| Basic info | Taxonomic history | Classification | Included Taxa |
| Morphology | Ecology and taphonomy | External Literature Search | Age range and collections |
Arctoperlaria
Taxonomy
Arctoperlaria was named by Zwick (1973). It is extant.
It was assigned to Plecoptera by Zwick (2000), Sroka et al. (2018).
It was assigned to Plecoptera by Zwick (2000), Sroka et al. (2018).
Synonyms
|
Synonymy list
| Year | Name and author |
|---|---|
| 1853 | Nemourina Newman |
| 1973 | Arctoperlaria Zwick |
| 1987 | Nemourina Sinitshenkova p. 45 |
| 1992 | Nemourina Sinitshenkova p. 50 |
| 2000 | Arctoperlaria Zwick |
| 2006 | Nemourina Liu et al. p. 550 |
| 2010 | Nemourina Sinitshenkova and Aristov p. 51 |
| 2018 | Arctoperlaria Sroka et al. p. 4 |
Is something missing? Join the Paleobiology Database and enter the data
|
|
If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subor. Arctoperlaria Zwick 1973
show all | hide all
Infraor. Euholognatha Zwick 1973
hide
Superfm. Nemouroidea Billberg 1820
+
Unr. Nemouromorpha Zwick 2000
+
Fm. Capniidae Banks 1900 [small winter stonefly]
+
G. †Dobbertiniopteryx Ansorge 1993
G. †Rovnocapnia Sinitshenkova 2009
Fm. Leuctridae Klapálek 1905 [rolled-winged stonefly]
+
G. †Baltileuctra Chen 2018
G. †Euroleuctra Chen 2018
G. Leuctra Stephens 1836
Subfm. Leuctrinae Klapálek 1905
G. †Lycoleuctra Sinitshenkova 1987
G. Megaleuctra Neave 1934
G. †Palaeoleuctra Sinitshenkova 2009
G. †Palaeopsole Caruso and Wichard 2011
G. Zealeuctra Ricker 1969
Fm. Nemouridae Billberg 1820 [spring stonefly]
+
Subfm. Amphinemurinae Baumann 1975
Subfm. Nemourinae Billberg 1820
G. †Parvinemoura Liu et al. 2008
Fm. Notonemouridae Ricker 1973
+
Subfm. †Paranotonemourinae Cui and Béthoux 2018
G. †Talbragaria Sroka and Prokop 2023
Fm. †Pronemouridae Liu et al. 2011
+
G. †Dimoula Sinitshenkova 2005
+
†Dimoula dimi Sinitshenkova 2005
†Dimoula khasurtensis Sinitshenkova 2011
G. †Pronemoura Liu et al. 2011
+
†Pronemoura angustithorax Liu et al. 2011
†Pronemoura longialata Liu et al. 2011
†Pronemoura minuta Liu et al. 2011
†Pronemoura peculiaris Liu et al. 2011
†Pronemoura shii Liu et al. 2011
Fm. Taeniopterygidae Klapálek 1905 [winter stonefly]
+
Subfm. Brachypterainae Zwick 1973
+
G. †Balticopteryx Chen 2018
G. Brachyptera Newport 1849
G. †Liaotaenionema Liu et al. 2008
G. †Gurvanopteryx Sinitshenkova 1986
+
†Gurvanopteryx effeta Sinitshenkova 1986
†Gurvanopteryx impleta Sinitshenkova 1986
G. †Jurataenionema Liu and Ren 2007
+
†Jurataenionema inornatus Liu and Ren 2007
†Jurataenionema rohdendorfi Sinitshenkova 2011
†Jurataenionema stigmaeus Liu and Ren 2007
G. †Mesotaeniopteryx Martynov 1937
+
†Mesotaeniopteryx elongata Martynov 1937
†Mesotaeniopteryx indefessa Sinitshenkova 1992
†Mesotaeniopteryx klapaleki Martynov 1937
†Mesotaeniopteryx semisessilis Sinitshenkova 1985
†Mesotaeniopteryx sinitsae Sinitshenkova 1987
†Mesotaeniopteryx splendida Martynov 1937
†Mesotaeniopteryx turgida Sinitshenkova 1987
Fm. †Perspicuusoperlidae Chen 2022
hide
Infraor. Systellognatha Enderlein 1969
hide
