †Oligokalotermes Nel 1987
+
G. †Otagotermes Engel and Kaulfuss 2016
+
†Otagotermes novazealandicus Engel and Kaulfuss 2016
G. †Proelectrotermes von Rosen 1913
+
†Proelectrotermes berendtii Pictet and Hagen 1856
†Proelectrotermes fodinae Scudder 1883
†Proelectrotermes holmgreni Engel et al. 2007
†Proelectrotermes roseni Armbruster 1941
+
Invalid names: Calotermes (Neotermites) frischi Armbruster 1941 [synonym]
†Proelectrotermes swinhoei Cockerell 1916
Invalid names: Neotermites Armbruster 1941 [synonym]
G. †Prokalotermes Emerson 1933
+
†Prokalotermes hagenii Scudder 1883
G. †Pterotermopsis Engel and Kaulfuss 2016
+
†Pterotermopsis fouldenica Engel and Kaulfuss 2016
G. †Taieritermes Engel and Kaulfuss 2016
+
†Taieritermes krishnai Engel and Kaulfuss 2016
G. †Valkyritermes Jouault et al. 2022
+
†Valkyritermes inopinatus Jouault et al. 2022
Invalid names: Calotermes (Calotermes) erythrops von Rosen 1913 [nomen nudum], Calotermes (Calotermes) resinatus von Rosen 1913 [nomen nudum], Electrotermitinae Emerson 1942 [invalid subgroup], Protermitidae Holmgren 1909 [synonym]
Unr. Neoisoptera Engel et al. 2009
+
Fm. †Archeorhinotermitidae Krishna and Grimaldi 2003
+
G. †Archeorhinotermes Krishna and Grimaldi 2003
+
†Archeorhinotermes rossi Krishna and Grimaldi 2003
Fm. Rhinotermitidae Froggatt 1897
+
+
G. Coptotermes Wasmann 1896
+
†Coptotermes hirsutus Krishna and Grimaldi 2009
†Coptotermes paleodominicanus Krishna and Grimaldi 2009
†Coptotermes priscus Emerson 1971
†Coptotermes sucineus Emerson 1971
G. Dolichorhinotermes Snyder and Emerson 1925
+
†Dolichorhinotermes apopnus Engel and Krishna 2007
†Dolichorhinotermes dominicanus Schlemmermeyer and Cancello 2000
Subfm. Heterotermitinae Froggatt 1897
+
G. Heterotermes Froggatt 1897
+
†Heterotermes eocenicus Engel 2008
†Heterotermes primaevus Snyder 1960
Invalid names: Leucotermes Silvestri 1901 [synonym]
G. †Lukotermes Perkovsky and Nel 2021
+
†Lukotermes milescaput Perkovsky and Nel 2021
G. Reticulitermes Holmgren 1913
+
†Reticulitermes antiquus Germar 1813
+
Invalid names: Maresa plebeja Giebel 1856 [synonym], Termes (Eutermes) moestus Giebel 1856 [synonym], Termes fossile Ouchakoff 1838 [synonym], Termes giebeli Handlirsch 1907 [synonym], Termes gracilis Pictet and Hagen 1856 [synonym]
†Reticulitermes creedei Snyder 1938
†Reticulitermes dofleini Armbruster 1941
†Reticulitermes fossarum Scudder 1883
+
Invalid names: Eutermes meadii Scudder 1883 [synonym]
†Reticulitermes grimaldii Jouault and Nel 2021
+
Invalid names: Termes adustus Statz 1939 [synonym], Termes aethiops Statz 1939 [synonym], Termes atomus Statz 1939 [synonym], Termes blandus Statz 1939 [synonym], Termes concinnus Statz 1939 [synonym], Termes contractulus Statz 1939 [synonym]
†Reticulitermes laurae Pierce 1958
†Reticulitermes minimus Snyder 1928
+
Invalid names: Leucotermes borussicus von Rosen 1913 [synonym]
Invalid names: Hemerobites Germar 1813 [nomen oblitum], Maresa Giebel 1856 [replaced], Reticulitermes tibialis dubitans Pierce 1958 [synonym]
