PBDB Taxon

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Polypodiidae

Polypodiopsida

Taxonomy

Polypodiidae was named by Poche (1914). It is extant. It was considered monophyletic by Cronquist et al. (1966) and I (2016).

It was reranked as the subclass Polypodiidae by I (2016).

It was assigned to Filicopsida by Kenrick and Crane (1997); and to Polypodiopsida by Cronquist et al. (1966) and I (2016).

Subtaxa

Cyatheales, Gleicheniales, Hymenophyllales, Osmundales, Polypodiales, Salviniales (syn. Hydropteridales), Schizaeales

Synonymy list

Year	Name and author
1914	Polypodiidae Poche p. 133
1966	Polypodiidae Cronquist et al.
1997	Polypodiidae Kenrick and Crane
2016	Polypodiidae I

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Rank	Name	Author
kingdom	Plantae
phylum	Pteridophyta
class	Polypodiopsida

Rank	Name	Author
subclass	Polypodiidae	()
subclass	Polypodiidae	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subcl. Polypodiidae Poche 1914

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Or. Cyatheales Pryor et al. 1995

Fm. Cyatheaceae Kaulfuss 1827 [scaly tree fern]

G. Alsophila Brown 1810

Alsophila andersonii Scott 1866

Alsophila spinulosa Wallich 1828

G. †Alsophilidites Cookson

†Alsophilidites bellus Venkatachala et al. 1968

†Alsophilidites densus Singh et al. 1964

†Alsophilidites exilis Sah and Jain 1965

†Alsophilidites grandis Sah and Jain 1965

†Alsophilidites kerguelensis Cookson 1947

†Alsophilidites pannuceus Brenner 1963

G. †Alsophilites Hirmer 1927

†Alsophilites polonica Raciborski 1927

G. †Alsophilocaulis Menendez 1961

†Alsophilocaulis calveloi Menendez 1961

G. †Cibotiocaulis Ogura 1927

†Cibotiocaulis tateiwae Ogura 1927

G. Cibotium Kaulfuss 1820 [manfern]

G. †Conantiopteris Lantz et al. 1999

†Conantiopteris schuchmanii Lantz et al. 1999

G. †Coniopteris Brongniart 1849

†Coniopteris burejensis Zalessky 1904

†Coniopteris frutiformis Douglas 1973

†Coniopteris hymenophylloides Brongniart 1828

Invalid names: Adiantites amurensis Heer 1876 [synonym]

†Coniopteris murrayana Brongniart 1835

Invalid names: Polystichites Presl 1838 [replaced]

G. Cyathea Smith 1793

†Cyathea siwalika Bera et al. 2014

†Cyathea tchihatcheffi Schmalhausen 1879

Invalid names: Cnemidaria Presl 1836 [synonym], Hemitelia Brown 1810 [synonym], Trichopteris Hall 1845 [synonym]