Superfm. Perloidea Latreille 1802
+
Fm. Chloroperlidae Okamoto 1912 [green stonefly]
+
G. †Dipsoperla Sinitshenkova 1987
+
†Dipsoperla kunikanensis Sinitshenkova 1990
†Dipsoperla serpentis Sinitshenkova 1987
Fm. Perlidae Latreille 1802 [common stonefly]
+
Subfm. Acroneuriinae Klapalek 1914
+
Tr. Acroneuriini Klapalek 1914
Tr. Anacroneuriini Stark and Gaufin 1976
G. †Billoperla Chen and Xu 2021
G. †Burmaperla Jouault and Nel 2022
G. †Burmesoperla Chen 2019
G. †Cretacroneuria Chen 2020
G. †Electroneuria Sroka et al. 2018
Tr. †Largusoperlini Chen 2018
Invalid names: Neoperlini Stark and Caufin 1976 [empty]
Fm. Perlodidae Klapalek 1912 [perlodid stonefly]
+
Superfm. Pteronarcyoidea Newman 1853
+
Fm. Peltoperlidae Claassen 1931
+
Subfm. †Petroperlinae Sroka et al. 2018
+
G. †Borisoperla Chen and Xu 2020
G. †Branchioperla Sroka and Staniczek 2019
G. †Dewaltoperla Chen 2023
G. †Graciloperla Chen et al. 2025
G. †Lapisperla Sroka et al. 2018
G. †Ovaloperla Chen and Xu 2022
G. †Petroperla Sroka et al. 2018
G. †Zwickoperla Chen and Wang 2020
Invalid names: Borisoperlinae Chen and Xu 2020 [synonym]
Fm. Pteronarcyidae Newman 1853 [salmonfly]
+
Invalid names: Perlomorpha Latreille 1802 [synonym]
Invalid names: Nemourina Newman 1853 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
|
|
||||
|
|
||||
|
|
||||
|
|
||||
|
|
||||
|
|
||||
| Source: o = order | |||||
| Reference: Tillyard 1921 | |||||
Age range
Maximum range based only on fossils: base of the Kungurian to the top of the Piacenzian or 283.30000 to 2.58000 Ma
Minimum age of oldest fossil (stem group age): 274.4 Ma
Minimum age of oldest fossil (stem group age): 274.4 Ma
Collections (175 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Kungurian | Russian Federation (Komi Republic) | Kaltanemoura derosa, Uralonympha vorkutica (151567) | |
| Kungurian | Russian Federation | Uralonympha varica (102504 196228 196229) | |
| Kungurian | Russian Federation (Suksun) | Tshekardoperla expulsa, Tshekardoperla depicta, Tshekardoperla squarrosa, Sylvoperlodes zhiltzovae, Rasilopsis irrita, Barathronympha victima (108206) | |
| Roadian | Russian Federation (Kemerovo) | Palaeoperla exacta, Palaeoperla prisca (122572) Palaeoperla exacta, Palaeoperla prisca, Palaeotaeniopteryx elegans (122568) Palaeoperla exacta, Palaeoperla prisca, Palaeotaeniopteryx elegans, Palaeotaeniopteryx kaltanicus, Palaeonemoura clara, Palaeonemoura altaica, Kaltanemoura depravata (104704) Palaeoperla prisca (122569) Palaeoperla prisca, Kaltanemoura depravata (122567) | |
| Wordian | Russian Federation (Udmurtiya) | Palaeonemourisca novojilovi (104751) | |
| Wordian | Russian Federation (Udmurtia) | Palaeonemoura furcata, Palaeonemoura abdita, Palaeonemoura benevola, Palaeotaeniopteryx trinerva, Palaeotaeniopteryx alligata, Palaeonemourisca corrugata, Votaknemoura persuasibil, Properla incrassata, Properla umbrosa, Kargaloperla grata, Kargaloperla fibrosa, Kargaloperla queata, Kargaloperla postica (104750) Palaeonemoura proximalis (103537) Palaeonemoura repleta, Palaeonemourisca strigosa (103502) | |