Subfm. Prorhinotermitinae Quennedey and Deligne 1975
+
G. †Zophotermes Engel 2011
+
†Zophotermes ashoki Engel and Singh 2011
G. Rhinotermes Hagen 1858
+
†Rhinotermes miocenicus Nel and Paicheler 1993
G. †Rhinotermites Armbruster 1941
+
†Rhinotermites dzierzoni Armbruster 1941
†Rhinotermites kuehni Armbruster 1941
†Rhinotermites wasmanni Armbruster 1941
Invalid names: Psammotermitinae Holmgren 1911 [empty]
Fm. Stylotermitidae Holmgren and Holmgren 1917
+
G. †Parastylotermes Snyder and Emerson 1949
+
†Parastylotermes calico Pierce 1958
†Parastylotermes frazieri Snyder 1955
†Parastylotermes krishnai Engel and Grimaldi 2011
†Parastylotermes nathani Perkovsky 2020
†Parastylotermes robustus von Rosen 1913
G. †Prostylotermes Engel and Grimaldi 2011
+
†Prostylotermes kamboja Engel and Grimaldi 2011
G. Stylotermes Holmgren and Holmgren 1917
Fm. Termitidae Latreille 1802 [termite]
+
Subfm. Apicotermitinae Grassé and Noirot 1955
+
G. Anoplotermes Muller 1873
+
†Anoplotermes bohio Krishna and Grimaldi 2009
†Anoplotermes cacique Krishna and Grimaldi 2009
†Anoplotermes carib Krishna and Grimaldi 2009
†Anoplotermes maboya Krishna and Grimaldi 2009
†Anoplotermes naboria Krishna and Grimaldi 2009
†Anoplotermes nitaino Krishna and Grimaldi 2009
†Anoplotermes quisqueya Krishna and Grimaldi 2009
†Anoplotermes taino Krishna and Grimaldi 2009
Subfm. Macrotermitinae Kemner 1934
+
G. Macrotermes Holmgren 1910
+
†Macrotermes pristinus Charpentier 1843
†Macrotermes scheuthlei Armbruster 1941
Subfm. Nasutitermitinae Hare 1937
+
G. Atlantitermes Fontes 1979
+
†Atlantitermes antillea Krishna and Grimaldi 2009
†Atlantitermes caribea Krishna and Grimaldi 2009
†Atlantitermes magnoculus Krishna and Grimaldi 2009
G. Caribitermes Roisin 1996
+
†Caribitermes hispaniola Krishna and Grimaldi 2009
G. Constrictotermes Holmgren 1910
+
G. Nasutitermes Dudley 1890
+
†Nasutitermes amplioculatus Krishna and Grimaldi 2009
†Nasutitermes crinitus Krishna and Grimaldi 2013
+
Invalid names: Nasutitermes pilosus Krishna and Grimaldi 2009 [replaced]
†Nasutitermes electrinus Krishna 1996
†Nasutitermes incisus Krishna and Grimaldi 2009
†Nasutitermes magnocellus Krishna and Grimaldi 2009
†Nasutitermes medioculatus Krishna and Grimaldi 2009
†Nasutitermes rotundicephalus Krishna and Grimaldi 2009
†Nasutitermes seminudus Krishna and Grimaldi 2009
G. Parvitermes Emerson 1949
+
†Parvitermes longinasus Krishna and Grimaldi 2009
G. Subulitermes Holmgren 1910
+
†Subulitermes hispaniola Krishna and Grimaldi 2009
†Subulitermes insularis Krishna and Grimaldi 2009
G. Trinervitermes Holmgren 1912
+
Invalid names: Eutermes (Trinervitermes) handlirschi von Rosen 1913 [nomen nudum]
G. Velocitermes Holmgren 1912
+
†Velocitermes bulbus Krishna and Grimaldi 2009
Subfm. Termitinae Latreille 1802
+
G. Amitermes Silvestri 1901
+
†Amitermes lucidus Krishna and Grimaldi 2009
G. Gnathamitermes Light 1932 [long jawed desert termite]
+
†Gnathamitermes rousei Pierce 1958
G. Gnathotermes Holmgren 1912
