PBDB Taxon

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Titanosauridae (titanosaurid)

Osteichthyes - Titanosauridae

Taxonomy

Titanosauridae was named by Lydekker (1885). Its type is Titanosaurus.

It was synonymized subjectively with Atlantosauridae by Charig (1967), Swinton (1970), Bonaparte (1978).

It was assigned to Homalosauropodidae by Huene (1948), Young (1958); to Cetiosauria by Colbert (1964); to Homalosauropodoidea by Kuhn (1966); to Saurischia by Madsen and Miller (1979); to Sauropoda by Bakker (1986); to Sauropodomorpha by Powell (1987); to Sauropoda by Molnar (1992), McIntosh (1997); to Sauropoda by Lydekker (1885), Marsh (1895), Marsh (1896), Marsh (1898), Huene (1927), Huene (1927), Huene (1929), Swinton (1936), Gilmore (1946), Romer (1956), Huene (1959), Romer (1966), Reig (1970), Rozhdestvensky (1977), Galton (1977), Ostrom (1980), Russell (1984), Carroll (1988), McIntosh (1990), Benton (1990), Jacobs et al. (1993), Hunt et al. (1995), Dong (1998); to Titanosauroidea by Upchurch (1995), Upchurch (1995), Upchurch (1997), Salgado (2003); to Eutitanosauria by Powell (2003); to Sauropoda by Lydekker (1893), Huene (1927), Huene (1929), Huene (1933), Roxo (1937), Hoffet (1942), Kuhn (1946), Lapparent (1947), Parsch (1963), Bonaparte (1980), Martinelli and Forasiepi (2004); and to Titanosauria by Bonaparte and Coria (1993), Bonaparte (1996), Salgado et al. (1997), Bonaparte (1999), Santucci and Bertini (2001), Smith et al. (2001), Calvo and González Riga (2003), González-Riga (2003), de Almeida et al. (2004), Salgado and Coria (2005), Apesteguía (2005), Novas et al. (2005), Moser et al. (2006), Calvo et al. (2007), Calvo et al. (2007), González Riga et al. (2009).

Subtaxa

Antarctosaurinae, Hypselosaurus, Titanosaurinae

Synonyms

Antarctosauridae was named by Olshevsky (1978).

It was synonymized subjectively with Titanosauridae by Dong (1998).
Hypselosaurus was named by Matheron (1869).

It was considered a nomen dubium by Le Loeuff (1993), Le Loeuff (1995); it was considered a nomen dubium by Casanovas-Cladellas et al. (1995); it was considered a nomen dubium by Upchurch et al. (2004), Le Loeuff (2005).

It was assigned to Crocodilia by Gervais (1869), Matheron (1869); to Hadrosauridae by Zittel (1890); to Atlantosauridae by Nopcsa (1901); to Morosauridae by Depéret (1900), Gautier (1901); to Sauropoda by Huene (1908), Huene (1909), von Zittel (1911); to Diplodocidae by Bataller (1960); to Sauropoda by Tatarinov (1964); to Titanosaurinae by Nopcsa (1928), Romer (1956), Charig (1967), Steel (1970), Swinton (1970), Sanz (1986), Casanovas et al. (1987); and to Titanosauridae by Huene (1927), Huene (1927), Huene (1927), Huene (1929), Huene (1941), Gilmore (1946), Young (1965), Romer (1966), Clottes and Raynaud (1983), Carroll (1988), Cousin et al. (1989), Buffetaut (1989), McIntosh (1990), McIntosh (1990), Buffetaut et al. (1991), Upchurch (1995), Jacobs et al. (1996), Sahni (1997), Powell (2003).
Hypselosaurus priscus was named by Matheron (1869). It is a 3D body fossil. Its type locality is Rognac train station, which is in a Maastrichtian terrestrial sandstone in France.

It was considered a nomen dubium by Le Loeuff (1993); it was considered a nomen dubium by Le Loeuff (1995); it was considered a nomen dubium by Le Loeuff (1998); it was considered a nomen dubium by Upchurch et al. (2004), Le Loeuff (2005); it was considered a nomen dubium by Tortosa et al. (2010), Le Loeuff (2010), Díez Díaz and Pereda Suberbiola (2010).