| Capitanian | Russian Federation (Orenburg) | Kargaloperla exuperata (104057) Palaeotaeniopteryx perlonga, Palaeotaeniopteryx distalis, Palaeonemoura lepida, Palaeonemoura petaloidea, Palaeonemoura duplicata, Palaeonemoura riparia, Palaeonemoura apicalis, Palaeonemoura remota, Palaeonemourisca subita, Palaeoperla perfracta (104717) Permoleuctropsis gracilis (104994) | |
| Wuchiapingian | Russian Federation (Vologda) | Palaeonemoura issadensis, Palaeotaeniopteryx laeta, Palaeotaeniopteryx triternata, Issadoperla permiana, Properla issadensis, Kargaloperla decipiens, Kargaloperla furcata, Mirumoperla multinerva, Palaeonemoura antrorsa, Palaeonemoura amica, Palaeotaeniopteryx dejecta, Palaeotaeniopteryx constricta, Palaeotaeniopteryx adjecta, Palaeotaeniopteryx succida, Palaeotaeniopteryx amissa, Palaeotaeniopteryx fixa, Palaeotaeniopteryx mira, Votaknemoura admiranda, Palaeonemourisca diluta, Palaeonemourisca formalis (104754) | |
| Wuchiapingian | Mongolia | Palaeonemoura finitima (97602) | |
| Lopingian | Antarctica | Uralonympha schopfi (124348) | |
| Wuchiapingian - Changhsingian | Russian Federation (Kemerovo) | Kaltanemoura repentina (151568) | |
| Changhsingian | Kazakhstan (Zaisan) | Kargaloperla avulsa, Kaltanemoura sharovi (105062) Palaeonemoura zwicki (107957) | |
| Changhsingian | Russian Federation | Properla tungussica (151553) Tungussonympha meyeni (151565) | |
| Ladinian | Kyrgyzstan (Osh) | Dicronemoura acaulis, Fritaniopsis brevicaulis, Fritaniopsis remota, Tritaniella mera, Tritaniella synneura (106088) Dicronemoura dira (113468) Dicronemoura dira, Dicronemoura acaulis, Dicronemoura declinata, Fritaniopsis brevicaulis, Fritaniopsis dependens, Fritaniopsis remota, Tritaniella mera, Tritaniella pectinata, Cristonemoura porrecta, Cristonemoura binerva (110763) Dicronemoura dira, Dicronemoura acaulis, Dicronemoura declinata, Fritaniopsis brevicaulis, Fritaniopsis dependens, Tritaniella mera, Tritaniella perlonga, Tritaniella synneura, Tritaniella pectinata (113406) Dicronemoura dira, Dicronemoura acaulis, Fritaniopsis brevicaulis, Tritaniella synneura, Triassonemoura ficteramosa (163987) Dicronemoura dira, Dicronemoura acaulis, Fritaniopsis remota, Tritaniella mera, Tritaniella synneura, Ramonemoura constricta (106090) Fritaniopsis brevicaulis (105007) Mesoleuctra brachypoda (151569 151570) | |
| Carnian | Argentina | Platyperla marquati (151962) | |
| Norian | South Korea | Baleyopterygidae indet. (125392) Platyperla sp. (191410) | |
| Norian | China (Yunnan) | Triassoperla yongrenensis (165249) | |
| Norian | Ukraine (Kharkiv) | Trianguliperla innoxia, Berekia neglecta (105057) | |
| Norian | Kazakhstan | Trianguliperla aequalis (105055 114782 151554 151566) | |
| Hettangian - Sinemurian | Kyrgyzstan (Tonskiy) | Karanemoura distalis (105084 105086 105087 113432) | |
| Early/Lower Jurassic - Middle Jurassic | Kazakhstan | Platyperla rigida (151561 151562 151563) | |
| Pliensbachian | Kyrgyzstan | Mesonemura turanica (114701) | |
