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Pachyrukhinae

Mammalia - Notoungulata

Taxonomy
Pachyrukhinae was named by Kraglievich (1934). It was considered monophyletic by Cerdeno and Bond (1998) and Croft and Anaya (2006).

It was reranked as the unranked clade Pachyrukhinae by Seoane et al. (2023).

It was assigned to Hegetotheriidae by Kraglievich (1934), Cerdeno and Bond (1998), Croft and Anaya (2006), Seoane et al. (2017), Vera and Ercoli (2018) and Seoane et al. (2019); and to Hegetotheriidae by Seoane et al. (2023).

Synonymy list
YearName and author
1934Pachyrukhinae Kraglievich
1998Pachyrukhinae Cerdeno and Bond p. 799
2006Pachyrukhinae Croft and Anaya pp. 393-396
2017Pachyrukhinae Seoane et al.
2018Pachyrukhinae Vera and Ercoli
2019Pachyrukhinae Seoane et al.
2023Pachyrukhinae Seoane et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
RankNameAuthor
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—AfrotheriaStanhope et al. 1998
—Paenungulatomorpha
—PaenungulataSimpson 1945
—Sudamericungulata
orderNotoungulata
suborderTypotheriaZittel 1892
—Hegetotheriidae(Ameghino 1894)
—Pachyrukhinae(Kraglievich 1934)
—Pachyrukhinae(Kraglievich 1934)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Pachyrukhinae Kraglievich 1934
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G. †Medistylus Stirton 1952
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†Medistylus dorsatus Ameghino 1904
Unr. †Pachyrukhini Seoane et al. 2023
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G. †Pachyrukhos Ameghino 1885
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†Pachyrucos elongatus Moreno 1888
†Pachyrucos moyani Ameghino 1885
†Pachyrucos politus Ameghino 1902
†Pachyrucos teres Ameghino 1889
†Pachyrukhos moyani Ameghino 1885
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Invalid names: Pachyrucos absis Ameghino 1889 [synonym], Pachyrucos naevius Ameghino 1889 [synonym], Pachyrucos trivius Ameghino 1889 [synonym], Pachyrukhos teres Ameghino 1889 [synonym]
†Pachyrukhos ngenwinkul Solórzano et al. 2023
†Pachyrukhos politus Ameghino 1901
†Pachyrukhos tribius Ameghino 1889
Invalid names: Pedotherium Burmeister 1888 [synonym], Propachyrucos depressus Roth 1898 [nomen dubium], Propachyrucos medianus Roth 1898 [nomen dubium]
G. †Paedotherium Burmeister 1888
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†Paedotherium bonaerense Ameghino 1887
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Invalid names: Pachyrucos maximus Ameghino 1908 [synonym], Pachyrucos miramarensis Ameghino 1908 [synonym], Paedotherium imperforatum Kraglievich 1926 [synonym], Paedotherium insigne Burmeister 1888 [synonym], Paedotherium isolinense Castellanos 1936 [objective synonym]
†Paedotherium borrelloi Zetti 1972
†Paedotherium ictus Ameghino 1889
†Paedotherium kakai Reguero et al. 2015
†Paedotherium minor Cabrera 1937
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Invalid names: Paedotherium affine Cabrera 1937 [synonym]
†Paedotherium typicum Ameghino 1887
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Invalid names: Paedotherium brusquitaense Ameghino 1908 [synonym], Paedotherium chapadmalense Ameghino 1908 [synonym], Paedotherium marplatense Ameghino 1908 [synonym]
Invalid names: Raulringueletia Zett 1972 [synonym]
G. †Tremacyllus Ameghino 1891
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†Tremacyllus impressus Ameghino 1888
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Invalid names: Tremacyllus chapalmalensis Ameghino 1908 [synonym], Tremacyllus diminutus Ameghino 1888 [synonym], Tremacyllus intermedius Rovereto 1914 [synonym], Tremacyllus novus Ameghino 1908 [synonym]
†Tremacyllus incipiens Rovereto 1914
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Invalid names: Tremacyllus latifrons Rovereto 1914 [synonym]
†Tremacyllus subdiminutus Rovereto 1914
G. †Propachyrucos Ameghino 1897
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G. †Prosotherium Ameghino 1897
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†Prosotherium garzoni Ameghino 1897
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Invalid names: Propachyrucos ameghinorum Simpson 1945 [synonym], Propachyrucos simpsoni Chaffee 1952 [synonym], Prosotherium quartum Ameghino 1901 [synonym], Prosotherium robustum Ameghino 1897 [synonym], Prosotherium triangulidens Ameghino 1897 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Maximum body mass: 1.3 kgs
Minimum body mass: 1.3 kgs
Body mass estimate: 1.3 kg
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: scansorialsubc
Diet: herbivoreo
Reproduction: viviparoussubc
Created: 2009-01-05 17:11:49
Modified: 2009-01-05 19:11:49
Source: s = species, o = order, subc = subclass, c = class, subp = subphylum
References: MacFadden et al. 1996, Carroll 1988, Solórzano et al. 2023, Lillegraven 1979, Hendy et al. 2009, Ji et al. 2002