G. †Cyatheites Goeppert 1836

†Cyatheites nebraskana Heer 1868

†Cyatheites schlotheimmii Goeppert 1836

G. †Cyatheopteris Schimper 1869

G. †Cyathocaulis Ogura 1927

†Cyathocaulis naktongensis Ogura 1927

G. †Cyathodendron Arnold 1945

†Cyathodendron texanum Arnold 1945

G. †Cyathoforma Bock 1969

†Cyathoforma carolinensis Emmons 1856

G. †Cyathoides Berry 1922

G. †Cyathorachis Ogura 1927

†Cyathorachis fujiiana Ogura 1927

G. †Dendropteridium Bancroft 1932

†Dendropteridium cyatheoides Bancroft 1932

G. †Dicksoniopsis Berry 1911

†Dicksoniopsis vernonensis Ward 1895

G. †Fasciostelopteris Stopes and Fujii 1910

†Fasciostelopteris tansleii Stopes and Fujii 1910

G. †Haydenia Seward 1912

†Haydenia thyrsopteroides Seward 1912

G. †Hemitelites Goeppert 1836

†Hemitelites hemitelioides Brongniart 1834

†Hemitelites torelli Heer 1869

G. †Kulyisporites Potonié 1956

†Kulyisporites lunaris Cookson and Dettmann 1958

G. †Protocyathea Feistmantel 1877

†Protocyathea rajmahalense Jacob 1937

†Protocyathea trichinopoliensis Feistmantel 1877

†Protocyathea ungeri Feistmantel 1877

G. †Pteridocaulis Bock 1969

†Pteridocaulis rhombiformis Bock 1969

G. †Tympanophora Lindley and Hutton 1835

†Tympanophora simplex Lindley and Hutton 1835

G. †Yezopteris Ogura 1930

†Yezopteris polycycloides Ogura 1930

G. †Hagiophyton Corsin 1948

Fm. †Loxsomataceae Presl

G. Loxsoma Hook and Brown 1847

G. †Loxsomopteris Skog 1976

Fm. Plagiogyriaceae Bower 1926

G. Plagiogyria Kunze 1858

Fm. Thyrsopteridaceae Presl 1847

G. Thyrsopteris Kunze 1834

†Thyrsopteris elegans Kunze 1818

†Thyrsopteris grevillioides Heer 1874

G. †Yavanna Vera 2013

†Yavanna chimaerica Vera 2013

Invalid names: Metaxyaceae Pichi-Sermolli 1970 [empty]

Or. Gleicheniales Jud 2011

Fm. †Dipteridaceae Seward and Dale

G. †Apachea Daugherty 1941

†Apachea arizonica Daugherty 1941

G. †Dictyophyllum Lindley and Hutton 1834

†Dictyophyllum ellenbergi Fabre and Greber 1960

†Dictyophyllum fuenzalidai Herbst 2000

†Dictyophyllum indicum Bose and Jana 1979

†Dictyophyllum nilssoni Brongniart 1836

†Dictyophyllum rugosum Lindley and Hutton 1834

†Dictyophyllum tenuifolium Stipanicic and Menendez 1949

G. Dipteris Reinw 1825 [fern]

G. †Hausmannia Dunker 1846

Subg. †Hausmannia (Protorhipis) Andrae 1853

†Hausmannia (Protorhipis) ussuriensis Kryshtofovich 1923

Invalid names: Hausmannia nariwaensis Oishi 1930 [synonym]

†Protorhipis fisheri Knowlton 1907

†Hausmannia buchii Andrae 1855

†Hausmannia californica Fontaine 1905

†Hausmannia crenata Nathorst 1878

Invalid names: Hausmannia indica Gupta 1955 [synonym]