G. †Metatermites Armbruster 1941
+
†Metatermites statzi Armbruster 1941
G. Microcerotermes Silvestri 1901
+
†Microcerotermes insulanus Krishna and Grimaldi 2009
†Microcerotermes setosus Krishna and Grimaldi 2009
Invalid names: Microcerotermes latinotus von Rosen 1913 [nomen nudum]
G. †Nanotermes Engel and Grimaldi 2011
+
†Nanotermes isaacae Engel and Grimaldi 2011
G. Termes Linnaeus 1758
+
Subg. Termes (Rhinotermes) Hagen 1858
Subg. Termes (Termes) Linnaeus 1758
†Termes primitivus Krishna and Grimaldi 2009
†Termes siruguei Nel 1984
Invalid names: Mirotermes Wasmann 1897 [synonym], Termes hassencampi Heer 1860 [nomen nudum]
Invalid names: Amitermitinae Kemner 1934 [synonym], Eutermes fraasi von Rosen 1913 [nomen dubium], Eutermes nickeli Armbruster 1941 [nomen dubium], Eutermes sachtlebini Armbruster 1941 [nomen dubium], Mirotermes resinatus von Rosen 1913 [nomen dubium], Termes (Eutermes) croaticus Heer 1849 [nomen dubium], Termes (Eutermes) obscurus Heer 1849 [nomen dubium], Termes (Termes) amicus von Rosen 1913 [nomen dubium], Termes buchii Heer 1864 [nomen dubium], Termes drabatyi Armbruster 1941 [nomen dubium], Termes giganteus Hagen 1855 [nomen nudum], Termes hauffi Armbruster 1941 [nomen dubium], Termes korschefskyi Armbruster 1941 [nomen dubium], Termes scheeri Armbruster 1941 [nomen dubium], Termes schleipi Armbruster 1941 [nomen dubium], Termes stitzi Armbruster 1941 [nomen dubium], Termes weismanni Armbruster 1941 [nomen dubium]
G. †Waipiatatermes Engel and Kaulfuss 2016
+
†Waipiatatermes matatoka Engel and Kaulfuss 2016
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Aptian to the top of the Holocene or 119.57000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 113.2 Ma

Collections (102 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Aptian119.57 - 113.2Brazil (Ceará) Cratokalotermes santanensis (122557 140371)
Early/Lower Cenomanian100.5 - 93.9Myanmar (Kachin) Kalotermes burmensis (104053)
Early/Lower Cenomanian100.5 - 93.9Myanmar Proelectrotermes holmgreni (111688) Termopsis swinhoei, Hodotermes tristis, Archeorhinotermes rossi (123020) Valkyritermes inopinatus (179227)
Selandian61.66 - 59.24France (Auvergne) Neotermes grassei (113685) Termopsis piacentinii (120351)
Ypresian56.0 - 48.07India (Gujarat) Rhinotermitidae indet., Parastylotermes krishnai, Prostylotermes kamboja, Nanotermes isaacae, Zophotermes sp., Zophotermes ashoki (124463)
Ypresian56.0 - 48.07France (Oise) Electrotermes flecki (123850) Kalotermitidae indet. (68174)
Lutetian48.07 - 41.03Germany (Sachsen-Anhalt) Termes sp. (155693)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Electrotermes affinis (124194 124205) Hemerobites antiquus, Termes girardi, Termes obscurus (147349) Heterotermes eocenicus (124370) Leucotermes robustus, Calotermes affinis, Leucotermes antiquus (140221) Proelectrotermes berendti (123927 124073) Reticulitermes antiquus (123493) Reticulitermes antiquus, Electrotermes girardi, Reticulitermes minimus (123771) Reticulitermes grimaldii (220536) Reticulitermes minimus (147652) Termes fossile (147350)