Age range: base of the Deseadan to the top of the Lujanian or 29.00000 to 0.01200 Ma

Collections (121 total)


Time interval Ma Country or state Original ID and collection number
Deseadan29.0 - 21.0Argentina (Mendoza) Prosotherium sp., Propachyrucos smithwoodwardi (176466)
Deseadan29.0 - 21.0Argentina (Chubut) Propachyrucos ameghinorum, Medistylus dorsatus (146358) Propachyrucos simpsoni, Prosotherium garzoni (49851) Propachyrucos sp. (176465) Prosotherium garzoni, Prosotherium triangulidens, Phanophilus dorsatus, Medistylus dorsatus (14211)
Colhuehuapian21.0 - 17.5Argentina (Chubut) Pachyrucos indet. (176441) Pachyrukhos politus (141368) Pachyrukhos sp. (232404)
Colhuehuapian - Burdigalian21.0 - 15.97Argentina (Chubut) Pachyrukhos sp. (204204)
Burdigalian20.44 - 15.97Argentina (Chubut) Pachyrukhos sp. (176205)
Burdigalian - Langhian20.44 - 13.82Argentina (Santa Cruz) Pachyrukhos moyani (197691)
Santacrucian17.5 - 16.3Argentina (San Juan) Pachyrukhinae indet. (197900)
Santacrucian17.5 - 16.3Argentina (Santa Cruz) Pachyrukhos moyani (87314 176589 176590 210637 210638) Pachyrukhos moyanoi (210153) Pachyrukhos sp. (176569 176570 176587 176588)
Santacrucian - Laventan17.5 - 11.0Chile (Bío Bío) Pachyrukhinae indet., Paedotherium minor (140105)
Friasian16.3 - 15.5Chile Pachyrukhos sp. (176666)
Friasian16.3 - 15.5Chile (Aisén) Pachyrukhos sp. (139845)
Langhian - Serravallian15.97 - 11.62Argentina (Chubut) Pachyrukhos moyani (215594 215595)
Colloncuran15.5 - 13.8Argentina (Río Negro) Pachyrukhos sp. (140691)
Serravallian - Tortonian13.82 - 7.246Argentina (La Pampa) Paedotherium sp. (231228)
Tortonian11.62 - 7.246Argentina (Chubut) Tremacyllus sp. (201906)
Late/Upper Miocene11.608 - 5.333Argentina (Salta) Paedotherium kakai (192105) Paedotherium minor, Paedotherium kakai (197784)
Late/Upper Miocene - Early/Lower Pliocene11.608 - 3.6Argentina (Catamarca) Tremacyllus incipiens (232594)
Chasicoan10.0 - 9.0Argentina (La Pampa) Paedotherium minor (206264)
Chasicoan10.0 - 9.0Argentina (Buenos Aires) Paedotherium minor (79180)
Chasicoan - Huayquerian10.0 - 5.0Argentina (La Pampa) Paedotherium minor, Tremacyllus impressus (204494)
Huayquerian8.0 - 5.0Argentina (La Pampa) Paedotherium minor (192135 192166 204361 204363 206260 206262 211563 230751) Paedotherium minor, Tremacyllus impressus (87198 151500 152061 152063 206031) Paedotherium sp. (151499) Paedotherium sp., Paedotherium minor, Tremacyllus impressus (140925 151498) Tremacyllus sp., Paedotherium sp., Paedotherium minor, Tremacyllus impressus (204598)
Huayquerian8.0 - 5.0Argentina (Mendoza) Paedotherium sp., Tremacyllus impressus (231327) Paedotherium typicum, Tremacyllus impressus (200444)
Huayquerian8.0 - 5.0Argentina (La Rioja) Paedotherium minor (151506) Paedotherium minor, Tremacyllus sp. (55596)
Huayquerian8.0 - 5.0Argentina (Buenos Aires) Paedotherium bonaerense (61300 61304 61306 198868) Paedotherium minor (198270 198689 206032) Paedotherium minor, Tremacyllus impressus (61292 61299) Paedotherium sp. (198862)
