PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Dortokidae

Osteichthyes - Testudines

Taxonomy

Dortokidae was named by Lapparent de Broin and Murelaga (1996). It is not extant.

It was reranked as the unranked clade Dortokidae by Cadena and Joyce (2015).

It was assigned to Pleurodira by de Lapparent de Broin and Murelaga (1999); to Megapleurodira by Gaffney et al. (2006); to Pleurodira by Lapparent de Broin et al. (2004), Vremir (2013); and to Panpleurodira by Pérez-García et al. (2012), Pérez-García et al. (2014), Cadena and Joyce (2015).

Subtaxa

Dortoka (syn. Ronella), Eodortoka, Muehlbachia, Muehlbachia nopcsai

Synonyms

Muehlbachia was named by Vremir and Codrea (2009). It is not extant.

It was considered a nomen nudum by Cadena and Joyce (2015).
Muehlbachia nopcsai was named by Vremir and Codrea (2009). It is not extant.

It was considered a nomen nudum by Vremir and Codrea (2011); it was considered a nomen dubium by Rabi et al. (2013); it was considered a nomen nudum by Cadena and Joyce (2015).

Synonymy list

Year	Name and author
1996	Dortokidae Lapparent de Broin and Murelaga
1999	Dortokidae de Lapparent de Broin and Murelaga
2004	Dortokidae Lapparent de Broin et al.
2006	Dortokidae Gaffney et al. p. 29
2009	Muehlbachia Vremir and Codrea
2009	Muehlbachia nopcsai Vremir and Codrea
2012	Dortokidae Pérez-García et al. p. 151
2013	Dortokidae Vremir
2014	Dortokidae Pérez-García et al.
2015	Dortokidae Cadena and Joyce

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
class	Osteichthyes
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda

Rank	Name	Author
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Testudinata	(Oppel 1811)
order	Testudines	Batsch 1788
—	Panpleurodira
—	Dortokidae	()
—	Dortokidae	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Dortokidae Lapparent de Broin and Murelaga 1996

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G. †Dortoka Lapparent de Broin and Murelaga 1996

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†Dortoka botanica Gheerbrant et al. 1999

†Dortoka vasconica Lapparent de Broin and Murelaga 1996

†Dortoka vremiri Augustin et al. 2021

Invalid names: Ronella Gheerbrant et al. 1999 [synonym]

G. †Eodortoka Pérez-García et al. 2014

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†Eodortoka morellana Pérez-García et al. 2014

Invalid names: Muehlbachia Vremir and Codrea 2009 [nomen nudum], Muehlbachia nopcsai Vremir and Codrea 2009 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialuc

Locomotion:

actively mobilec

Created:

2017-04-17 15:15:35

Modified:

2017-04-17 15:15:35

Source: c = class, subp = subphylum, uc = unranked clade

References: Carroll 1988, Hendy et al. 2009, Kiessling 2004

Age range: base of the Early/Lower Barremian to the top of the Lutetian or 125.77000 to 41.03000 Ma

Collections (42 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Hauterivian - Early/Lower Barremian	132.6 - 121.4	Spain (Teruel)	Dortokidae indet. (187331)
Barremian	125.77 - 121.4	United Kingdom (England)	Eodortoka morellana (230379)
Early/Lower Barremian	125.77 - 121.4	Spain (Aragón)	Dortokidae indet. (28196)
Late/Upper Barremian	125.77 - 121.4	Spain (Valenciana)	Solemydidae indet. (152639)
Early/Lower Cenomanian	100.5 - 93.9	France (Poitou-Charentes)	Dortokidae indet. (77231 77233)
Santonian	85.7 - 83.6	Hungary (Veszprém)	Dortokidae indet. (48722)
Early/Lower Campanian	83.6 - 72.2	France (Languedoc-Roussillon)	Dortoka sp. (47782)
Early/Lower Campanian	83.6 - 72.2	Austria (Niederosterreich)	Dortokidae indet. (48720)
Late/Upper Campanian	83.6 - 72.2	Spain (Castilla y Leon)	Chelonia indet. (48861) Dortoka vasconica (28406 28765)
Middle Campanian	83.6 - 72.2	France (Provence-Alpes-Côte d'Azur)	Dortoka sp. (48849)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	France (Languedoc-Roussillon)	Dortoka sp. (74660)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Spain (Valencia)	Dortoka sp. (53918)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	France (Languedoc-Roussillon)	Dortoka sp. (59687) Dortoka vasconica (48870)
Late/Upper Campanian - Late/Upper Maastrichtian	83.6 - 66.0	France (Provence-Alpes-Cote d'Azur)	Dortoka sp. (59778)
Early/Lower Maastrichtian	72.2 - 66.0	Romania (Alba)	Dortokidae indet. (80858 80859 153475) Muehlbachia nopcsai (93729 97813 224587)
Early/Lower Maastrichtian	72.2 - 66.0	Romania	Muehlbachia sp. (115004)
Early/Lower Maastrichtian	72.2 - 66.0	Romania (Sebeş)	Dortokidae indet. (156124)
Early/Lower Maastrichtian	72.2 - 66.0	Romania (Hunedoara)	Dortoka vremiri (221768) Dortokidae indet. (13334 59410 67811 163322)
Late/Upper Maastrichtian	72.2 - 66.0	Romania	Dortokidae indet. (132350 181030)
Late/Upper Maastrichtian	72.2 - 66.0	Romania (Alba)	Muehlbachia sp. (156125)
Thanetian	59.24 - 56.0	Romania	Ronella botanica (83307 138239 138240 138242 138243 138244 138245)
Ypresian - Lutetian	56.0 - 41.03	Romania	Ronella sp. (178481)