| Pliensbachian | Russian Federation (Kemerovo) | Mesoleuctra gigantea (125341) Mesoleuctra gigantea, Baleyopteryx orthoclada (117990) Mesoleuctra gigantea, Mesoleuctra tibialis (125496) | |
| Pliensbachian - Toarcian | Russian Federation (Novokuznetsk) | Baleyopteryx orthoclada (105018) | |
| Pliensbachian - Toarcian | Kazakhstan (Karagandy) | Mesoleuctra gigantea (125495) | |
| Early/Lower Toarcian | Germany (Mecklenburg-Vorpommern) | Perlariopseidae indet., Dicronemoura furcata (114993) | |
| Early/Lower Toarcian | Germany | Dobbertiniopteryx capniomimus (114991) | |
| Toarcian | Russian Federation (Krasnoyarsk) | Baleyopteryx altera (110819) | |
| Toarcian | Kyrgyzstan (Batken) | Fritaniopsis inflata (105088) Karanemoura brevis (105124) Karanemoura brevis, Fritaniopsis inflata (110817) Mesoleuctra quadrata, Mesoleuctra exserta (113381) Perlariopsis gravis, Karanemoura brevis, Fritaniopsis inflata (105003) | |
| Toarcian | Russian Federation (Irkutsk) | Mesoleuctra gracilis, Platyperla platypoda, Mesonemura maakii (114696) Mesotaeniopteryx semisessilis (104995) Perlisca aufuga (125228 125494) Perlisca aufuga, Baleyopteryx orthoclada (122950) | |
| Toarcian | Russian Federation (Kemerovo) | Platyperla caudiculata (125493) | |
| Toarcian | Kyrgyzstan | Mesotaeniopteryx elongata, Mesotaeniopteryx klapaleki (114702) Mesotaeniopteryx elongata, Mesotaeniopteryx splendida (114703) Taeniopterygidae indet. (114704) | |
| Toarcian - Aalenian | Russian Federation (Kemerovo) | Baleyopteryx orthoclada (125340) | |
| Aalenian | Mongolia | Platyperla admissa, Mesotaeniopteryx turgida (123958) | |
| Bajocian - Bathonian | Mongolia (Uver-Khangay Aymag) | Plutopteryx beata (118011) | |
| Bajocian - Bathonian | Mongolia | Platyperla propera (151564) | |
| Bajocian - Bathonian | Mongolia (Bayan-Khongor) | Mesotaeniopteryx sinitsae (124340) Perlariopsis fidelis (114615) | |
| Bajocian - Bathonian | Russian Federation (Krasnoyarsk) | Udopteryx kovalevi, Plutopteryx popovi, Platyperla conferta (118010) | |
| Bathonian | Mongolia | Mesoleuctroides latus, Trianguliperla orbiculata (119621) | |
| Callovian - Oxfordian | China (Nei Mongol) | Aristoleuctra yehae (113376) Paranotonemoura zwicki (183155) Pronemoura shii, Pronemoura longialata, Pronemoura peculiaris, Pronemoura angustithorax, Pronemoura minuta, Dobbertiniopteryx juracapnia, Karanemoura manca, Karanemoura abrupta, Jurataenionema inornatus, Jurataenionema stigmaeus, Protaenionema fuscalatus, Mengitaenioptera multiramis, Noviramonemoura trinervis (112663) Pteroliriope sinitshenkovae (113378 155751 172659) | |
| Callovian - Oxfordian | China (Hebei) | Sinotaeniopteryx luanpingensis (154786) | |
| Callovian - Oxfordian | China (Jilin) | Platyperla platypoda (154827) | |
| Callovian - Oxfordian | China (Liaoning) | Mesoleuctra peipiaoensis, Perlariopsis peipiaoensis (156597) Platyperla platypoda (154823 173107) Rectonemoura yujiagouensis (154729) Sinoperla abdominalis (156596) Sinotaeniopteryx chengdeensis (113475) | |