Huayquerian8.0 - 5.0Argentina (Catamarca) Paedotherium sp. (157794) Tremacyllus incipiens (198547 206028)
Messinian7.246 - 5.333Argentina (La Pampa) Paedotherium minor (140932) Paedotherium sp., Paedotherium minor, Tremacyllus impressus (152067) Tremacyllus sp., Paedotherium minor, Tremacyllus impressus (141939)
Messinian7.246 - 5.333Argentina (Catamarca) Tremacyllus incipiens (198820)
Messinian - Zanclean7.246 - 3.6Argentina (Catamarca) Tremacyllus incipiens (204710)
Messinian - Zanclean7.246 - 3.6Argentina (Mendoza) Pachyrukhinae indet. (230923) Tremacyllus impressus (200118)
Early/Lower Pliocene5.333 - 3.6Argentina (Buenos Aires) Paedotherium sp. (191764 191767 191768 191769 191770 191773)
Zanclean5.333 - 3.6Argentina (Catamarca) Tremacyllus latifrons, Tremacyllus incipiens (198549)
Zanclean5.333 - 3.6Argentina (Buenos Aires) Paedotherium bonaerense (191775) Paedotherium bonaerense, Tremacyllus impressus (191776) Paedotherium sp. (191774)
Montehermosan5.0 - 3.8Argentina (Buenos Aires) Paedotherium bonaerense (198684 198685 198686 198687) Paedotherium insigne (145757) Paedotherium sp. (152064 152086) Paedotherium sp., Paedotherium bonaerense (152062) Paedotherium sp., Paedotherium bonaerense, Tremacyllus impressus (152087) Paedotherium sp., Paedotherium typicum, Paedotherium bonaerense, Tremacyllus impressus (152081 152084 152085 152088) Paedotherium typicum, Paedotherium bonaerense, Tremacyllus impressus (198863)
Montehermosan5.0 - 3.8Argentina (Mendoza) Tremacyllus sp., Pachyrukhinae indet. (189520)
Montehermosan - Chapadmalalan5.0 - 3.0Argentina (Córdoba) Paedotherium bonaerense (197904)
Montehermosan - Chapadmalalan5.0 - 3.0Argentina (Catamarca) Tremacyllus latifrons (157795)
Chapadmalalan4.0 - 3.0Argentina (Córdoba) Tremacyllus impressus (202212)
Chapadmalalan4.0 - 3.0Argentina (Buenos Aires) Paedotherium chapadmalense (198844) Paedotherium insigne (198699) Paedotherium typicum (202340)
Chapadmalalan4.0 - 3.0Argentina (San Luis) Paedotherium sp., Paedotherium typicum, Paedotherium insigne, Tremacyllus impressus (152073)
Late/Upper Pliocene3.6 - 2.588Argentina (Entre Rios) Paedotherium typicum (79631 207682)
Late/Upper Pliocene3.6 - 2.588Argentina (Buenos Aires) Paedotherium sp. (191760 191763 191771 199160)
Late/Upper Pliocene3.6 - 2.588Argentina (Córdoba) Paedotherium sp., Paedotherium isolinense (211573)
Late/Upper Pliocene3.6 - 2.588Argentina (Cordoba) Tremacyllus impressus (191133)
Piacenzian3.6 - 2.588Argentina (Córdoba) Tremacyllus impressus (225604)
Early/Lower Pleistocene2.588 - 0.781Argentina (Buenos Aires) Tremacyllus sp., Paedotherium sp. (71304)
Lujanian0.781 - 0.012Argentina (Buenos Aires) Paedotherium sp., Paedotherium typicum, Paedotherium insigne, Tremacyllus impressus (202096)
Lujanian0.781 - 0.012Argentina (Entre Ríos) Paedotherium typicum (212840)