†Hausmannia crookshankii Shah and Singh 1964

†Hausmannia dentata Oishi 1932

†Hausmannia dichotoma Dunker 1846

†Hausmannia leeiana Sze 1933

†Hausmannia montanensis Lapasha and Miller 1985

†Hausmannia pachyderma Sukh-Dev 1961

†Hausmannia papilio Herbst 1960

†Hausmannia papilionaceae Liu 1980

G. †Thaumatopteris Goeppert 1841

†Thaumatopteris barrealensis Stipanicic and Menendez 1949

†Thaumatopteris muensteri Goeppert 1841

†Thaumatopteris rothii Frenguelli 1941

Fm. Matoniaceae Presl 1847

G. †Alstaettia Remy and Remy 1969

†Alstaettia andersoni Remy and Remy 1969

G. †Barrealia Frenguelli 1942

†Barrealia dichotoma Frenguelli 1942

G. †Boseisporites Bharadwaj and Kumar 1972

†Boseisporites indicus Kumar 1973

†Boseisporites insignitus Venkatachala 1968

†Boseisporites lobatus Venkatachala 1968

†Boseisporites minutus Venkatachala et al. 1968

†Boseisporites praeclarus Sukh-Dev 1961

†Boseisporites punctatus Venkatachala 1968

G. †Dandotiaspora Sah

G. †Delosorus Skog 1988

†Delosorus heterophyllus Skog 1988

G. †Dictyophyllidites Couper 1958

†Dictyophyllidites arcuatus Pocknall and Mildenhall 1984

†Dictyophyllidites chiquimilense Mautino and Anzótegui 1998

†Dictyophyllidites complex Cookson and Dettmann 1958

†Dictyophyllidites concavus Harris 1965

†Dictyophyllidites crenatus Dettmann 1963

†Dictyophyllidites decus Kar 1970

†Dictyophyllidites equiexinus Dettman 1963

†Dictyophyllidites harrisii Couper 1958

†Dictyophyllidites mortoni Playford and Dettmann 1965

†Dictyophyllidites mortonii Playford and Dettmann 1965

†Dictyophyllidites pectinaeformis Dettmann 1963

†Dictyophyllidites speciosus Cookson and Dettman 1958

†Dictyophyllidites sukhdevii Maheshwari 1974

†Dictyophyllidites venkatachalai Ramanujam 1974

G. †Knowltonella Berry 1911

†Knowltonella maxoni Berry 1911

G. †Laccopteris Presl 1838

†Laccopteris elegans Presl 1838

†Laccopteris lanceolata Göppert 1838

G. Matonia Brown

G. †Matonidium Schenk 1871

†Matonidium althausii Dunker 1844

†Matonidium cingulatum Zeba-Bano and Bose 1981

†Matonidium goepperti Ettingshausen 1871

†Matonidium indicum Sahni 1936

G. †Matoniella Hirmer and Hoerhammer 1936

†Matoniella wiesneri Krasser 1936

G. †Nathorstia Heer 1880

†Nathorstia alata Halle 1913

†Nathorstia angustifolia Heer 1880

G. †Phlebomeris Saporta 1894

†Phlebomeris spectanda Saporta 1894

G. †Phlebopteris Brongniart 1836

†Phlebopteris arcuata Sukh-Dev 1961

†Phlebopteris athgarhensis Jain 1968

†Phlebopteris branneri White 1913

†Phlebopteris minutifolius Banerji 1982

†Phlebopteris polypodioides Brongniart 1836

G. †Piazopteris Lorch 1969

†Piazopteris branneri White 1913

G. †Weichselia Stiehler 1857

†Weichselia ludovicae Stiehler 1857

†Weichselia reticulata Stokes and Webb 1824

Or. Hymenophyllales Frank 1877

Fm. †Hymenophyllaceae Link

G. †Eusphenopteris Weiss 1882

†Eusphenopteris tenella Brongniart 1882

G. †Excipulites Goeppert 1836

†Excipulites neesii Goeppert 1836

G. Mecodium Presl 1937

G. †Trichomanites Goeppert 1836

†Trichomanites myriophyllum Goeppert 1836

Invalid names: Trichomanoideae Presl 1843 [empty]

Or. Polypodiales Link 1833

Subor. †Aspleniineae Schneid and Rothf

Fm. Aspleniaceae Newman 1840 [spleenwort]

G. Asplenium Linnaeus 1753

†Asplenium albertum Dawson 1885

†Asplenium cecilensis Berry 1911

†Asplenium coloradense Knowlton 1917

†Asplenium crossii Knowlton 1898

†Asplenium dicksonianum Heer 1874

†Asplenium dubium Fontaine 1889

†Asplenium eolignitica Berry 1916

†Asplenium foersteri Debey and Ettingshausen 1859

†Asplenium haguei Knowlton 1899

†Asplenium hurleyensis Berry 1916

†Asplenium iddingsi Knowlton 1899

†Asplenium issiacense Fritel 1910

†Asplenium jerseyensis Berry 1911

†Asplenium magnum Knowlton 1899

†Asplenium martinianum Dawson 1885

†Asplenium neomexicanum Knowlton 1916

†Asplenium niobrara Dawson 1882

†Asplenium primero Knowlton 1917

†Asplenium raritanense Berry 1909

†Asplenium remotidens Knowlton 1899

†Asplenium subcretaceum Saporta 1868

†Asplenium tenellum Knowlton 1900

†Asplenium wyomingense Knowlton 1900

G. Cystocarpium Fraser-Jenkins 2008

Fm. Onocleaceae Pichi-Sermolli 1970

G. Onoclea Linnaeus 1753

†Onoclea fecunda Lesquereux 1872

†Onoclea hesperia Hickey 1977

†Onoclea inquirenda Hollick 1904

†Onoclea minima Knowlton 1899

†Onoclea neomexicana Knowlton 1916

Onoclea sensibilis Linnaeus 1753

Fm. Thelypteridaceae Pichi-Sermolli 1970

Subfm. Phegopteridoideae Salino et al. 2016

G. Phegopteris Fée 1852

Subfm. Thelypteridoideae Reed 1963

G. Cyclosorus Link 1833

G. Thelypteris Schmidel 1763

Fm. Athyriaceae Alston 1956

G. †Dickwhitea Karafit et al. 2006

†Dickwhitea allenbyensis Karafit et al. 2006

G. Diplazium Swartz 1800 [twinsorus fern]

Subor. Dennstaedtiineae Schwartsb and Hovenkamp 2016

Fm. Dennstaedtiaceae Lotsy 1909

G. †Dennastra McIver and Basinger 1993

G. Dennstaedtia Bernhardi 1800

†Dennstaedtia americana Knowlton 1910

Invalid names: Asplenium tenerum Lesquereux 1883 [synonym], Sphenopteris blomstrandi Heer 1868 [synonym]

