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Homosphyronida

Arachnida - Pseudoscorpiones

Taxonomy
Homosphyronida was named by Chamberlin (1929). It is extant.

It was assigned to Pseudoscorpiones by Benavides et al. (2019).

Synonymy list
YearName and author
1929Homosphyronida Chamberlin
2019Homosphyronida Benavides et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
subphylumChelicerataHeymons 1901
—EuchelicerataWeygoldt and Paulus 1979
RankNameAuthor
—Prosomapoda
—Planaterga
—Dekatriata
—SclerophorataKamenz et al. 2011
classArachnidaLamarck 1801
—HaplocnemataBorner 1904
orderPseudoscorpionesLatreille 1817
—HomosphyronidaChamberlin 1929
—HomosphyronidaChamberlin 1929

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Homosphyronida Chamberlin 1929
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Subor. Atoposphyronida Harvey 2019
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Superfm. Feaelloidea Ellingsen 1906
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Fm. Feaellidae Ellingsen 1906
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Subfm. †Archaeofeaellinae Kolesnikov et al. 2022
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G. †Archaeofeaella Kolesnikov et al. 2022
Subfm. †Protofeaellinae Kolesnikov et al. 2022
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G. †Protofeaella Henderickx 2016
Fm. Pseudogarypidae Chamberlin 1923
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G. Pseudogarypus Ellingsen 1909
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†Pseudogarypus extensus Beier 1937
†Pseudogarypus hemprichii Koch and Berendt 1854
†Pseudogarypus minor Beier 1947
†Pseudogarypus pangaea Henderickx 2006
†Pseudogarypus synchrotron Henderickx 2012
Subor. Iocheirata Harvey 1992
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Infraor. Hemictenata Harvey 1992
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Superfm. Neobisioidea Chamberlin 1930
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Fm. Hyidae Chamberlin 1930
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G. Hya Chamberlin 1930
Fm. Ideoroncidae Chamberlin 1930
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G. †Proalbiorix Geißler et al. 2021
Fm. Neobisiidae Chamberlin 1930
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G. †Electrobisium Cockerell 1917
G. Microcreagris Balzan 1892
G. Neobisium Chamberlin 1930
G. Roncus Koch 1873
Invalid names: Bochicidae Chamberlin 1930 [empty], Gymnobisiidae Beier 1947 [empty], Parahyidae Harvey 1992 [empty], Syarinidae Chamberlin 1930 [empty]
Infraor. Panctenata Harvey 1992
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Superfm. Cheiridioidea Hansen 1894
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Fm. Cheiridiidae Hansen 1894
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G. Cheiridium Menge 1855
G. Cryptocheiridium Chamberlin 1931
Fm. Pseudochiridiidae Chamberlin 1923
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G. Pseudochiridium With 1906
Superfm. Cheliferoidea Risso 1826
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Fm. Atemnidae Kishida 1929
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Subfm. Atemninae Kishida 1929
G. †Progonatemnus Beier 1955
Fm. Cheliferidae Risso 1826
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Subfm. Cheliferinae Risso 1827
G. †Dichela Menge 1854
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Invalid names: Oligochelifer Beier 1937 [synonym]
G. †Electrochelifer Beier 1937
G. †Heurtaultia Judson 2009
G. Parawithius Chamberlin 1931
Subfm. Philomaoriinae Chamberlin 1931
G. †Pycnochelifer Beier 1937
G. †Trachychelifer Hong 1983
Fm. Chernetidae Menge 1855
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G. Americhernes Muchmore 1976
G. Byrsochernes Beier 1959
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Invalid names: Mayachernes Riquelme et al. 2014 [synonym]
G. Lustrochernes Beier 1932
G. †Oligochernes Beier 1937
G. Pachychernes Beier 1932
Fm. Withiidae Chamberlin 1931
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G. †Beierowithius Mahnert 1979
G. Chelifer Geoffroy 1762
G. Dolichowithius Beier 1932
G. Withius Kew 1911
Superfm. Garypinoidea Daday 1888
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Fm. Garypinidae Daday 1888
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Subfm. Amblyolpiinae Harvey 2023
Subfm. Garypininae Daday 1889
Subfm. Protogarypininae Harvey 2023
Invalid names: Larcidae Chamberlin 1930 [empty]
Superfm. Garypoidea Simon 1879
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Fm. Garypidae Simon 1879
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G. †Garypus Koch 1873
Fm. Geogarypidae Chamberlin 1930
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G. †Geogaranya Agnihotri et al. 2024
G. Geogarypus Chamberlin 1930
Invalid names: Hesperolpiidae Chamberlin 1930 [empty], Menthidae Chamberlin 1930 [empty], Olpiidae Banks 1895 [empty]
Superfm. Sternophoroidea Chamberlin 1923
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Fm. Sternophoridae Chamberlin 1923
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G. Idiogaryops Hoff 1963
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Grouping: solitaryc
Environment: terrestrialc
Locomotion: actively mobilec
Created: 2017-04-17 09:55:58
Modified: 2017-04-17 09:55:58
Source: c = class
Reference: Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Norian to the top of the Holocene or 227.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 205.7 Ma