| Oxfordian | Russian Federation (Buryat) | Mesoleuctroides saturatus, Perlariopsis stipitatus, Spinoperla spinosa (105017) Perlariopsis stipitatus (104949) | |
| Oxfordian | Russian Federation | Trianguliperla attenuata, Baleyopteryx subitus (124813) | |
| Oxfordian | Russian Federation (Chita) | Mesoleuctroides saturatus (125499 125500 125501 125502) | |
| Oxfordian | Russian Federation (Buryatia) | Baleyopteryx blanda (124814) Mesoleuctroides saturatus (123194 123195 125403 125497) Perlariopsis sp. (125498) Perlomimus multus (125229) | |
| Oxfordian - Kimmeridgian | Russian Federation (Buryat) | Chloroperloides fusiformis, Udopteryx complicata (104950) | |
| Oxfordian - Kimmeridgian | Russian Federation (Chita) | Mogzonoperla truncata (114661) | |
| Late/Upper Jurassic | China (Xinjiang) | Sinoephemera kingi (124528) | |
| Kimmeridgian | Kazakhstan | Karanemoura abrupta (106102) Karanemoura abrupta, Perlariopsis mediana (105971) Perlariopsis martynovi, Karanemoura divaricata (105426) Perlariopsis mediana (106101) Udopteryx expansa, Karanemoura abrupta (113439) Udopteryx expansa, Karanemoura divaricata (106279) | |
| Tithonian | Mongolia (Dund-Gov’ Aymag) | Karanemoura desiliens, Perlariopsis basalis, Perlariopsis terna, Karanemoura appropinquata (122321) | |
| Tithonian | Australia (New South Wales) | Talbragaria australis (128121) | |
| Tithonian | Russian Federation (Chita) | Dimoula dimi, Dipsoperla serpentis, Nemourisca diligens, Lycoleuctra lupina (114307) Dipsoperla serpentis, Positopteryx dubia (123197) Pectinoperla notabilis (125407) Perlitodes aenigmaticus (119397) Savina laeta (140476) | |
| Tithonian | Russian Federation | Dipsoperla kunikanensis (124812) Lycoleuctra lupina (138151) Positopteryx dubia (123198) | |
| Tithonian | Mongolia | Baleyopteryx urocornis (151571) Bestioperlisca inulata, Trianguliperla optanda (121394) Platyperla mendosa, Trianguliperla optanda, Mesoleuctroides derelictus, Dicronemoura attenuata, Karanemoura perpropinqua, Mongolonemoura inversa (104748) | |
| Berriasian - Barremian | Russian Federation (Yakutiya) | Baissoleuctra conspecta, Isoperlodes perstrictus, Perlitodes borealis, Dicronemoura sibirica, Dicronemoura pusilla, Dicronemoura muscosa, Karanemoura probata, Mesotaeniopteryx indefessa, Fritaniopsis extricata, Baissoleuctra breviuscula (110818) | |
| Late/Upper Berriasian | United Kingdom (England) | Ecdyoperla fairlightensis (123449) | |
| Valanginian - Barremian | Mongolia (Dornogov) | Trianguliperla quassa (113407) | |
| Hauterivian | Russian Federation (Chita) | Platyperla parricidalis (152177) | |
| Hauterivian - Barremian | Mongolia (Ovorkhangai) | Accretonemoura radiata (113357) | |
| Barremian | Russian Federation (Chita) | Trianguliperla limosa, Mesoleuctrisca pronotalis (125782) Trianguliperla limosa, Trianguliperla volucris, Mesoleuctrisca pronotalis, Karanemoura curvata (113599) Trianguliperla volucris (125784) | |
| Late/Upper Barremian | China (Liaoning) | Archaeoperla rarissimus, Parvinemoura parvus (140490) Liaotaenionema tenuitibia (140491) Sinosharaperla zhaoi (118075) | |