Synonymy list

Year	Name and author
1869	Hipselosaurus Gervais p. 215
1869	Hipselosaurus priscus Gervais p. 215
1869	Hypselosaurus Matheron p. 370
1869	Hypselosaurus priscus Matheron p. 370
1869	Hypselosaurus Matheron p. 790
1869	Hypselosaurus priscus Matheron p. 790
1873	Hypselosaurus priscus Gervais p. 469
1885	Titanosauridae Lydekker p. 37
1890	Hypselosaurus Zittel p. 766
1893	Titanosauridae Lydekker p. 3
1895	Titanosauridae Marsh
1896	Titanosauridae Marsh p. 242
1898	Titanosauridae Marsh p. 158
1900	Hypselosaurus priscus Depéret p. 637
1900	Hypselosaurus Depéret pp. 638-639
1901	Hypselosaurus priscus Gautier p. 113
1901	Hypselosaurus Gautier p. 115
1901	Hypselosaurus Nopcsa p. 205
1901	Hypselosaurus priscus Nopcsa p. 205
1908	Hypselosaurus Huene p. 348
1909	Hypselosaurus Huene p. 15
1911	Hypselosaurus von Zittel p. 288
1924	Hypselosaurus priscus Joleaud p. 44
1927	Titanosauridae Huene p. 125
1927	Hypselosaurus Huene p. 126
1927	Titanosauridae Huene p. 256
1927	Hypselosaurus Huene p. 270
1927	Hypselosaurus priscus Huene p. 270
1927	Titanosauridae Huene p. 467
1927	Hypselosaurus Huene p. 469
1928	Hypselosaurus Nopcsa p. 184
1929	Hypselosaurus priscus Huene p. 94
1929	Hypselosaurus Huene p. 107
1929	Titanosauridae Huene p. 155
1931	Hypselosaurus priscus Jepsen p. 13
1932	Hypselosaurus priscus Stromer p. 6
1933	Titanosauridae Huene p. 70
1936	Titanosauridae Swinton p. 209
1937	Titanosauridae Roxo p. 65
1941	Hypselosaurus Huene p. 90
1941	Hypselosaurus priscus Huene p. 90
1942	Titanosauridae Hoffet p. 53
1946	Titanosauridae Gilmore p. 29
1946	Hypselosaurus Gilmore p. 30
1946	Titanosauridae Kuhn p. 66
1947	Titanosauridae Lapparent p. 17
1947	Hypselosaurus priscus Lapparent p. 21
1948	Titanosauridae Huene p. 93
1948	Hypselosaurus priscus Lapparent p. 1186
1956	Titanosauridae Romer p. 621
1956	Hypselosaurus Romer p. 622
1958	Titanosauridae Young p. 25
1959	Titanosauridae Huene p. 121
1960	Hypselosaurus Bataller p. 159
1960	Hypselosaurus priscus Jeletzky p. 34
1961	Hypselosaurus priscus Schwarz et al. p. 346
1963	Titanosauridae Parsch p. 4
1964	Titanosauridae Colbert p. 22 fig. 6
1964	Hypselosaurus Tatarinov p. 552
1965	Hypselosaurus Young p. 169
1965	Hypselosaurus priscus Young p. 169
1966	Titanosauridae Kuhn p. 99
1966	Hypselosaurus Romer p. 370
1966	Titanosauridae Romer p. 370
1967	Hypselosaurus Charig p. 713
1967	Hypselosaurus priscus Charig p. 713
1967	Hypselosaurus priscus Lapparent p. 10
1970	Titanosauridae Reig p. 279
1970	Hypselosaurus Steel p. 76
1970	Hypselosaurus priscus Steel p. 76
1970	Hypselosaurus Swinton p. 183
1971	Hypselosaurus priscus Dinca et al. p. 90
1972	Hypselosaurus priscus Erben p. 17
1976	Hypselosaurus priscus Dughi and Sirugue p. 3
1977	Titanosauridae Galton p. 230
1977	Hypselosaurus priscus Krampitz et al. p. 583
1977	Titanosauridae Rozhdestvensky p. 113
1978	Antarctosauridae Olshevsky p. 27