†Dennstaedtia concava Bolchovitina 1956

†Dennstaedtia fremonti Hall 1843

†Dennstaedtia rajmahalensis Sharma 1975

†Dennstaedtia sorimarginata McIver and Basinger 1993

G. Histiopteris Smith 1875

G. Hypolepis Bernhardi 1805

†Hypolepis coloradensis Cockerell 1909

G. Microlepia Presl 1836

G. †Microlepiopsis Serbet and Rothwell 2003

†Microlepiopsis aulenbackii Serbet and Rothwell 2003

†Microlepiopsis bramanii Serbet and Rothwell 2003

G. Pteridium Gleditsch and Scopoli 1760

G. †Karstenia Goeppert 1836

†Karstenia omphalostigma Goeppert 1836

Subor. Lindsaeineae Lehtonen and Tuomisto 2016

Fm. Lindsaeaceae [lace fern]

G. †Odontosorites Kobayashi and Yosida 1944

†Odontosorites heerianus Yokoyama 1944

G. Sphenomeris Maxon 1913

G. Tapeinidium Christensen 1906

†Tapeinidium undulatum Hall 1842

Subor. Polypodiineae Dumort 1829

Fm. Davalliaceae Schomb 1848

G. Davallia Smith 1793

†Davallia montana Knowlton 1899

†Davallia walkeri Conran et al. 2010

Invalid names: Humata Cavanilles 1802 [synonym]

Fm. Dryopteridaceae Herter 1949

G. Athyrium Roth 1800

G. †Ctenis Lindley and Hutton 1834

†Ctenis auriculata Fontaine 1896

†Ctenis clarnoensis Erdei and Manchester 2015

†Ctenis falcata Lindley and Hutton 1834

†Ctenis grandifolia Fontaine 1896

†Ctenis imbricata Fontaine 1889

†Ctenis imjhiriensis Bose and Zeba-Bano 1979

†Ctenis orovillensis Fontaine 1896

†Ctenis sulcicaulis Phillips 1829

†Ctenis wardii Fontaine 1896

Subfm. Dryopteridoideae Link 1841

G. Dryopteris Adanson 1763

G. Polystichum Roth 1799

G. Nephrodium Smith 1841

G. †Olfersites Guembel 1859

†Olfersites dichotomus Guembel 1859

Invalid names: Schizeites Guembel 1859 [replaced]

Invalid names: Elaphoglossoideae Pichi-Sermolli 1975 [empty], Polybotryoideae Liu and Zhang 2016 [empty]

Fm. Nephrolepidaceae Pichi-Sermolli 1975

G. Nephrolepis Schott 1834

Fm. Oleandraceae Pichi-Sermolli 1965

G. Oleandra Cavanilles 1801

†Oleandra arctica Heer 1874

†Oleandra graminaefolia Knowlton 1907

Invalid names: Lomariopsidaceae Alston 1956 [empty]

G. †Polypodiisporites Potonié 1931

†Polypodiisporites echinatus Jaramillo and Dilcher 2001

†Polypodiisporites favus Potonie 1956

†Polypodiisporites impariter Potonié and Sah 1960

Invalid names: Polypodiaceaesporites tuberculensis Baksi 1962 [synonym]