| Early/Lower Aptian | Mongolia (Ara-Hangayn aymag) | Derancheperla collaris, Baleyopteryx bibula (123464) | |
| Aptian | Mongolia (Govi-Altai) | Gurvanopteryx effeta, Gurvanopteryx impleta (122332) | |
| Aptian | Russian Federation (Buryatia) | Accretonemoura grata (126000) Baissoleuctra irinae, Baissoleuctra sparsa (105536) Oecanthoperla sibirica (105969) Rasnitsyrina culonga, Dimoula khasurtensis, Jurataenionema rohdendorfi, Perlomorpha indet. (113349) Rasnitsyrina culonga, Dimoula khasurtensis, Jurataenionema rohdendorfi, Siberiopelta bashkuevi (113350) | |
| Early/Lower Cenomanian | Myanmar | Acroneuriinae indet., Largusoperla sp., Petroperla mickjaggeri, Lapisperla keithrichardsi, Electroneuria ronwoodi, Largusoperla charliewattsi, Largusoperla billwymani, Largusoperla micktaylori, Largusoperla brianjonesi, Branchioperla ianstewarti (123053) Burmaperla pouilloni (212258) Largusoperla acus, Largusoperla flata, Largusoperla arcus, Largusoperla reni, Starkoperla longusocollum, Zwickoperla brevicauda, Borisoperla kondratieffi, Billoperla starki, Ovaloperla staniczeki (161726) Perlidae indet., Pinguisoperla yangzhouensis, Largusoperla crassus, Largusoperla difformitatem, Burmesoperla expansa, Largusoperla dewalti, Largusoperla borisi, Burmacroneuria projecta, Cretacroneuria angularis, Burperla decolorata, Perspicuusoperla lata, Largusoperla zhouchangfai, Kachinoperla zwicki, Dewaltoperla edwardi, Cavoperla excavata, Crossoperla teslenkoae, Largusoperla vena, Graciloperla bulbosa, Graciloperla stylata (194479) | |
| Priabonian | Russian Federation | Podmosta attenuata (123419) | |
| Priabonian | Poland | Nemoura sp., Perla succinica, Perla resinata, Nemoura lata, Leuctra minuscula (120811) Perla succinica, Perla resinata, Taeniopteryx ciliata, Taeniopteryx elongata, Nemoura affinis, Nemoura puncticollis, Nemoura ocularis, Leuctra fusca, Leuctra gracilis, Leuctra linearis, Perla prisca (109625) | |
| Priabonian | United Kingdom (England) | Nemoura priscula (21976) | |
| Priabonian | Ukraine | Rovnocapnia ambita (113621) Rovnocapnia atra, Palaeoleuctra acuta (122997) | |
| Priabonian | Russian Federation (Kaliningrad) | Isoperla baltica (123936) Lednia zilli (110735) Leuctridae indet., Baltileuctra foraminis, Euroleuctra gillesi, Balticopteryx dui, Balticonemoura bulbosus, Baltileuctra dewalti, Brachyptera dewalti (194875) Megaleuctra neavei (148889) Nemoura sp. (123927) Palaeopsole weiterschani (120814) Zealeuctra cornuta (124468) | |
| Oligocene | Mexico (Puebla) | Euperlida parvicercifera (31519) | |
| MP 30 | Germany | Leuctra antiqua (123945) | |
| Burdigalian - Langhian | Dominican Republic | Dominiperla antigua (122473) | |
| MN 3 | Czechia | Perla burmeisteriana (105508) | |
| Langhian | USA (Idaho) | Megaleuctra jewetti (97616) | |
| Langhian | China (Fujian) | Amphinemura himberii (220273) | |
| Messinian | France (Cantal) | Perlomyia cantalensis (122195) | |
| Piacenzian | Japan | Perlodidae indet. (188973) | |
| Piacenzian | Germany (Niedersachsen) | Brachyptera schmidti (109196) |