1979	Titanosauridae Madsen and Miller p. 5
1980	Titanosauridae Bonaparte p. 82
1980	Titanosauridae Ostrom p. 22
1981	Hypselosaurus priscus Borgomanero and Leonardi p. 215
1983	Hypselosaurus Clottes and Raynaud pp. 10-11
1983	Hypselosaurus priscus Clottes and Raynaud p. 10
1984	Hypselosaurus priscus Brinkmann p. 297
1984	Titanosauridae Russell p. 29
1985	Hypselosaurus priscus Penner p. 668
1986	Titanosauridae Bakker p. 461
1986	Hypselosaurus priscus Ginsburg p. 144
1986	Hypselosaurus Sanz p. 85
1986	Hypselosaurus priscus Villatte et al. p. 93
1987	Hypselosaurus Casanovas et al. p. 97
1987	Hypselosaurus priscus Casanovas et al. p. 100
1987	Titanosauridae Powell p. 148
1988	Hypselosaurus Carroll
1988	Titanosauridae Carroll
1988	Hypselosaurus priscus Masriera and Ullastre p. 74
1988	Hypselosaurus priscus Vianey-Liaud et al. p. 416
1989	Hypselosaurus Buffetaut p. 438
1989	Hypselosaurus Cousin et al. p. 165
1989	Hypselosaurus priscus Cousin et al. p. 165
1989	Hypselosaurus priscus Le Loeuff et al. p. 851
1990	Titanosauridae Benton p. 19
1990	Hypselosaurus McIntosh p. 57
1990	Titanosauridae McIntosh p. 394
1990	Hypselosaurus McIntosh p. 398
1990	Hypselosaurus priscus McIntosh p. 398
1991	Hypselosaurus Buffetaut et al. p. 34
1991	Hypselosaurus priscus Buffetaut et al. p. 34
1992	Hypselosaurus priscus Buffetaut p. 130
1992	Titanosauridae Molnar p. 266
1993	Titanosauridae Bonaparte and Coria
1993	Hypselosaurus priscus Buffetaut et al. p. 170
1993	Hypselosaurus priscus Currie and Eberth p. 139
1993	Titanosauridae Jacobs et al. p. 524
1994	Hypselosaurus priscus Britt and Naylor p. 257
1995	Titanosauridae Hunt et al. p. 264
1995	Titanosauridae Upchurch p. 253
1995	Hypselosaurus Upchurch p. 380
1996	Titanosauridae Bonaparte p. 102
1996	Hypselosaurus Jacobs et al. p. 602
1997	Hypselosaurus priscus Erickson p. 4
1997	Titanosauridae McIntosh p. 657
1997	Hypselosaurus Sahni p. 364
1997	Titanosauridae Salgado et al. p. 20
1997	Titanosauridae Upchurch p. 82A
1998	Titanosauridae Dong p. 85
1999	Titanosauridae Bonaparte pp. 160-161
2001	Hypselosaurus priscus Hirsch p. 381
2001	Titanosauridae Santucci and Bertini p. 308
2001	Titanosauridae Smith et al. p. 1705 fig. 3
2003	Titanosauridae Calvo and González Riga p. 335
2003	Titanosauridae González-Riga
2003	Titanosauridae Powell pp. 21–22, 156
2003	Hypselosaurus Powell p. 75
2003	Hypselosaurus priscus Powell p. 75
2003	Titanosauridae Salgado p. 17 fig. 1
2004	Titanosauridae Martinelli and Forasiepi
2004	Titanosauridae de Almeida et al. p. 242
2005	Titanosauridae Apesteguía pp. 261-262
2005	Titanosauridae Novas et al. p. 37
2005	Titanosauridae Salgado and Coria p. 435
2006	Titanosauridae Moser et al. p. 46
2007	Titanosauridae Calvo et al. p. 497 fig. 14
2007	Titanosauridae Calvo et al. p. 533 fig. 3
2007	Hypselosaurus priscus Malkani
2009	Titanosauridae González Riga et al. p. 144 fig. 11