Priabonian37.71 - 33.9Poland Electrotermes affinis (120811) Termes affinis, Termes berendtii, Termes antiquus (109625)
Priabonian37.71 - 33.9Denmark Electrotermes affinis, Proelectrotermes berendti (123762)
Priabonian37.71 - 33.9Ukraine Parastylotermes nathani (113621 122997) Parastylotermes nathani, Lukotermes milescaput (196704)
Priabonian37.71 - 33.9France (Gard) Kalotermitidae indet. J (144529)
Priabonian37.71 - 33.9United Kingdom (England) Reticulitermes sp. (21976) Sisyra disrupta (123959) Termitidae sp.A (123960)
Chadronian37.0 - 33.9USA (Colorado) Parotermes fodinae (106381) Parotermes hagenii, Parotermes fodinae, Eutermes fossarum, Eutermes meadii (106384)
Rupelian33.9 - 27.3France (Alpes-de-Haute) Kalotermitidae indet. I (114016)
Whitneyan31.8 - 29.5USA (Montana) Prokalotermes alderensis (120623)
Chattian27.3 - 23.04France (Provence-Alpes-Côte d'Azur) Termes siruguei (122489) Termitidae indet. (122487)
Chattian27.3 - 23.04Germany (North Rhine-Westphalia) Eotermes grandaeva, Calotermes rhenanus, Termes holmgreni, Termes aethiops, Termes blandus, Termes concinnus, Termes contractulus, Termes adustus, Termes atomus (122010)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Oligokalotermes fischeri (127415)
Late/Upper Oligocene27.3 - 23.04USA (Colorado) Reticulitermes creedei (121977)
MP 3027.3 - 23.04Germany Calotermes rhenanus (123945)
Aquitanian23.04 - 20.45Türkiye Termitidae indet. A (123307)
Aquitanian23.04 - 20.45Germany Kalotermes sp. (123872)
Aquitanian23.04 - 20.45New Zealand Waipiatatermes matatoka, Taieritermes krishnai, Otagotermes novazealandicus, Pterotermopsis fouldenica, Kalotermitidae indet. (50035)
Early/Lower Miocene23.04 - 15.98USA (California) Parastylotermes frazieri (152010)
Early/Lower Miocene23.04 - 15.98Mexico Heterotermes sp., Coptotermes sp., Calcaritermes sp. (21977)
Early/Lower Miocene23.04 - 15.98Mexico (Chiapas) Calcaritermes vetus (140212) Coptotermes sucineus (124469 138882) Coptotermes sucineus, Calcaritermes vetus (138881) Coptotermes sucineus, Heterotermes primaevus, Incisitermes krishnai, Kalotermes nigritus (124623) Dolichorhinotermes apopnus (124928) Heterotermes primaevus (140313) Nasutitermes electrinus (106432 124470 140369) Nasutitermes sp. (123456)
Early/Lower Miocene23.04 - 15.98Ethiopia Glyptotermes abyssinicus (224753)
Burdigalian20.45 - 15.98China (Shandong) Kalotermes nigellus, Kalotermes fossus (138945) Kalotermes nisus (138946) Kalotermes shandongianus (128552)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Constrictotermes electroconstrictus (123422) Coptotermes priscus, Coptotermes hirsutus, Coptotermes paleodominicanus, Anoplotermes bohio, Anoplotermes cacique, Anoplotermes carib, Anoplotermes maboya, Anoplotermes naboria, Anoplotermes nitaino, Anoplotermes quisqueya, Anoplotermes taino, Amitermes lucidus, Microcerotermes insulanus, Microcerotermes setosus, Termes primitivus, Atlantitermes antillea, Atlantitermes