†Polypodiisporites inangahuensis Pocknall and Mildenhall 1984

†Polypodiisporites ipsviciensis Playford and Dettmann 1965

†Polypodiisporites irregularis Pocknal and Mildenhall 1984

†Polypodiisporites mawkmaensis Dutta and Sah 1970

†Polypodiisporites minimus Khan and Martin 1971

†Polypodiisporites oligocenicus Sah and Dutta 1968

†Polypodiisporites pachyexinatus Jaramillo and Dilcher 2001

†Polypodiisporites radiatus Pocknall and Mildenhall 1984

†Polypodiisporites rajasthanensis Srivastava 1966

†Polypodiisporites speciosus Sah and Dutta 1968

Subor. Pteridineae Prado and Schuettp 2016

Fm. Pteridaceae Reichenbach

G. †Adiantopteris Vassilevskaja 1963

†Adiantopteris tripinnata Cladera et al. 2007

Subfm. Cheilanthoideae Horvat 1927

G. Cheilanthes Swartz 1806 [lip fern]

G. Pellaea Link 1841 [cliffbrake]

G. †Contignisporites Dettmann 1963

†Contignisporites burgeri Filatoff and Price 1988

†Contignisporites burgerii Filatoff and Price 1988

†Contignisporites cooksonae Dettmann 1963

†Contignisporites cooksoniae Dettmann 1963

†Contignisporites cooksonii Dettmann 1963

†Contignisporites crenatus Varma and Ramanujam 1984

†Contignisporites dettmannii Singh and Kumar 1966

†Contignisporites fornicatus Dettmann 1963

†Contignisporites glebulentus Dettmann 1963

†Contignisporites kutchensis Venkatachala et al. 1968

†Contignisporites medeae Filatoff and Price 1988

†Contignisporites multimuratus Dettmann 1963

†Contignisporites psilatus Singh and Kumar 1966

†Contignisporites triletus Venkatachala et al. 1968

Subfm. Cryptogrammoideae Lindsay 2009

G. Coniogramme Fée 1852

G. Cryptogramma Brown 1823 [fern]

G. Eriosorus Fée 1852

G. †Kolymella Samylina and Filipova 1970

†Kolymella raevskii Samylina and Filipova 1970

G. Onychium Kaulfuss 1820

Onychium japonicum Thunberg 1784

Subfm. Pteridoideae Link 1841

G. Pteris Linnaeus 1753

G. †Pterisisporites Sung and Zheng

Subfm. Vittarioideae Link 1841

G. Adiantum Linnaeus 1753

Subor. Saccolomatineae Hovenkamp 2016

Fm. Saccolomataceae Doweld 2008

G. Saccoloma Kaulfuss 1820

†Saccoloma gardneri Lesquereux 1873

G. †Saportaea Fontaine and White 1880

†Saportaea dichotoma Frenguelli 1942

†Saportaea salisburioides Fontaine and White 1880

Invalid names: Drynariaceae Ching 1978 [empty]

Or. Salviniales Link 1833

Fm. †Azollaceae Wettstein 1903

G. †Azollopsis Hall 1968

†Azollopsis intermedia Sweet and Hills 1974

†Azollopsis tormentosa Hall 1968

G. †Paleoazolla Archangelsky et al. 1999

†Paleoazolla patagonica Archangelsky et al. 1999

G. †Parazollopsis Lapasha and Miller 1985

Fm. †Salviniaceae Lestiboudois

G. †Ariadnaesporites Potonié 1956

†Ariadnaesporites micromedusus Stough 1968

†Ariadnaesporites spinocaperatus Phillips and Felix 1971

†Ariadnaesporites spinosus Salami 1983

G. Azolla Lamarck 1783

†Azolla andreisii De Benedetti et al. 2021

†Azolla colhuehuapensis Vallati et al. 2017

†Azolla coloniensis De Benedetti and Zamaloa 2018

†Azolla deccanensis Patil and Upadhya 1980

†Azolla indica Trivedi and Verma 1971

†Azolla intertrappea Sahni and Rao 1934

†Azolla keuja Hermsen et al. 2019

†Azolla nana Dorofeev 1980

†Azolla primaeva Arnold 1955

†Azolla prisca Reid and Chandler 1926

†Azolla schopfii Dijkstra 1961

†Azolla stanleyi Jain and Hall 1969

†Azolla velus Jain and Hall 1969

†Azolla ventricosa Dorofeev 1955

G. †Azollophyllum Penhallow 1890

†Azollophyllum primaevum Penhallow 1890

G. †Massulites Sahni and Rao 1943

†Massulites coelatus Sahni and Rao 1943

G. Salvinia Adanson 1763 [watermoss]

†Salvinia coahuilensis Weber 1973

†Salvinia elliptica Newberry 1894

†Salvinia formosa Heer 1859

†Salvinia graui Herbst et al. 1987

†Salvinia hainanensis Wang et al. 2014

†Salvinia intertrappea Mahabale 1950

†Salvinia preauriculata Berry 1925

Invalid names: Sausarospermum Sahni and Srivastava 1940 [nomen nudum]

Invalid names: Hydropteridales Willdenow [synonym]

Or. Schizaeales Jud 2011

Fm. Anemiaceae Link 1841

G. Anemia Swartz 1806

†Anemia elongata Newberry 1863

Invalid names: Anemia haydenii Lesquereux 1871 [synonym], Anemia perplexa Hollick 1898 [synonym]