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
class	Osteichthyes
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)

Rank	Name	Author
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Saurischia	()
—	Sauropodomorpha	(Huene 1932)
—	Massopoda
—	Sauropodiformes
—	Sauropoda	()
—	Gravisauria
—	Eusauropoda
—	Neosauropoda
—	Macronaria
—	Titanosauriformes
—	Somphospondyli
—	Titanosauria
family	Titanosauridae
family	Titanosauridae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Titanosauridae Lydekker 1885 [titanosaurid]

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Invalid names: Antarctosauridae Olshevsky 1978 [synonym], Antarctosaurinae Powell 1986 [empty], Hypselosaurus Matheron 1869 [nomen dubium], Titanosaurinae Lydekker 1893 [empty]

Diagnosis

Reference	Diagnosis
R. Lydekker 1893	Sauropodous Dinosaurs in which the caudal vertebrae, with the excepcion [sic] of the firt, are procoelous, the presacrals opisthocoelous, and the lumbars without lateral cavities; the superior extremities of the chevron-bones being open.
F. v. Huene 1948	First caudal biconvex, all vertebrae in front of it opisthocoelous, behind it procoelous. No bifid neural spines; cavernous, lamellar structure of neural arches; quadrate ham-shaped; at least 12 cervicals, 6 sacrals. Originating from Cetiosaurs. Upper Jurassic to Upper Cretaceous.
L. Salgado et al. 1997	Absence of hyposphene-hypantrum articulation in posterior trunk vertebrae. Six sacral vertebrae. Anterior caudals strongly procoelous having "ball and socket" articular faces. Semilunar sternal plates. Claw on manual digit I absent. Manual phalanges absent. Preacetabular lobe of ilium nearly horizontal, outwardly projected.
J. E. Powell 2003	Quadrupedal herbivorous dinosaurs, of medium to gigantic size. Skull with very long and recurved paroccipital processes. Cylindrical teeth with open roots, slightly spatulated when unworn, except in Ampelosaurus. Short basipterygoid processes. Thirteen cervical vertebrae, probably 12 dorsals, 6 sacrals and more than 30 caudals. The opisthocoelous presacral vertebrae have cancellous structure in the centra and bulky parts of the neural arch (neural arch peduncles, ends of neurapophyses and diapophyses). Dorsal vertebrae without hyposphene-hypantrum except in the primitive Epachthosaurus. Sacrum with completely fused vertebrae. Sacral ribs with an 'I'-shaped cross section, participating in the conformation of the acetabular articular surface. First caudal biconvex or procoelous. Remaining caudals with a posterior articular condyle, with some occasional amphicoelic elements on the distal region of the tail. Neural arches joined to the anterior portion of the centrum in the middle and posterior caudals. Hemapophyses proximally open. Scapula with the glenoid articular facet oriented forward, down and inwards. Quadrangular coracoids. Sternal plates long and relatively broad, with a concave lateral margin. Fore limbs shorter than the hind limbs. Straight femur, somewhat anteroposteriorly flattened. Fibula with a marked lateral process. Astragalus relatively narrow.

Measurements

No measurements are available

Composition:

hydroxyapatiteinfrao

Entire body:

yesinfrao

Adult length:

10 to < 100infrao

Adult width:

10 to < 100infrao

Adult height:

10 to < 100infrao

Maximum body mass:

10000 kgg

Minimum body mass:

3000 kgg

Thickness:

thickinfrao

Architecture:

compact or denseinfrao

Form:

sphericalinfrao

Ontogeny:

accretion, modification of partsinfrao

Grouping:

gregariousinfrao

Environment:

terrestrialinfrao

Locomotion:

actively mobileinfrao

Life habit:

ground dwellinginfrao

Diet:

herbivoreinfrao

Reproduction:

oviparousinfrao

Dispersal:

direct/internalinfrao

Dispersal 2:

mobileinfrao

Created:

2004-12-14 12:49:10

Modified:

2004-12-14 14:49:10

Source: g = genus, infrao = infraorder

References: Kiessling 2004, Marsh 1875, Peczkis 1995

Age range: base of the Early/Lower Kimmeridgian to the top of the Maastrichtian or 154.80000 to 66.00000 Ma

Collections (157 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Kimmeridgian	154.8 - 152.21	France (Haute-Normandie)	Streptospondylus cuvieri (48856)
Early/Lower Berriasian - Late/Upper Albian	143.1 - 100.5	Mali (Gao)	Sauropoda indet. (25201)
Early/Lower Cretaceous	143.1 - 100.5	Kenya	Titanosauridae indet. (135809)
Cretaceous	143.1 - 66.0	Brazil (Mato Grosso)	Titanosauridae indet. (58812)
Aptian	121.4 - 113.2	Malawi (Northern)	Sauropodomorpha indet. (38916) Titanosauridae indet. (38912)
Aptian - Early/Lower Albian	121.4 - 110.1	USA (Wyoming)	Titanosauridae indet. (50355)
Aptian - Albian	121.4 - 100.5	USA (Utah)	Titanosauridae indet. (201626)
Late/Upper Aptian - Early/Lower Albian	119.57 - 110.1	South Korea (Gyeongsangnam-do)	Titanosauridae indet. (52863)
Albian - Cenomanian	113.2 - 93.9	Australia (Queensland)	Titanosauridae indet. (68155)
Cenomanian	100.5 - 93.9	Brazil (Maranhão)	Titanosauria indet. (56365)
Early/Lower Cenomanian	100.5 - 93.9	Morocco (Drâa-Tafilalet)	Titanosauridae indet. (155212 163917)
Early/Lower Cenomanian	100.5 - 93.9	Brazil (Rio Grande do Norte)	Titanosauridae indet. (224253)
Late/Upper Cenomanian - Early/Lower Turonian	100.5 - 89.8	Argentina (Chubut)	Titanosauridae indet. (51619)
Middle Cenomanian - Late/Upper Turonian	100.5 - 89.8	Argentina (Chubut)	Titanosaurinae indet. (68151)
Late/Upper Cretaceous	100.5 - 66.0	Chile (Copiapó)	Titanosauridae indet. (222946)
Late/Upper Cretaceous	100.5 - 66.0	Brazil (Mato Grosso)	Titanosauridae indet. (32335)
Late/Upper Turonian - Early/Lower Coniacian	93.9 - 85.7	Argentina (Neuquén)	Titanosauridae indet. (48174 49415 55443)
Late/Upper Coniacian - Santonian	89.8 - 83.6	Niger (Tahoua)	Titanosaurinae indet. (48867)
Coniacian - Maastrichtian	89.8 - 66.0	Brazil (Minas Gerais)	Titanosauridae indet. (51957)
Coniacian - Maastrichtian	89.8 - 66.0	Brazil (São Paulo)	Titanosauridae indet. (51626)
Early/Lower Santonian	85.7 - 83.6	South Africa (KwaZulu-Natal)	Titanosaurinae indet. (38158)
Santonian	85.7 - 83.6	Chile (Coquimbo)	Titanosauridae indet., Titanosaurinae indet., Antarctosaurus wichmannianus (51669)
Santonian	85.7 - 83.6	Argentina (Neuquén)	Titanosauridae indet. (51569 58749 84310)
Campanian	83.6 - 72.2	Argentina (Salta)	Titanosauridae indet. (51567)
Campanian	83.6 - 72.2	Bolivia (Chuquisaca)	Sauropoda indet. (36232)
Early/Lower Campanian - Middle Campanian	83.6 - 72.2	Argentina (Neuquén)	Titanosauridae indet. (51578 61454 61455 61457)
Late/Upper Campanian	83.6 - 72.2	France (Aquitaine-Limousin-Poitou-Charentes)	Titanosauridae indet. (178473)
Late/Upper Campanian	83.6 - 72.2	France (Aquitaine)	Titanosauridae indet. (68000)
