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Nanophoca vitulinoides

Mammalia - Carnivora - Phocidae

Taxonomy
Phoca vitulinoides was named by Van Beneden (1876). Its type specimen is neotype: IRSNB M2276a–q, a set of vertebrae (partial skeleton), and it is a 3D body fossil. Its type locality is Borsbeek, Fort 3, which is in a Tortonian lagoonal horizon in the Diest Formation of Belgium.

It was recombined as Phoca (Phoca) vitulinoides by Trouessart (1898) and Friant (1947); it was considered a nomen dubium by Koretsky and Ray (2008); it was recombined as Nanophoca vitulinoides by Dewaele et al. (2017), Berta et al. (2022) and Rule and Park (2024).

Synonymy list
YearName and author
1874Phoca vitulinoides Vanden Broeck p. 199
1876Phoca vitulinoides Van Beneden p. 800
1898Phoca (Phoca) vitulinoides Trouessart p. 385
1904Phoca vitulinoides Trouessart p. 286
1922Phoca vitulinoides Kellogg p. 118
1947Phoca (Phoca) vitulinoides Friant p. 7
1976Phoca vitulinoides Ray p. 394
2001Phoca vitulinoides Koretsky p. 85
2017Nanophoca vitulinoides Dewaele et al. p. 13
2022Nanophoca vitulinoides Berta et al. p. 7 figs. Table 1.1
2024Nanophoca vitulinoides Rule and Park p. 10 figs. Table 2

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
classMammalia
RankNameAuthor
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivora
suborderCaniformiaKretzoi 1943
infraorderCanoidea(Simpson 1931)
superfamilyArctoideaFlower 1869
PanpinnipediaWolsan et al. 2020
Pinnipedimorpha
Pinnipedia()
familyPhocidae()
subfamilyPhocinae
genusNanophoca
speciesvitulinoides()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Nanophoca vitulinoides Van Beneden 1876
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Diagnosis
ReferenceDiagnosis
L. Dewaele et al. 2017Nanophoca vitulinoides is a small seal, estimated to have reached a length of approximately
one meter, which is slightly smaller than members of the genus Pusa (1.3 m for male Pusa sibirica Ciesielski et al., 2006). It differs from other genera of Phocinae in the following characteristics: sacrum with three (also in Monachinae Gray, 1869) to four fused sacral vertebrae (also in other Phocinae); sacral spinous processes fused and dorsally elongate; prominent hook-like ischiatic spine; and a low proximodistally oriented ridge just proximal to the medial condyle of the femur, less than one millimeter raised over the condyle. Additionally, the following characteristics have also been observed in other Phocinae, but their combination is unique to N. vitulinoides: a scapular spine and subspinous ridge (see definition below) fuse at scapular neck (also in H. grypus, Phoca, and Pusa) lesser tubercle of humerus at same level as humeral head (also in Cryptophoca maeotica); greater trochanter of femur higher than head (also in Praepusa vindobonenesis, Pusa caspica, and Pusa sibirica); head of femur on narrow, long neck (also in C. maeotica, L. proxima, Monachopsis pontica, Praepusa vindobonensis, and Sarmatonectes sintsovi).
Measurements
No measurements are available
Composition: hydroxyapatiteuc
Ontogeny: modification of partsuc
Environment: marineuc
Locomotion: actively mobilec
Life habit: amphibiousuc
Diet: piscivoref
Diet 2: carnivoref
Reproduction: viviparousuc
Created: 2005-06-08 10:11:09
Modified: 2005-06-08 12:11:09
Source: f = family, c = class, uc = unranked clade
References: Nowak 1991, Gingerich 2003, Hendy et al. 2009, Carreño and Cronin 1993

Age range: base of the Serravallian to the top of the Zanclean or 13.82000 to 3.60000 Ma

Collections (7 total)


Time interval Ma Country or state Original ID and collection number
Miocene23.03 - 5.333Netherlands Phoca vitulinoides (190171)
Langhian - Serravallian15.97 - 11.62Belgium Nanophoca vitulinoides (91520)
Serravallian13.82 - 11.62Netherlands Phoca vitulinoides (75152)
Tortonian11.62 - 7.246Belgium Nanophoca vitulinoides (116630 217346)
Tortonian11.62 - 7.246Belgium (Antwerpen) Phoca vitulinoides (type locality: 52049)
Zanclean5.333 - 3.6Belgium (Antwerpen) Phoca vitulinoides (57701)