†Anemia eocenica Berry 1916

†Anemia fremonti Knowlton 1917

†Anemia gracillima Lesquereux 1883

†Anemia hesperia Knowlton 1916

†Anemia occidentalis Knowlton 1917

†Anemia poolensis Chandler 1955

†Anemia robusta Hollick 1902

†Anemia subcretacea Gard. and Ett. 1880

†Anemia supercretacea Hollick 1902

G. †Palaeomohria Archangelsky 2009

†Palaeomohria patagonica Archangelsky 2009

G. Appendicisporites Weyland and Greifeld 1953

†Appendicisporites concentricus Kemp 1970

†Appendicisporites dentimarginatus Brenner 1963

†Appendicisporites distocarinatus Dettmann and Playford 1968

†Appendicisporites erdtmanii Pocock 1964

†Appendicisporites ethmos Delcourt and Sprumont 1959

†Appendicisporites giganticus Groot and Groot 1962

†Appendicisporites insignis Chlonova 1983

†Appendicisporites jansonii Pocock 1962

†Appendicisporites matesovae Norris 1967

†Appendicisporites potomacensis Brenner 1963

†Appendicisporites pschekhaensis Pocock 1964

†Appendicisporites robustus Kemp 1970

†Appendicisporites segmentus Brenner 1963

†Appendicisporites sellingi Pocock 1964

†Appendicisporites stylosus Deak 1963

†Appendicisporites tricornitatus Weyland and Greifeld 1953

†Appendicisporites unicus Markova 1961

G. †Convolutispora Hoffmeister et al. 1955

†Convolutispora perfecta Kumaran 1980

Fm. Lygodiaceae Roem 1840

G. Lygodium Swartz 1800 [climbing fern]

†Lygodium binervatum Lesquereux 1869

†Lygodium coloradensis Knowlton 1930

†Lygodium compactum Lesquereux 1868

†Lygodium dentoni Lesquereux 1875

†Lygodium dinmorphyllum Churchill 1969

†Lygodium kaulfussi Heer

Invalid names: Lygodium neuropteroides Lesquereux 1870 [synonym]

†Lygodium marvinei Lesquereux 1875

†Lygodium poolensis Chandler 1955

†Lygodium prestwichii Gardner and Ettingshausen 1880

†Lygodium trichomanoides Lesquereux 1874

G. †Nodosisporites Dettmann and Clifford 1992

†Nodosisporites crenimurus Davies 1986

†Nodosisporites macrobaculatus Archangelsky and Llorens 2005

G. †Plicatella Maljavkina 1949

†Plicatella baqueroensis Davies 1985

†Plicatella pseudotripartita Archangelsky and Archangelsky 2010

†Plicatella trichacantha Maljavkina 1949

†Plicatella tricornitata Potonié 1960

G. †Ruffordiaspora Dettmann and Clifford 1992

†Ruffordiaspora australiensis Dettmann and Clifford 1992

†Ruffordiaspora cardielensis Archangelsky and Archangelsky 2010

†Ruffordiaspora crassiterminatus Archangelsky and Archangelsky 2010

†Ruffordiaspora ludbrookiae Dettmann and Clifford 1992

†Ruffordiaspora ticoensis Archangelsky et al. 2008

Fm. †Schizaeaceae Kaulfuss

G. †Acrostichopteris Fontaine 1889

†Acrostichopteris adiantifolia Fontaine 1889

†Acrostichopteris cyclopteroides Fontaine 1889

Invalid names: Baieropsis denticulata Fontaine 1889 [synonym]

†Acrostichopteris expansa Fontaine 1889

†Acrostichopteris fimbriata Knowlton 1907

†Acrostichopteris longipennis Fontaine 1889

Invalid names: Acrostichopteris densifolia Fontaine 1889 [synonym], Baieropsis foliosa Fontaine 1889 [synonym]

†Acrostichopteris parvifolia Fontaine 1889

Invalid names: Acrostichopteris parcelobata Fontaine 1889 [synonym]

†Acrostichopteris pluripartita Fontaine 1889

Invalid names: Baieropsis longifolia Fontaine 1889 [synonym]