Late/Upper Campanian	83.6 - 72.2	USA (New Mexico)	Titanosauridae indet. (52035 222078)
Middle Campanian	83.6 - 72.2	France (Provence-Alpes-Côte d'Azur)	Titanosauridae indet. (48849)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	France (Languedoc-Roussillon)	Titanosauridae indet. (74660)
Campanian - Maastrichtian	83.6 - 66.0	Kazakhstan (Almaty)	Titanosauridae indet. (53143 59732)
Campanian - Maastrichtian	83.6 - 66.0	Spain (Vasco-Cantabrica)	Titanosauridae indet. (208188)
Campanian - Maastrichtian	83.6 - 66.0	Kazakhstan (Aqmola)	Titanosauridae indet. (53140)
Campanian - Maastrichtian	83.6 - 66.0	Chile (Copiapó)	Titanosauridae indet. (223014)
Campanian - Maastrichtian	83.6 - 66.0	Sudan (Ash Shamaliyah)	Titanosauridae indet. (150678) Titanosaurus sp. (38682)
Campanian - Maastrichtian	83.6 - 66.0	China (Xinjiang)	Titanosauridae indet. (45171)
Campanian - Maastrichtian	83.6 - 66.0	Uruguay (Río Negro)	Dinosauria indet. (52094)
Campanian - Maastrichtian	83.6 - 66.0	Uruguay (Soriano)	Dinosauria indet. (219562)
Campanian - Maastrichtian	83.6 - 66.0	Chile (Atacama)	Titanosauridae indet. (35282 51671)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Spain (Castilla y Leon)	Dinosauria indet. (68458)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	France (Languedoc-Roussillon)	Sauropoda indet. (68875) Titanosauridae indet. (48870 59887)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	France (Provence-Alpes-Côte d'Azur)	Dinosauria indet. (114997) Hypselosaurus priscus, Titanosaurus indicus (48864)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Brazil (São Paulo)	Titanosauridae indet. (56653 56997 57000 57003 232337)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Argentina (Río Negro)	Titanosauridae indet. (31079 51593 51594 51595 51596 231684) Titanosauridae indet., Aeolosaurus rionegrinus (45087)
Late/Upper Campanian - Late/Upper Maastrichtian	83.6 - 66.0	France (Provence-Alpes-Cote d'Azur)	Titanosauridae indet. (59778)
Late/Upper Campanian - Maastrichtian	83.6 - 66.0	Argentina (Río Negro)	Titanosauridae indet. (51583)
Early/Lower Maastrichtian	72.2 - 66.0	Romania (Hunedoara)	Titanosauridae indet. (67811)
Early/Lower Maastrichtian	72.2 - 66.0	Spain (Cataluña)	Hypselosaurus sp. (183662) Titanosauridae indet. (59090)
Early/Lower Maastrichtian	72.2 - 66.0	France (Languedoc-Roussillon-Midi-Pyrénées)	Hypselosaurus priscus (91745)
Early/Lower Maastrichtian	72.2 - 66.0	France (Languedoc-Roussillon)	Dinosauria indet. (53483 55388 91093 91094) Hypselosaurus priscus (53480 91744) Titanosauridae indet. (113294)
Early/Lower Maastrichtian	72.2 - 66.0	France (Aude)	Titanosauridae indet. (48862)
Early/Lower Maastrichtian	72.2 - 66.0	France (Midi-Pyrénées)	Mammalia indet. (140926)
Early/Lower Maastrichtian	72.2 - 66.0	France (Provence-Alpes-Côte d'Azur)	Titanosauridae indet. (47893)
Late/Upper Maastrichtian	72.2 - 66.0	USA (New Mexico)	Titanosauridae indet. (56013 59068)