Collections (34 total)


Time interval Ma Country or state Original ID and collection number
Norian227.3 - 205.7Ukraine (Kharkiv) Archaeofeaella henderickxi (105057)
Late/Upper Albian106.3 - 100.5France (Poitou-Charentes) Cheliferidae indet., Heurtaultia rossiorum (99114)
Late/Upper Albian106.3 - 100.5Spain (Álava) Cretogarypinus zaragozai, Ithioreolpium alavensis (107098)
Late/Upper Albian106.3 - 100.5Spain (Aragón) Pseudogarypus sp. (91626)
Early/Lower Cenomanian100.5 - 93.9Myanmar Electrobisium acutum, Garypus burmiticus (123020) Hya fynni (165561) Proalbiorix gracilis (172900) Protofeaella peetersae (125393)
Santonian85.7 - 83.6Hungary Ajkagarypinus stephani (224054)
Middle Campanian83.6 - 72.2Canada (Alberta) Chernetidae indet. (34896)
Ypresian56.0 - 48.07India (Gujarat) Geogaranya valiyaensis (183742)
Priabonian37.71 - 33.9Germany (Halle) Baltamblyolpium grabenhorsti (147606)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Baltamblyolpium gizmotum (110735) Chernetidae indet., Pseudogarypus minor, Electrochelifer bachofeni, Electrochelifer rapulitarsatus, Cheiridium hartmanni, Oligochernes bachofeni, Oligochelifer berendtii (123870) Microcreagris koellneri, Cheiridium sp. (120814) Neobisium henderickxi (123936) Pseudogarypus synchrotron (154709) Pseudogarypus synchrotron, Pseudogarypus pangaea (154708) Pycnochelifer sp., Cheiridium hartmanni, Neobisium exstinctum, Roncus succineus, Geogarypus macrodactylus, Progonatemnus succineus, Oligowithius sieboldi, Electrochelifer balticus (123771)
Priabonian37.71 - 33.9Russian Federation Garypinus electri (123419)
Priabonian37.71 - 33.9Ukraine Geogarypus gorskii (154710)
Priabonian37.71 - 33.9Poland Chelifer hemprichii, Chelifer kleemanni, Obisium rathkii (109625) Geogarypus gorskii (124519) Neobisium sp., Chelifer hemprichii, Chelifer kleemanni, Obisium rathkii, Chelifer wigandi, Chelifer hartmanni, Dichela berendtii, Obisium sieboldtii, Garypinus electri, Geogarypus macrodactylus, Geogarypus major, Pseudogarypus extensus, Oligochelifer gracilis, Oligochelifer serratidentatus, Oligochelifer granulatus, Electrochelifer mengei (120811)
Early/Lower Miocene23.04 - 15.98Mexico Chernetidae indet., Lustrochernes sp. 1 (21977) Paratemnoides sp., Lustrochernes sp. 2 (146607)
Early/Lower Miocene23.04 - 15.98Mexico (Chiapas) Cheliferini indet. (178679) Mayachernes maatiatus (163578)
Burdigalian20.45 - 15.98New Zealand (Otago) Philomaoria sp. (191650)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Idiogaryops pumilus (123422) Pachychernes sp., Pachychernes effossus, Americhernes sp., Cryptocheiridium antiquum, Parawithius sp. (122474) Pseudochiridium lindae (156229)
Holocene0.0117 - 0.0Colombia (Santander) Paratemnoides nidificator, Pachychernes subrobustus (147613)
Holocene0.0117 - 0.0Tanzania Chelifer eucarpus (152160)