G. †Baieropsis Fontaine 1889

G. †Cicatricosisporites Potonie and Gelletich 1933

†Cicatricosisporites angustus Singh 1971

†Cicatricosisporites annulatus Archangelsky and Gamerro 1966

†Cicatricosisporites apicanalis Phillips and Felix 1971

†Cicatricosisporites aralica Brenner 1963

†Cicatricosisporites augustus Singh 1971

†Cicatricosisporites auritus Singh 1971

†Cicatricosisporites australiensis Potonié 1956

†Cicatricosisporites avnimelechi Horowitz 1970

†Cicatricosisporites baconicus Deak 1963

†Cicatricosisporites baqueroensis Archangelsky and Gamerro 1966

†Cicatricosisporites brevilaesuratus Kemp 1970

†Cicatricosisporites claricanalis Phillips and Felix 1971

†Cicatricosisporites cuneiformis Pocock 1964

†Cicatricosisporites dorogensis Potonie and Gelletich 1933

†Cicatricosisporites dunrobinensis Couper 1958

†Cicatricosisporites eocenicus Jansonius and Hill 1976

†Cicatricosisporites ethmos Archangelsky and Gamerro 1966

†Cicatricosisporites furcatus Deak 1963

†Cicatricosisporites giganteus Archangelsky and Gamerro 1966

†Cicatricosisporites hallei Delcourt and Sprumont 1955

†Cicatricosisporites hughesi Dettmann 1963

†Cicatricosisporites hughesii Dettmann 1963

†Cicatricosisporites kashmiriensis Ghosh and Lukose 1967

†Cicatricosisporites kedvesii Schrank 2000

†Cicatricosisporites ludbrookiae Dettmann et al. 1987

†Cicatricosisporites macrocostatus Baksi 1962

†Cicatricosisporites magnus Doring 1965

†Cicatricosisporites mediostriatus Pocock 1964

†Cicatricosisporites microstriatus Jardiné and Magloire 1965

†Cicatricosisporites minutaestriatus Pocock 1964

†Cicatricosisporites mohrioides Delcourt and Sprumont 1955

†Cicatricosisporites nuni Horowitz 1975

†Cicatricosisporites orbiculatus Singh 1964

†Cicatricosisporites patapscoensis Brenner 1963

†Cicatricosisporites potomacensis Brenner 1963

†Cicatricosisporites pramparoana Archangelsky and Archangelsky 2010

†Cicatricosisporites proxiradiatus Kemp 1970

†Cicatricosisporites pseudotripartitus Dettmann 1963

†Cicatricosisporites purbeckensis Norris 1969

†Cicatricosisporites recticicatricosus Doring 1965

†Cicatricosisporites salardii Dejax 1985

†Cicatricosisporites sewardi Delcourt and Sprumont 1955

†Cicatricosisporites sinuosus Hunt 1985

†Cicatricosisporites sprumontii Döring 1965

†Cicatricosisporites subrotundus Brenner 1963

†Cicatricosisporites tersus Wang 1990

†Cicatricosisporites ticoensis Archangelsky and Gamerro 1966

†Cicatricosisporites venustus Deak 1963

†Cicatricosisporites wakkachuensis Ghosh and Lukose 1967

G. †Cicatricososporites Pflug and Thomson 1953

†Cicatricososporites decussatus Jaramillo and Dilcher 2001

†Cicatricososporites eocenicus Jansonius and Hills 1976

G. †Distaltriangulisporites Singh 1971

†Distaltriangulisporites pelliculus Scott 1976

G. †Klukisporites Couper 1958

†Klukisporites antarcticus Cao 1994

†Klukisporites apunctus Venkatachala et al. 1968

†Klukisporites areolatus Singh 1971

†Klukisporites foveolatus Pocock 1965

†Klukisporites labiatus Baldoni and Archangelsky 1983

†Klukisporites lacunus Filatoff 1975

†Klukisporites neovariegatus Filatoff 1975

†Klukisporites pseudoreticulatus Couper 1958

†Klukisporites punctatus Venkatachala 1968

†Klukisporites scaberis Dettmann 1963

†Klukisporites tuberosus Archangelsky 1983

†Klukisporites variegatus Couper 1958

†Klukisporites venkatachalae Tripathi 1990

G. †Lygodiumsporites Potonié 1956

†Lygodiumsporites edelii Mathur and Chopra 1982

†Lygodiumsporites eocenicus Dutta and Sah 1970

†Lygodiumsporites lakiensis Sah and Kar 1969

G. †Microfoveolatosporites Nakoman 1966

†Microfoveolatosporites skottsbergii Srivastava 1971

G. †Mintopteris Radforth and Walton 1960

†Mintopteris hirsuta Radforth and Walton 1960

G. Mohria Swartz 1806

G. †Mohriopsis Appert 1973

†Mohriopsis plastica Appert 1973

G. †Neyvelia Thiergart and Frantz 1963

†Neyvelia riedeli Thiergart and Frantz 1963

G. †Norimbergia Gothan 1914

†Norimbergia braunii Goeppert 1914

G. †Pastoria Thiergart and Frantz 1963

†Pastoria schizoneura Thiergart and Frantz 1963

G. †Pelletieria Seward 1913

†Pelletieria valdensis Seward 1913

G. †Pelletixia Watson and Hill 1982

†Pelletixia amelguita Skog 1982

G. †Protornithopteris Reed 1947

†Protornithopteris fremonti Knowlton 1947

G. †Ruffordia Seward 1894

†Ruffordia acrodenta Fontaine 1890

†Ruffordia goepperti Dunker 1844

Invalid names: Cladophlebis sphenopteroides Fontaine 1889 [synonym], Sphenopteris pachyphylla Fontaine 1889 [synonym], Sphenopteris spatulata Fontaine 1889 [synonym], Sphenopteris thysopteroides Fontaine 1889 [synonym], Thyrsopteris heteromorpha Fontaine 1889 [synonym]