Late/Upper Maastrichtian	72.2 - 66.0	Pakistan (Balochistan)	Titanosauridae indet. (79666)
Late/Upper Maastrichtian	72.2 - 66.0	Brazil (Minas Gerais)	Titanosauria indet. (34410) Titanosauridae indet. (51662 51663 51959 219556 219726)
Late/Upper Maastrichtian	72.2 - 66.0	France (Haute-Garonne)	Titanosauridae indet. (67195 67196 118194 148954)
Late/Upper Maastrichtian	72.2 - 66.0	Brazil (São Paulo)	Titanosauridae indet. (32323 83263)
Late/Upper Maastrichtian	72.2 - 66.0	India (Tamil Nadu)	Titanosauridae indet. (53922)
Late/Upper Maastrichtian	72.2 - 66.0	Spain (Cataluña)	Hypselosaurus sp. (36381) Titanosauridae indet. (38893)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Texas)	Titanosauridae indet. (156122)
Maastrichtian	72.2 - 66.0	Mexico (Coahuila)	Titanosauridae indet. (99140)
Maastrichtian	72.2 - 66.0	India (Gujarat)	Ankylosauria indet. (95807)
Maastrichtian	72.2 - 66.0	Brazil (São Paulo)	Titanosaurinae indet. (51666)
Maastrichtian	72.2 - 66.0	Bolivia (Potosí)	Sauropoda indet. (36238) Titanosauridae indet. (219411)
Maastrichtian	72.2 - 66.0	France (Occitanie)	Titanosauridae indet. (199762)
Maastrichtian	72.2 - 66.0	Uruguay (Durazno)	Dinosauria indet. (223924) Titanosauridae indet. (219561)
Maastrichtian	72.2 - 66.0	France (Hérault)	Dinosauria indet. (59769 59770)
Maastrichtian	72.2 - 66.0	France (Var)	Titanosauridae indet. (13332)
Maastrichtian	72.2 - 66.0	France (Languedoc-Roussillon)	Titanosauridae indet. (68050)
Maastrichtian	72.2 - 66.0	Bolivia (Chuquisaca)	Titanosauridae indet. (219427)
Maastrichtian	72.2 - 66.0	Uruguay (Soriano)	Dinosauria indet. (52092 223918 223919) Laplatasaurus araukanicus (59147) Titanosauridae indet. (224202)
Maastrichtian	72.2 - 66.0	India (Madhya Pradesh)	Dinosauria indet. (222692) Titanosauridae indet. (26373) Titanosauridae indet. I, Titanosauridae indet. II (178977)
Maastrichtian	72.2 - 66.0	India (Maharashtra)	Titanosauridae indet. (68619)
Maastrichtian	72.2 - 66.0	Argentina (Santa Cruz)	Titanosauridae indet. (51624)
Maastrichtian	72.2 - 66.0	Brazil (Minas Gerais)	Titanosauridae indet. (51958 51960 219558)
Maastrichtian	72.2 - 66.0	Argentina (Chubut)	Titanosauridae indet. (231410)
Maastrichtian	72.2 - 66.0	Uruguay (Río Negro)	Dinosauria indet. (223920 223921 223925 224246) Laplatasaurus sp. (179789)
Maastrichtian	72.2 - 66.0	France (Provence-Alpes-Cote d'Azur)	Mastodontidae indet. (59781)
Maastrichtian	72.2 - 66.0	China (Jiangxi)	Titanosaurinae indet. (58443)
Maastrichtian	72.2 - 66.0	Madagascar (Mahajanga)	Titanosauridae indet. (55268 55269 55270 55271)
Maastrichtian	72.2 - 66.0	Uruguay (Canelones)	Titanosauridae indet. (219560)
Maastrichtian	72.2 - 66.0	Mexico (Puebla)	Sauropoda indet. (59365)
Maastrichtian	72.2 - 66.0	Uruguay (Paysandú)	Dinosauria indet. (204439 223923 224247)
Maastrichtian	72.2 - 66.0	France (Provence-Alpes-Côte d'Azur)	Hypselosaurus priscus (48863 53482 174674) Hypselosaurus sp. (59782 141628 141629) Rhabdodon priscum (141598) Titanosauridae indet. (51692)