caribea, Atlantitermes magnoculus, Nasutitermes amplioculatus, Nasutitermes incisus, Nasutitermes magnocellus, Nasutitermes medioculatus, Nasutitermes pilosus, Nasutitermes seminudus, Subulitermes insularis, Subulitermes hispaniola, Nasutitermes rotundicephalus, Parvitermes longinasus, Caribitermes hispaniola, Velocitermes bulbus, Constrictotermes electroconstrictus, Nasutitermes electronasutus, Cryptotermes glaesarius, Cryptotermes yamini, Glyptotermes grimaldii, Glyptotermes paleoliberatus, Incisitermes peritus (122982) Cryptotermes pouilloni (228917) Dolichorhinotermes dominicanus (124282)
Hemingfordian18.5 - 16.3USA (California) Cryptotermes ryshkoffi (144521) Cryptotermes sp., Parastylotermes sp., Gnathotermes sp. (106431) Parastylotermes calico (144523) Reticulitermes laurae (144525) Reticulitermes laurae, Gnathamitermes magnoculus, Reticulitermes tibialis (144524) Reticulitermes sp. (144522)
Langhian15.98 - 13.82USA (Idaho) Eotermes sp. (97616)
Langhian15.98 - 13.82USA (Washington) Stylotermes washingtonensis (140652)
MN 515.98 - 13.82Germany (Baden-Württemberg) Calotermes roseni, Calotermes frischi, Calotermes assmuthi, Leucotermes dofleini, Rhinotermites dzierzoni, Rhinotermites kuehni, Rhinotermites wasmanni, Termes scheuthlei, Metatermites statzi, Eutermes nickeli, Eutermes sachtlebini, Termes drabatyi, Termes korschefskyi, Termes scheeri, Termes schleipi, Termes stitzi, Termes weismanni (127614) Eutermes fraasi (109058) Eutermes nickeli, Termes scheeri (155661) Termes scheuthlei, Termes hauffi (155659)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Termes hartungi, Termes buchii (117252)
Sarmatian12.8 - 11.63Croatia Termes obscurus, Termes croaticus (117250) Termes pristinus (113693 117246 156758)
Late/Upper Miocene11.63 - 5.333Japan Rhinotermitidae indet. (152335)
Turolian8.725 - 5.09France (Ardèche) Calotermes sp. (172761) Macrotermes scheuthlei, Kalotermitidae indet., Reticulitermes hartungi (156164) Reticulitermes sp., Termitidae indet., Kalotermitidae indet. F, Kalotermitidae indet. G (113893)
Messinian7.246 - 5.333France (Ardeche) Kalotermitidae indet. A, Kalotermitidae indet. B (122194)
Messinian7.246 - 5.333Italy Kalotermitidae indet. H (157195)
Messinian7.246 - 5.333Italy (Piedmont) Rhinotermes miocenicus (113998)
Messinian7.246 - 5.333Italy (Tuscany) Calotermes bosniaskii (113751)
Messinian7.246 - 5.333France (Cantal) Kalotermitidae indet. C, Kalotermitidae indet. D, Kalotermitidae indet. E (122195)
Pliocene5.333 - 2.58Tanzania (Eastern Drift Valley) Termitidae indet. (32050)
Piacenzian3.6 - 2.58Germany (Niedersachsen) Reticulitermes pliozaenicus (109196)
Gelasian2.58 - 1.8France (Cantal) Termitidae indet. B (157197)
Holocene0.0117 - 0.0Tanzania Calotermes batheri, Calotermes erythrops, Termes amicus, Mirotermes resinatus, Microcerotermes latinotus (150372) Termes diaphanus, Termes punctatus (150374) Termes pusillus (123418) Termes sp. (123991)
Holocene0.0117 - 0.0Benin Calotermes resinatus, Eutermes handlirschi (150373)