G. Schizaea Smith 1793

G. †Schizaeangium Bohra and Sharma 1978

†Schizaeangium jurassica Bohra and Sharma 1978

G. †Schizaeoisporites Delcourt and Sprumont 1955

†Schizaeoisporites crassimurus Dutta and Sah 1970

†Schizaeoisporites dorogensis Potonié 1951

†Schizaeoisporites ghoshii Ramanujam 1967

†Schizaeoisporites minimus Ramanujam 1967

†Schizaeoisporites minor Pocock 1964

†Schizaeoisporites phaseolus Delcourt and Sprumont 1955

†Schizaeoisporites pseudorogensis Weyland and Greifeild 1953

†Schizaeoisporites sinuata Ramanujam 1967

G. †Schizaeopsis Berry 1911

†Schizaeopsis americana Berry 1917

Invalid names: Baieropsis macrophylla Fontaine 1889 [synonym], Schizaeopsis expansa Fontaine 1911 [synonym]

G. †Schizaeopteris Stopes and Fujii 1910

†Schizaeopteris mesozoica Stopes and Fujii 1910

G. †Senftenbergia Corda 1845

†Senftenbergia elegans Corda 1845

G. †Undulatisporites Pflug 1953

†Undulatisporites microcutis Thomson and Pflug 1953

†Undulatisporites pannuceus Singh 1971

†Undulatisporites sinuosis Groot and Groot 1962

†Undulatisporites undulapolus Brenner 1963

G. †Waldenburgia Gothan 1950

†Waldenburgia corynepteroides Gothan 1950

Invalid names: Anemieae Swartz [empty], Lygodieae [empty]

Invalid names: Osmundales [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

No ecological data are available

Age range

Maximum range based only on fossils: base of the Givetian to the top of the Holocene or 387.70000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 383.7 Ma

Collections (2616 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Early/Lower Givetian	388.1 - 383.7	United Kingdom (Scotland)	Convolutispora cerebra (22772)
Givetian	387.7 - 382.7	United Kingdom (Scotland)	Convolutispora disparalis (22794)
Givetian	387.7 - 382.7	USA (Arizona)	Convolutispora sp. (22756)
Givetian - Early/Lower Frasnian	387.7 - 382.4	Belgium	Convolutispora sp. (22904)
Givetian - Frasnian	387.7 - 372.2	Canada (Northwest Territories)	Convolutispora flexosa (23422)
Middle Givetian	383.0 - 380.2	United Kingdom (Scotland)	Convolutispora cerebra (22763 22766 22770)
Frasnian	382.7 - 372.2	Canada (Quebec)	Convolutispora caperata (179176)
Frasnian - Late/Upper Devonian	382.7 - 358.9	USA (Maryland)	Convolutispora subtilis (23578)
Late/Upper Devonian	382.7 - 358.9	USA (Pennsylvania)	Convolutispora opressa (23012)
Late/Upper Devonian - Carboniferous	382.7 - 298.9	Australia (Western Australia)	Convolutispora fromensis (23153)
Late/Upper Givetian - Early/Lower Frasnian	380.2 - 382.4	Australia (Western Australia)	Convolutispora caementosa (23199 23201 23265 23267 23268 23269 23272)
Early/Lower Famennian	376.1 - 370.6	Brazil	Convolutispora caementosa (23271)
Late/Upper Famennian - Tournaisian	364.7 - 346.7	Australia (Western Australia)	Convolutispora ampla, Convolutispora fromensis, Convolutispora jugosa, Convolutispora mellita, Convolutispora sp. (23096) Convolutispora fromensis (23146)
Tournaisian	358.9 - 346.7	Australia (Western Australia)	Convolutispora mellita, Convolutispora jugosa, Convolutispora sp. (23141)
Mississippian	358.9 - 323.2	USA (Illinois)	Eusphenopteris morrowensis (13511)
Tournaisian - Serpukhovian	358.9 - 323.2	USA (Illinois)	Senftenbergia sp. (23914)
Carboniferous	358.9 - 298.9	Canada (New Brunswick)	Mintopteris hirsuta (12696) Pecopteris hemitelioides (12724) Saportaea dispar (12721) Saportaea dispar, Pecopteris hemitelioides, Mintopteris hirsuta (12426)
Carboniferous	358.9 - 298.9	USA (Indiana)	Senftenbergia pennaeformis (12668)
Carboniferous	358.9 - 298.9	USA (Georgia)	Eusphenopteris aldrichii (12700)
Carboniferous	358.9 - 298.9	USA (Alabama)	Eusphenopteris lobata (28985 28987 28988 28989 28990 28991 28992 31028 31029 31030 31031 31032 31033 31034 31035)